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A Taxonomic Revision of Hymenophyllaceae
BLUMEA 51: 221–280 Published on 27 July 2006 http://dx.doi.org/10.3767/000651906X622210 A TAXONOMIC REVISION OF HYMENOPHYLLACEAE ATSUSHI EBIHARA1, 2, JEAN-YVES DUBUISSON3, KUNIO IWATSUKI4, SABINE HENNEQUIN3 & MOTOMI ITO1 SUMMARY A new classification of Hymenophyllaceae, consisting of nine genera (Hymenophyllum, Didymoglos- sum, Crepidomanes, Polyphlebium, Vandenboschia, Abrodictyum, Trichomanes, Cephalomanes and Callistopteris) is proposed. Every genus, subgenus and section chiefly corresponds to the mono- phyletic group elucidated in molecular phylogenetic analyses based on chloroplast sequences. Brief descriptions and keys to the higher taxa are given, and their representative members are enumerated, including some new combinations. Key words: filmy ferns, Hymenophyllaceae, Hymenophyllum, Trichomanes. INTRODUCTION The Hymenophyllaceae, or ‘filmy ferns’, is the largest basal family of leptosporangiate ferns and comprises around 600 species (Iwatsuki, 1990). Members are easily distin- guished by their usually single-cell-thick laminae, and the monophyly of the family has not been questioned. The intrafamilial classification of the family, on the other hand, is highly controversial – several fundamentally different classifications are used by indi- vidual researchers and/or areas. Traditionally, only two genera – Hymenophyllum with bivalved involucres and Trichomanes with tubular involucres – have been recognized in this family. This scheme was expanded by Morton (1968) who hierarchically placed many subgenera, sections and subsections under -
New Species and Records of Tree Ferns (Cyatheaceae, Pteridophyta) from the Northern Andes
Org. Divers. Evol. 6, Electr. Suppl. 13: 1 - 11 (2006) © Gesellschaft für Biologische Systematik URL: http://www.senckenberg.de/odes/06-13.htm URN: urn:nbn:de:0028-odes0613-1 New species and records of tree ferns (Cyatheaceae, Pteridophyta) from the northern Andes Marcus Lehnert Albrecht-von-Haller Institut, Abt. Systematische Botanik, Universität Göttingen, Untere Karspüle 2, 37073 Göttingen, Germany e-mail: [email protected] Received 7 September 2005 • Accepted 6 December 2005 Abstract Four new species of Cyatheaceae from Ecuador are described: Alsophila conantiana Lehnert, Cyathea brucei Lehnert, C. mora- nii Lehnert, and C. sylvatica Lehnert. Range extensions are documented for Alsophila esmeraldensis R.C. Moran and Cyathea macrocarpa (C. Presl) Domin. Keywords: Alsophila; Cyathea; Andes; Colombia; Ecuador; Guayana Highlands Introduction The pteridophyte flora of Ecuador is one of the richest of most species. These advances enable us to resurrect in the world. About 1300 species have been registe- some species that had been united with others; they red (Jørgensen and León-Yánez 1999), including 177 also allow several new species to be described. endemic species (Valencia et al. 2000). Though mem- bers of the tree fern family were collected and studied New species frequently in the past (Tryon 1970, 1971, 1976, 1986; Gastony 1973; Stolze 1974; Barrington 1978; Conant Alsophila conantiana Lehnert, sp. nov. 1983; Tryon and Stolze 1989), new discoveries are (Fig. 1) still being made (Moran 1991, 1995a, 1998; Lehnert Etymology. This species is named for David S. Co- 2003, 2004). The complex taxonomy of the tree ferns, nant, Lyndon State College, Vermont, to honor his fragmentary collections, inadequate descriptions, and work on Cyatheaceae and especially Alsophila, from special descriptive vocabulary all contribute to our which my studies have greatly benefitted. -
