Insecta: Holometabola: Neuropterida: Neuroptera)
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UFRJ a Paleoentomofauna Brasileira
Anuário do Instituto de Geociências - UFRJ www.anuario.igeo.ufrj.br A Paleoentomofauna Brasileira: Cenário Atual The Brazilian Fossil Insects: Current Scenario Dionizio Angelo de Moura-Júnior; Sandro Marcelo Scheler & Antonio Carlos Sequeira Fernandes Universidade Federal do Rio de Janeiro, Programa de Pós-Graduação em Geociências: Patrimônio Geopaleontológico, Museu Nacional, Quinta da Boa Vista s/nº, São Cristóvão, 20940-040. Rio de Janeiro, RJ, Brasil. E-mails: [email protected]; [email protected]; [email protected] Recebido em: 24/01/2018 Aprovado em: 08/03/2018 DOI: http://dx.doi.org/10.11137/2018_1_142_166 Resumo O presente trabalho fornece um panorama geral sobre o conhecimento da paleoentomologia brasileira até o presente, abordando insetos do Paleozoico, Mesozoico e Cenozoico, incluindo a atualização das espécies publicadas até o momento após a última grande revisão bibliográica, mencionando ainda as unidades geológicas em que ocorrem e os trabalhos relacionados. Palavras-chave: Paleoentomologia; insetos fósseis; Brasil Abstract This paper provides an overview of the Brazilian palaeoentomology, about insects Paleozoic, Mesozoic and Cenozoic, including the review of the published species at the present. It was analiyzed the geological units of occurrence and the related literature. Keywords: Palaeoentomology; fossil insects; Brazil Anuário do Instituto de Geociências - UFRJ 142 ISSN 0101-9759 e-ISSN 1982-3908 - Vol. 41 - 1 / 2018 p. 142-166 A Paleoentomofauna Brasileira: Cenário Atual Dionizio Angelo de Moura-Júnior; Sandro Marcelo Schefler & Antonio Carlos Sequeira Fernandes 1 Introdução Devoniano Superior (Engel & Grimaldi, 2004). Os insetos são um dos primeiros organismos Algumas ordens como Blattodea, Hemiptera, Odonata, Ephemeroptera e Psocopera surgiram a colonizar os ambientes terrestres e aquáticos no Carbonífero com ocorrências até o recente, continentais (Engel & Grimaldi, 2004). -
From Chewing to Sucking Via Phylogeny—From Sucking to Chewing Via Ontogeny: Mouthparts of Neuroptera
Chapter 11 From Chewing to Sucking via Phylogeny—From Sucking to Chewing via Ontogeny: Mouthparts of Neuroptera Dominique Zimmermann, Susanne Randolf, and Ulrike Aspöck Abstract The Neuroptera are highly heterogeneous endopterygote insects. While their relatives Megaloptera and Raphidioptera have biting mouthparts also in their larval stage, the larvae of Neuroptera are characterized by conspicuous sucking jaws that are used to imbibe fluids, mostly the haemolymph of prey. They comprise a mandibular and a maxillary part and can be curved or straight, long or short. In the pupal stages, a transformation from the larval sucking to adult biting and chewing mouthparts takes place. The development during metamorphosis indicates that the larval maxillary stylet contains the Anlagen of different parts of the adult maxilla and that the larval mandibular stylet is a lateral outgrowth of the mandible. The mouth- parts of extant adult Neuroptera are of the biting and chewing functional type, whereas from the Mesozoic era forms with siphonate mouthparts are also known. Various food sources are used in larvae and in particular in adult Neuroptera. Morphological adaptations of the mouthparts of adult Neuroptera to the feeding on honeydew, pollen and arthropods are described in several examples. New hypoth- eses on the diet of adult Nevrorthidae and Dilaridae are presented. 11.1 Introduction The order Neuroptera, comprising about 5820 species (Oswald and Machado 2018), constitutes together with its sister group, the order Megaloptera (about 370 species), and their joint sister group Raphidioptera (about 250 species) the superorder Neuropterida. Neuroptera, formerly called Planipennia, are distributed worldwide and comprise 16 families of extremely heterogeneous insects. -
Comparative Study of Sensilla and Other Tegumentary Structures of Myrmeleontidae Larvae (Insecta, Neuroptera)