Papers and Proceedings of the Royal Society of Tasmania, Volume 127, 1993 23
Papers and Proceedings of the Royal Society of Tasmania, Volume 127, 1993 23 RAINFOREST IN EASTERN TASMANIA - FLORISTICS AND CONSERVATION by M.G. Neyland and M.J. Brown (with two tables, four text-figures and one appendix) NEYLAND, M. G. & BROWN, M. J., 1993 (31:viii): Rainforest in eastern Tasmania - floristics and conservation. Pap. Proc. R. Soc. Tasm. 127: 23-32. https://doi.org/10.26749/rstpp.127.23 ISSN 0080-4703. Parks and Wildlife Service, Department of Environment and Land Management, GPO Box 44A, Hobart, Tasmania 7001 (MGN); Forestry Commission, Macquarie Street, Hobart, Tasmania 7000 (MJB). Six floristic communities are described from rainforest in northern and eastern Tasmania. The communities occur in lower rainfall areas, where they are often restricted ro fire-protected sites. They have climatic envelopes which are significantly distinct from each other and from rainforest in higher rainfall areas. The conservation status of the communities is assessed. Key Words: rainforest, Tasmania, conservation, relicts. INTRODUCTION METHODS Temperate rainforests worldwide are restricted mainly to the TASFORHAB profiles (Peters 1984) were collected from coastal and maritime zones, and generally occur in areas of relict rainforest patches throughout the study area. These high rainfall (Kellogg 1992). All the rainforests of Tasmania profiles record the floristics, species abundance and the are relicts of extensive rainforest which once occurred on the structure of the forest. A number of profiles already on the ancient continent Gondwana (Hill 1990, Nelson 1981). TASFORHAB data base were used to cross-check results, Many of the genera which are characteristic of rainforest in and to locate potential rainforest sites. -
Cyathea Cunninghamii (Slender Treefern)
CyatheaListing Statement cunninghamii for Cyathea cunninghamii (slender treefern) slender treefern T A S M A N I A N T H R E A T E N E D F L O R A L I S T I N G S T A T E M E N T Image by Mike Garrett Scientific name: Cyathea cunninghamii Hook.f., Icon . Pl. 10, t.985 (1854) Common name: slender treefern (Wapstra et al. 2005) Group: vascular plant, pteridophyte, family Cyatheaceae Status: Threatened Species Protection Act 1995 : endangered Environment Protection and Biodiversity Conservation Act 1999 : Not Listed Distribution: Endemic: Not endemic to Tasmania Tasmanian NRM Regions: Cradle Coast, North and South Figure 1. Distribution of Cyathea cunninghamii in Plate 1. Cyathea cunninghamii : habit Tasmania (image by Oberon Carter) 1 Threatened Species Section – Department of Primary Industries, Parks, Water & Environment Listing Statement for Cyathea cunninghamii (slender treefern) IDENTIFICATION AND ECOLOGY black, dull, with numerous, very small, sharp Cyathea cunninghamii is a tall treefern in the tubercles. The scales at the base of the stipe are Cyatheaceae family. It has a slender trunk and papery, shiny, pale fawn to light brown (often small crown, and typically occurs along creeks with dark central streaks), 1 to 4 cm long, ovate in sheltered coastal fern gullies (Plate 1). to linear with hair-like tips (Figure 2). Recruitment is from spore, with plants reaching Lamina are dark green, sub-triangular to sub- maturity at an age of about 25 to 30 years. lanceolate, 3-pinnate with pinnae shorter near Cyathea cunninghamii may be recognised in the the stipe. -
Fern Gazette
THE FERN GAZETTE Edited by BoAoThomas lAoCrabbe & Mo6ibby THE BRITISH PTERIDOLOGICAL SOCIETY Volume 14 Part 3 1992 The British Pteridological Society THE FERN GAZETTE VOLUME 14 PART 3 1992 CONTENTS Page MAIN ARTICLES A Revised List of The Pteridophytes of Nevis - B.M. Graham, M.H. Rickard 85 Chloroplast DNA and Morphological Variation in the Fern Genus Platycerium(Polypodiaceae: Pteridophyta) - Johannes M. Sandbrink, Roe/and C.H.J. Van Ham, Jan Van Brederode 97 Pteridophytes of the State of Veracruz, Medico: New Records - M6nica Pa/acios-Rios 119 SHORT NOTES Chromosome Counts for Two Species of Gleichenia subgenus Mertensiafrom Ecuador - Trevor G. Walker 123 REVIEWS Spores of The Pteridophyta - A. C. Jermy 96 Flora Malesiana - A. C. Jermy 123 The pteridophytes of France and their affinities: systematics. chorology, biology, ecology. - B. A. Thoinas 124 THE FERN GAZ ETTE Volume 14 Pa rt 2 wa s publis hed on lO Octobe r 1991 Published by THE BRITISH PTERIDOLOGICAL SOCIETY, c/o Department of Botany, The Natural History Museum, London SW7 580 ISSN 0308-0838 Metloc Printers Ltd .. Caxton House, Old Station Road, Loughton, Essex, IG10 4PE ---------------------- FERN GAZ. 14(3) 1992 85 A REVISED LIST OF THE PTERIDOPHYTES OF NEVIS BMGRAHAM Polpey, Par, Cornwall PL24 2T W MHRICKARD The Old Rectory, Leinthall Starkes, Ludlow, Shropshire SY8 2HP ABSTRACT A revised list of the pteridophytes of Nevis in the Lesser Antilles is given. This includes 14 species not previously recorded for the island. INTRODUCTION Nevis is a small volcanic island in the West Indian Leeward Islands. No specific li st of the ferns has ev er been pu blished, although Proctor (1977) does record each of the species known to occur on the island. -
Tmesipteris Parva
Listing Statement for Tmespteris parva (small forkfern) Small forkfern Tmesipteris parva T A S M A N I A N T H R E A T E N E D F L O R A L I S T I N G S T A T E M E N T Image by Matthew Larcombe Scientific name: Tmesipteris parva , N.A.Wakef., Vict. Nat. 60: 143 (1944) Common name: small forkfern (Wapstra et al. 2005) Group: vascular plant, pteridophyte, family Psilotaceae Status: Threatened Species Protection Act 1995 : vulnerable Environment Protection and Biodiversity Conservation Act 1999 : Not Listed Distribution: Endemic status: Not endemic to Tasmania Tasmanian NRM Regions: North and Cradle Coast Figure 1 . Distribution of Tmesipteris parva in Tasmania Plate 1. Tmesipteris parva detail (Image by Matthew Larcombe) 1 Threatened Species Section – Department of Primary Industries, Parks, Water and Environment Listing Statement for Tmespteris parva (small forkfern) IDENTIFICATION AND ECOLGY Naracoopa on King Island. The species has also Tmesipteris parva is a small fern in the Psilotaceae been collected from a tributary of the Grassy family, known in Tasmania from Flinders River on King Island (Garrett 1996, Chinnock Island and King Island. The species occurs in 1998) (see Table 1). The linear extent of the three sites in Tasmania is 350 km, the extent of sheltered fern gullies, where it grows on the 2 trunks of treeferns (Plates 1 and 2). occurrence 2,800 km (which includes large areas of sea), and the area of occupancy is less The species has rhizomes that are buried deeply than 1 ha. within the fibrous material of treefern trunks. -
PTERIDOLOGIST 2012 Contents: Volume 5 Part 5, 2012 Scale Insect Pests of Ornamental Ferns Grown Indoors in Britain