Received: 30 April 2020 Revised: 17 June 2020 Accepted: 11 July 2020 DOI: 10.1002/jmor.21240 RESEARCH ARTICLE Comparative study of sensilla and other tegumentary structures of Myrmeleontidae larvae (Insecta, Neuroptera) Fernando Acevedo Ramos1,2 | Víctor J. Monserrat1 | Atilano Contreras-Ramos2 | Sergio Pérez-González1 1Departamento de Biodiversidad, Ecología y Evolución, Unidad Docente de Zoología y Abstract Antropología Física, Facultad de Ciencias Antlion larvae have a complex tegumentary sensorial equipment. The sensilla and Biológicas, Universidad Complutense de Madrid, Madrid, Spain other kinds of larval tegumentary structures have been studied in 29 species of 2Departamento de Zoología, Instituto de 18 genera within family Myrmeleontidae, all of them with certain degree of Biología- Universidad Nacional Autónoma de psammophilous lifestyle. The adaptations for such lifestyle are probably related to México, Mexico City, Mexico the evolutionary success of this lineage within Neuroptera. We identified eight types Correspondence of sensory structures, six types of sensilla (excluding typical long bristles) and two Fernando Acevedo Ramos, Departamento de Biodiversidad, Ecología y Evolución, Unidad other specialized tegumentary structures. Both sensilla and other types of structures Docente de Zoología y Antropología Física, that have been observed using scanning electron microscopy show similar patterns in Facultad de Ciencias Biológicas, Universidad Complutense de Madrid, Madrid, Spain. terms of occurrence and density in all the studied -
Fossil Calibrations for the Arthropod Tree of Life
bioRxiv preprint doi: https://doi.org/10.1101/044859; this version posted June 10, 2016. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY 4.0 International license. FOSSIL CALIBRATIONS FOR THE ARTHROPOD TREE OF LIFE AUTHORS Joanna M. Wolfe1*, Allison C. Daley2,3, David A. Legg3, Gregory D. Edgecombe4 1 Department of Earth, Atmospheric & Planetary Sciences, Massachusetts Institute of Technology, Cambridge, MA 02139, USA 2 Department of Zoology, University of Oxford, South Parks Road, Oxford OX1 3PS, UK 3 Oxford University Museum of Natural History, Parks Road, Oxford OX1 3PZ, UK 4 Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK *Corresponding author: [email protected] ABSTRACT Fossil age data and molecular sequences are increasingly combined to establish a timescale for the Tree of Life. Arthropods, as the most species-rich and morphologically disparate animal phylum, have received substantial attention, particularly with regard to questions such as the timing of habitat shifts (e.g. terrestrialisation), genome evolution (e.g. gene family duplication and functional evolution), origins of novel characters and behaviours (e.g. wings and flight, venom, silk), biogeography, rate of diversification (e.g. Cambrian explosion, insect coevolution with angiosperms, evolution of crab body plans), and the evolution of arthropod microbiomes. We present herein a series of rigorously vetted calibration fossils for arthropod evolutionary history, taking into account recently published guidelines for best practice in fossil calibration. -
Neuroptera, Insecta)
Arthropod Structure & Development 37 (2008) 410–417 Contents lists available at ScienceDirect Arthropod Structure & Development journal homepage: www.elsevier.com/locate/asd Sperm ultrastructure and spermiogenesis of Coniopterygidae (Neuroptera, Insecta) Z.V. Zizzari, P. Lupetti, C. Mencarelli, R. Dallai* Department of Evolutionary Biology, University of Siena, Via Aldo Moro 2, I-53100 Siena, Italy article info abstract Article history: The spermiogenesis and the sperm ultrastructure of several species of Coniopterygidae have been ex- Received 16 January 2008 amined. The spermatozoa consist of a three-layered acrosome, an elongated elliptical nucleus, a long Accepted 17 March 2008 flagellum provided with a 9þ9þ3 axoneme and two mitochondrial derivatives. No accessory bodies were observed. The axoneme exhibits accessory microtubules provided with 13, rather than 16, protofilaments in their tubular wall; the intertubular material is reduced and distributed differently from that observed Keywords: in other Neuropterida. Sperm axoneme organization supports the isolated position of the family Insect spermiogenesis previously proposed on the basis of morphological data. Insect sperm ultrastructure 2008 Elsevier Ltd. All rights reserved. Electron microscopy Ó Insect phylogeny 1. Introduction appear normal in the larval instars, but progressively degenerate in the pupa so that in the adult only a ventral oval receptacle Neuropterida (Neuroptera sensu lato) comprise the orders filled with spermatozoa and secretion is evident; this single re- Raphidioptera (snakeflies), Megaloptera (alderflies and dobson- ceptacle is considered to be a seminal vesicle. A wide ductus flies) and the extremely heterogeneous Neuroptera (lacewings). ejaculatorius leads from the vesicula seminalis to the penis The first modern approach towards systematization of the Neuro- (Withycombe, 1925; Meinander, 1972). -
1 Universidade Federal Do Ceará Centro De Ciências
1 UNIVERSIDADE FEDERAL DO CEARÁ CENTRO DE CIÊNCIAS DEPARTAMENTO DE GEOLOGIA PROGRAMA DE PÓS-GRADUAÇÃO EM GEOLOGIA LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL FORTALEZA 2019 2 LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL Tese apresentada ao Programa de Pós- Graduação em Geologia da Universidade Federal do Ceará, como requisito parcial à obtenção do título de doutor em Geologia. Área de concentração: Geologia Sedimentar e Paleontologia. Orientador: Prof. Dr. Geraldo Jorge Barbosa de Moura. Coorientador: Prof. Dr. César Ulisses Vieira Veríssimo. FORTALEZA 2019 3 4 LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL Tese apresentada ao Programa de Pós- Graduação em Geologia da Universidade Federal do Ceará, como requisito parcial à obtenção do título de doutor em Geologia. Área de concentração: Geologia Sedimentar e Paleontologia. Aprovada em: 18/01/2019. BANCA EXAMINADORA ________________________________________ Prof. Dr. Geraldo Jorge Barbosa de Moura (Orientador) Universidade Federal Rural de Pernambuco (UFRPE) _________________________________________ Prof. Dr. Marcio Mendes Universidade Federal do Ceará (UFC) _________________________________________ Prof. Dr. Marcos Antônio Leite do Nascimento Universidade Federal do Rio Grande do Norte (UFRN) _________________________________________ Prof. Dr Kleberson de Oliveira Porpino Universidade do Estado do Rio Grande do Norte (UERN) ________________________________________ Dra Pâmela Moura Universidade Federal do Ceará (UFC) 5 A Deus. -
Psyche 85:265
AN UNUSUAL ASCALAPHID LARVA (NEUROPTERA: ASCALAPHIDAE) FROM SOUTHERN AFRICA, WITH COMMENTS ON LARVAL EVOLUTION WITHIN THE MYRMELEONTOIDEA* BY CHARLESS. HENRY Biological Sciences Group, University of Connecticut, Storrs 06268 Ascalaphidae is a fairly large family of planipennian Neuroptera that has received little attention from taxonomists since Weele's 1908 monograph. Life history and behavioral studies of the group have suffered even greater neglect; what is known has been sum- marized in a previous work (Henry, 1977). As an order, Neuroptera is taxonomically intractable, largely due to difficulty in interpreting wing venation: extreme convergence is common in distantly related families, yet divergence often occurs within a single subfamily or genus. For this reason, the immature stages have proven to be more reliable indicators of relationship than the adults, and our present understanding of evolution within the order is based more on larval than adult features (Withycombe, 1925; MacLeod, 1970). Since such considerations apply as strongly to Ascalaphidae as they do to other Neuroptera, all larval or life-history data are of paramount importance to phylogenetic studies of the family. Described here is a peculiar ascalaphid larva from Mkuze Game Park, Natal, collected in November of 1967 by J. A. Slater and T. Schuh. Although it shows all the diagnostic features of the family, certain details of its setal morphology and of the form and distribution of its thoracic and abdominal extensions (scoli) are unique and of great phylogenetic importance. As the larva is (necessarily) unassociated with an adult, I will describe its major features informally and compare them critically with those of larvae of known taxa in an attempt to deduce the general systematic position of the insect. -
Phylogeny and Bayesian Divergence Time Estimates of Neuropterida (Insecta) Based on Morphological and Molecular Data
Systematic Entomology (2010), 35, 349–378 DOI: 10.1111/j.1365-3113.2010.00521.x On wings of lace: phylogeny and Bayesian divergence time estimates of Neuropterida (Insecta) based on morphological and molecular data SHAUN L. WINTERTON1,2, NATE B. HARDY2 and BRIAN M. WIEGMANN3 1School of Biological Sciences, University of Queensland, Brisbane, Australia, 2Entomology, Queensland Primary Industries & Fisheries, Brisbane, Australia and 3Department of Entomology, North Carolina State University, Raleigh, NC, U.S.A. Abstract. Neuropterida comprise the holometabolan orders Neuroptera (lacewings, antlions and relatives), Megaloptera (alderflies, dobsonflies) and Raphidioptera (snakeflies) as a monophyletic group sister to Coleoptera (beetles). The higher-level phylogenetic