PTERIDOLOGIST 2012 Contents: Volume 5 Part 5, 2012 Scale insect pests of ornamental ferns grown indoors in Britain. Dr. Chris Malumphy 306 Familiar Ferns in a Far Flung Paradise. Georgina A.Snelling 313 Book Review: A Field Guide to the Flora of South Georgia. Graham Ackers 318 Survivors. Neill Timm 320 The Dead of Winter? Keeping Tree Ferns Alive in the U.K. Mike Fletcher 322 Samuel Salt. Snapshots of a Victorian Fern Enthusiast. Nigel Gilligan 327 New faces at the Spore Exchange. Brian and Sue Dockerill 331 Footnote: Musotima nitidalis - a fern-feeding moth new to Britain. Chris Malumphy 331 Leaf-mining moths in Britain. Roger Golding 332 Book Review: Ferns of Southern Africa. A Comprehensive Guide. Tim Pyner 335 Stem dichotomy in Cyathea australis. Peter Bostock and Laurence Knight 336 Mrs Puffer’s Marsh Fern. Graham Ackers 340 Young Ponga Frond. Guenther K. Machol 343 Polypodium Species and Hybrids in the Yorkshire Dales. Ken Trewren 344 A Challenge to all Fern Lovers! Jennifer M. Ide 348 Lycopodiums: Trials in Pot Cultivation. Jerry Copeland 349 Book Review: Fern Fever. Alec Greening 359 Fern hunting in China, 2010. Yvonne Golding 360 Stamp collecting. Martin Rickard 365 Dreaming of Ferns. Tim Penrose 366 Variation in Asplenium scolopendrium. John Fielding 368 The Case for Filmy Ferns. Kylie Stocks 370 Polystichum setiferum ‘Cristato-gracile’. Julian Reed 372 Why is Chris Page’s “Ferns” So Expensive? Graham Ackers 374 A Magificent Housefern - Goniophlebium Subauriculatum. Bryan Smith 377 A Bolton Collection. Jack Bouckley 378 360 Snails, Slugs, Grasshoppers and Caterpillars. Steve Lamont 379 Sphenomeris chinensis. -
Cyathea Medullaris
Cyathea medullaris COMMON NAME Black tree fern, mamaku, black mamaku SYNONYMS Sphaeropteris medullaris (G.Forst.) Bernh.; Cyathea medullaris var. polyneuron (Colenso) C.Chr.; Cyathea medullaris var. integra Hook.; Cyathea polyneuron Colenso; Polypodium medullare G.Forst.; FAMILY Cyatheaceae AUTHORITY Cyathea medullaris (G. Forst.) Sw. FLORA CATEGORY Vascular – Native ENDEMIC TAXON No ENDEMIC GENUS No ENDEMIC FAMILY No STRUCTURAL CLASS Ferns NVS CODE Stokes Valley. Dec 2004. Photographer: CYAMED Jeremy Rolfe CHROMOSOME NUMBER 2n = 138 CURRENT CONSERVATION STATUS 2012 | Not Threatened PREVIOUS CONSERVATION STATUSES 2009 | Not Threatened 2004 | Not Threatened BRIEF DESCRIPTION Large tree fern with black-stalked leaves to 5m long. Trunk with obvious scars from old leaves, to 20m tall. Leaf stems covered in small scales that have a spiny edge (lens needed). Sporangia arranged in small round capsules underneath leaves. DISTRIBUTION Indigenous. Occurring form the Three Kings Islands south to Stewart and the main Chatham Islands. Uncommon in the drier eastern portion of the South Island, and apparently absent from Canterbury and Otago. HABITAT Mamaku. Photographer: DoC Common in lowland forest throughout the North Island. Primarily in wetter coastal areas of the South Island. FEATURES Tree fern up to 20 m tall. Trunk black covered with hexagonal stipe bases. Stipes thick, black, harsh to touch, covered in black scales. Scales with marginal spines. Fronds up to 5 m long, arching upwards from crown, 3- pinnate, leathery, dead fronds falling (except in very young plants). Longest primary pinnae 0.4-1 m long, undersurfaces bearing scales with marginal spines. Indusia completely covering sori at maturity, splitting irregularly. SIMILAR TAXA Easily recognised by trunk with its distinctive hexagonal stipe scars and by the scales which possess marginal spines. -