relationships among these groups, as well as the family-level hierarchy of Neuroptera, have to date proved difficult to reconstruct. We used morphological data and multi-locus DNA sequence data to infer Neuropterida relationships. Nucleotide sequences were obtained for fragments of two nuclear genes (CAD, 18S rDNA) and two mitochondrial genes (COI, 16S rDNA) for 69 exemplars representing all recently recognized families of Neuropterida as well as outgroup exemplars from Coleoptera. The joint posterior probability of phylogeny and divergence times was estimated using a Bayesian relaxed-clock inference method to establish a temporal sequence of cladogenesis for the group over geological time. Megaloptera were found to be paraphyletic with respect to the rest of Neuropterida, calling into question the validity of the ordinal status for Megaloptera as presently defined. Ordinal relationships were weakly supported, and monophyly of Megaloptera was not recovered in any total- evidence analysis; Corydalidae were frequently recovered as sister to Raphidioptera. Only in relaxed-clock inferences were Raphidioptera and a paraphyletic Megaloptera recovered with strong support as a monophyletic group sister to Neuroptera. -
Evaluation of Tolerance and Susceptibility to Conventional and Biorational Insecticides in Chrysoperla Externa and C
Evaluation insecticides of tolerance in in Chrysoperla externa Chrysoperla and susceptibility Chrysopidae and to ) C. C. conventional asoralis ( Neuroptera and biorational : Evaluation of tolerance and susceptibility to Marina Haramboure conventional and biorational insecticides in Chrysoperla externa and C. asoralis (Neuroptera: Chrysopidae) Marina Haramboure 2016 ISBN 978-90-5989-874-5 Evaluation of tolerance and susceptibility to conventional and biorational insecticides in Chrysoperla externa and C. asoralis (Neuroptera: Chrysopidae) Marina Haramboure Thesis submitted for the degree of Doctor (PhD) in Applied Biological Sciences at the Faculty of Bioscience Engineering of Ghent University, Belgium, with co-tutelage with the Post-Graduate Program of the National University of La Plata, Argentina “the top environmental problems are selfishness, greed and apathy… … and to deal with those we need a spiritual and cultural transformation.” Gus Speth Promoters: Prof. Dr. ir. Guy Smagghe Department of Crop Protection Faculty of Bioscience Engineering Ghent University, Ghent, Belgium Dr. Marcela Inés Schneider Center of Parasitological and Vectors Studies Faculty of Natural Sciences and Museum National University of La Plata, Argentina Dr. Raúl Adolfo Alzogaray Scientific and Technical Research for Defence National University of San Martín, Argentina Chair of the examination Prof. Dr. Els J.M. Van Damme commitee: Department of Molecular Biotechnology Faculty of Bioscience Engineering Ghent University, Ghent, Belgium Members of the examination Prof. Dr. Nancy Greco commitee: Pests Ecology and Biological Control Center of Parasitological and Vectors Studies Faculty of Natural Sciences and Museum National University of La Plata, Argentina Prof. Dr. Gabriela Luna Pests Ecology and Biological Control Center of Parasitological and Vectors Studies Faculty of Natural Sciences and Museum National University of La Plata, Argentina Prof. -
Neuroptera: Myrmeleontidae) ⇑ ⇑ Bruno Michel A, , Anne-Laure Clamens B, Olivier Béthoux C,D, Gael J
Molecular Phylogenetics and Evolution 107 (2017) 103–116 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev A first higher-level time-calibrated phylogeny of antlions (Neuroptera: Myrmeleontidae) ⇑ ⇑ Bruno Michel a, , Anne-Laure Clamens b, Olivier Béthoux c,d, Gael J. Kergoat b,1, Fabien L. Condamine e, ,1 a CIRAD, UMR 1062 CBGP (INRA, IRD, CIRAD, Montpellier SupAgro), 755 Avenue du Campus Agropolis, 34988 Montferrier-sur-Lez, France b INRA, UMR 1062 CBGP (INRA, IRD, CIRAD, Montpellier SupAgro), 755 Avenue du Campus Agropolis, 34988 Montferrier-sur-Lez, France c Sorbonne Universités, UPMC Univ Paris 06, MNHN, CNRS, Centre de Recherche sur la Paléobiodiversité et les Paléoenvironnements (CR2P), Paris, France d Muséum National d’Histoire Naturelle, 57 rue Cuvier, CP38, F-75005 Paris, France e CNRS, UMR 5554 Institut des Sciences de l’Evolution (Université de Montpellier), Place Eugène Bataillon, 34095 Montpellier, France article info abstract Article history: In this study, we reconstruct the first time-calibrated phylogeny of the iconic antlion family, the Received 