Fern Gazette
THE .. FERN GAZETTE VOLUME ELEVEN PART FOUR 1976 THEJOURNAL OF THE BRITISHPTERIDOLOGICAL SOCIETY THE FERN GAZETTE VOLUME 11 PART 4 1976 CONTENTS Page MAIN ARTICLES Notes on some Mascarene species of Elaphoglossum - D. Lorence 199 Studying ferns in the Cameroons I. The lava ferns and their occurrence on Cameroon Mountain - G. Ben/ 207 The position of the megaprothallus of Salvinia natans - J.J. Schneller 217 A scanning electron microscope investigation of the spores of the genus Cystoperis - Ronald W. Pearman 221 Ecology and biogeography of New Zealand pteridophytes - B.S. Parris 231 Morphology of the sporophyte of the Vittarioid fern Ananthacorus Subhash Chandra 247 Six new species of Se/aginella from tropical South America - J.A. Crabbe and A. C. Jermy 255 Dryopteris caucasica, and thecytology of its hybrids - C.R. Fraser-Jenkins 263 SHORT NOTES Selaginella in Rajasthan, India - O.P. Sharma and T. N. Bhardwaja 268 ECOLOGICAL NOTES Ferns in canal navigations in Birmingham -A. R. Busby 269 REVIEWS. 205, 206, 216, 220, 246,253, 270 THE FERN GAZETTE Volume 11 Parts 2 & 3 was published 30th July, 1975 Published by THE BRITISH PTERIDOLOGiCAL SOCIETY, c/o Department of Botany, British Museum (Natural History), London SW7 5BD. Printed ECONOPRI NT L TO., Street, Edinburgh by 42A Albany ERRATUM in Fern Gaz.11:201 (1976): Amend caption to read: ...(d, fertile frond;) e, scale from rhizome x 25; f, scale from stipe x 50; g, scale from sterile lamina x 50. FERN GAZ. 11(4) 1976 199 NOTES ON SOME MASCARENE SPECIES OF ELAPHOGLOSSUM (LOMARIOPSIDOIDEAE SENSU HOL TTUM) D. -
Trichomanes Venosum
Trichomanes venosum COMMON NAME Veined bristle fern, veined filmy fern SYNONYMS Crepidomanes venosum (R.Br.) Bostock, Phlebiophyllum venosum (R.Br.) Bosch; Polyphlebium venosum (R.Br.) Copel. FAMILY Hymenophyllaceae AUTHORITY Trichomanes venosum R.Br. FLORA CATEGORY Vascular – Native ENDEMIC TAXON No Mangaone Track. Apr 1984. Photographer: Jeremy Rolfe ENDEMIC GENUS No ENDEMIC FAMILY No STRUCTURAL CLASS Ferns NVS CODE TRIVEN CHROMOSOME NUMBER 2n = 72 Coromandel. Photographer: John Smith- Dodsworth CURRENT CONSERVATION STATUS 2012 | Not Threatened PREVIOUS CONSERVATION STATUSES 2009 | Not Threatened 2004 | Not Threatened DISTRIBUTION Indigenous. New Zealand: Kermadec (Raoul Island), North, South, Stewart and Chatham Islands. Also Australia HABITAT Coastal to montane. A widespread and common filmy fern in the wetter parts of New Zealand. This species is usually found in closed forest where it the characteristic epiphyte of tree fern (Cyathea and Dicksonia spp.) trunks, though it will colonise other suitable trees. It is also found on rock faces, amongst moss on boulders, in tree caves and very rarely on damp, moss and liverwort encrusted ground. FEATURES Epiphytic (rarely terrestrial) fern usually forming dense carpets on suitable substrates. Rhizomes 0.2-0.8 mm diameter, widely creeping, much-branched and interwoven; densely hairy, hairs long, golden brown. Fronds 20-180 mm long, bright green, translucent, venation conspicuous. Stipe 8-55 mm long, slender, not winged. Rachises winged. Lamina 20-110 × 15-65 mm, lanceolate to elliptic. somewhat irregular, 1-pinnatifid, 1(-2)-pinnate, glabrescent (hairs when present unbranched). Ultimate segments 1.5-6.0 mm wide; margins crenate; apex obtuse, truncate, often notched; veins pinnately branched; false veins absent. -
3193VEA Otway Report PDF