25 May 2016 Myrmeleontidae (Neuroptera: Myrmeleontiformia). We use maximum likelihood and Bayesian inference Revised 20 October 2016 to analyse a molecular dataset based on seven mitochondrial and nuclear gene markers. The dataset Accepted 21 October 2016 encompasses 106 species of Neuroptera, including 94 antlion species. The resulting phylogenetic frame- Available online 22 October 2016 work provides support for a myrmeleontid classification distinguishing four subfamilies: Acanthaclisinae, Myrmeleontinae, Palparinae, and Stilbopteryginae. Within Myrmeleontinae, Myrmecaelurini and Keywords: Nemoleontini are recovered as monophyletic clades; Gepini also appears as a valid tribe, distinct from Early Cretaceous Myrmecaelurini whereas Myrmecaelurini and Nesoleontini on one hand and Brachynemurini and Fossil calibrations Higher-level phylogeny Dendroleontini on the other hand, appear closely related. -
Neuroptera: Myrmeleontidae
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Center for Systematic Entomology, Gainesville, Insecta Mundi Florida March 1994 Reclassification of the New orldW antlion genera formerly included in the tribe Brachynemurini (Neuroptera: Myrmeleontidae Lionel A. Stange Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services, Gainesville, FL Follow this and additional works at: https://digitalcommons.unl.edu/insectamundi Part of the Entomology Commons Stange, Lionel A., "Reclassification of the New orldW antlion genera formerly included in the tribe Brachynemurini (Neuroptera: Myrmeleontidae" (1994). Insecta Mundi. 295. https://digitalcommons.unl.edu/insectamundi/295 This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Vol. 8, No. 1 - 2, March - June, 1994 67 Reclassification of the New World antlion genera formerly included in the tribe Brachynemurini (Neuroptera: Myrmeleontidae) Lionel A. Stange Florida State Collection of Arthropods Department of Agriculture & Consumer Services P.O. Box 147100 Gainesville, FL, 32614-7100 U.S.A. Abstract A cladistic analysis of the New World tribe Brachynemurini has resulted in several new taxonomic designations. The tribe is divided into 3 tribes, 2 of which are newly described. The Brachynemurini S.S. now contains 12 genera of which Argentoleon, Atricholeon, Mmleon and Venezueleon are newly described. The Gnopholeontini (NEW TRIBE) includes 4 North American genera whereas the Lemolemini (NEW TRIBE) contains 6 South American genera of which Ecualwn and Galapagolwn are newly described. -
The Function and Phylogenetic Implications of the Tentorium in Adult Neuroptera (Insecta)
Arthropod Structure & Development 40 (2011) 571e582 Contents lists available at ScienceDirect Arthropod Structure & Development journal homepage: www.elsevier.com/locate/asd The function and phylogenetic implications of the tentorium in adult Neuroptera (Insecta) Dominique Zimmermann a,*, Susanne Randolf a, Brian D. Metscher b, Ulrike Aspöck a a Natural History Museum, 2nd Zoological Department, Burgring 7, 1010 Vienna, Austria b Department of Theoretical Biology, University of Vienna, Althanstrasse 14, 1090 Wien, Austria article info abstract Article history: Despite several recent analyses on the phylogeny of Neuroptera some questions still remain to be Received 11 April 2011 answered. In the present analysis we address these questions by exploring a hitherto unexplored Accepted 12 June 2011 character complex: the tentorium, the internal cuticular support structure of the insect head. We described in detail the tentoria of representatives of all extant neuropteran families and the muscles Keywords: originating on the tentorium using 3D microCT images and analyzed differences in combination with Neuroptera a large published matrix based on larval characters. We find that the tentorium and associated Tentorium musculature are a source of phylogenetically informative characters. The addition of the tentorial Musculature Phylogeny characters to the larval matrix causes a basad shift of the Sisyridae and clearly supports a clade of all Function Neuroptera except Sisyridae and Nevrorthidae. A sister group relationship of Coniopterygidae and the Laminatentorium dilarid clade is further corroborated. A general trend toward a reduction of the dorsal tentorial arms and the development of laminatentoria is observed. In addition to the phylogenetic analysis, a correlation among the feeding habits, the development of the maxillary muscles, and the laminatentoria is demonstrated.