ANGAHOOK-OTWAY INVESTIGATION DISCUSSION PAPER VICTORIAN ENVIRONMENTAL ASSESSMENT COUNCIL SEPTEMBER 2003 WHAT IS VEAC? The Victorian Environmental Assessment Council was established in 2002 – under the Victorian Environmental Assessment Council Act 2001 – to replace the Environment Conservation Council (ECC) as the body providing the State Government with independent advice on strategic public land-use planning. The five members of VEAC are: (Chair) Dr Brian Robinson,AM, FTSE, B.Sc., Ph.D. – The former chair of the Environment Protection Authority; a wealth of experience in sustainable development and natural resources management. Dr Sarah Ewing, B.Sc (Hons), M.Sc., Ph.D., Grad. Dip. Ed. – Currently a member of the Victorian Catchment Management Council and formerly a deputy member of the Australian Landcare Council; many years experience in catchment and natural resource management issues. Mr Duncan Malcolm – Currently the Chairman of the Gippsland Coastal Board and Watermark Inc.; immediate past Chairman of Lakes and Wilderness Tourism Association, immediate past chair and current board member of Irrigation Association of Australia and member of Victorian Coastal Council. Dr David Mercer, BA (Hons), Ph.D., Dip.Ed. – Fellow of The Environment Institute of Australia and New Zealand; broad expertise in natural resource management, recreation and tourism and also highly regarded academic experience. Mrs Eda Ritchie, Grad. Dip. Bus. – Formerly a member of the Environment Conservation Council,Trust for Nature Board and the Chairperson of the Western Region Coastal Board; has a strong background in farming in Western Victoria. SUBMISSIONS INVITED Submissions are now invited from interested groups and individuals for consideration for the Draft Proposals Paper to be published early in 2004. -
Conservation Status of New Zealand Indigenous Vascular Plants, 2012
NEW ZEALAND THREAT CLASSIFICATION SERIES 3 Conservation status of New Zealand indigenous vascular plants, 2012 Peter J. de Lange, Jeremy R. Rolfe, Paul D. Champion, Shannel P. Courtney, Peter B. Heenan, John W. Barkla, Ewen K. Cameron, David A. Norton and Rodney A. Hitchmough Cover: The Nationally Critical shrub Pittosporum serpentinum from the Surville Cliffs is severely affected by possums, and no seedlings have been found during recent surveys. Photo: Jeremy Rolfe. New Zealand Threat Classification Series is a scientific monograph series presenting publications related to the New Zealand Threat Classification System (NZTCS). Most will be lists providing NZTCS status of members of a plant or animal group (e.g. algae, birds, spiders). There are currently 23 groups, each assessed once every 3 years. After each 3-year cycle there will be a report analysing and summarising trends across all groups for that listing cycle. From time to time the manual that defines the categories, criteria and process for the NZTCS will be reviewed. Publications in this series are considered part of the formal international scientific literature. This report is available from the departmental website in pdf form. Titles are listed in our catalogue on the website, refer www.doc.govt.nz under Publications, then Science & technical. © Copyright August 2013, New Zealand Department of Conservation ISSN 2324–1713 (web PDF) ISBN 978–0–478–14995–1 (web PDF) This report was prepared for publication by the Publishing Team; editing by Amanda Todd and layout by Lynette Clelland. Publication was approved by the Deputy Director-General, Science and Capability Group, Department of Conservation, Wellington, New Zealand.