Carlquist Revisited: History, Success, and Applicability of a Natural History Model
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KS2 Tortoise Shapes and Sizes
TORTOISE SHAPES AND SIZE... ? Not all tortoises look the same. What can you learn about a tortoise from looking at the shape of its shell? Some Galapagos tortoises have Some Galapagos tortoises have domed shells like this saddleback shells like this An adaptation is a feature an animal has which helps it survive. If an animal has a greater chance of surviving then it has a greater chance of having more babies who are also likely to have the same adaptation. In the space below, draw one of our Galapagos giant tortoises. Make notes describing the TASK 1 different adaptations of the tortoise and how those features might help the tortoise survive in the wild. Worksheet 2 KS2 1 ? Not all the Galapagos Islands have the same habitat. What can you tell about the habitat of a tortoise by looking at the shape of its shell? Some of the Galapagos Islands are Some of the Galapagos Islands are smaller and dryer, where tall cacti larger and wetter, where many plants grow. plants grow close to the ground. TASK 2 Of the two tortoise shell shapes, which is likely to be better for reaching tall cacti plants? ______________________________________________________________________________________________ ______________________________________________________________________________________________ ______________________________________________________________________________________________ Which of these island types is likely to provide enough food for tortoises to grow to large sizes? ______________________________________________________________________________________________ -
An Introduction to Lemurs for Teachers and Educators
AN INTRODUCTION TO LEMURS FOR TEACHERS AND EDUCATORS WELCOME TO THE WORLD OF AKO THE AYE-AYE The Ako the Aye-Aye Educator’s Guide introduces you to the remarkable world of lemurs. This guide provides background information about the biological concepts conveyed through the 21 Ako lessons. These lessons were created to accompany the Ako books. The Ako book series were developed by renowned primatologist Alison Jolly for students in Madagascar to inspire understanding and appreciation for the unique primates that share their island home. In addition to the books there is also a set of posters which showcase the habitat of each lemur species and their forest “neighbors.” GOALS OF THE AKO LESSONS: • Inspire students to make a positive difference for lemurs and other wildlife. • Promote environmental awareness, understanding and appreciation. • Provide activities that connect students to nature and motivate conservation action. HOW TO USE THIS GUIDE Each lesson aligns with a specific grade level (Kindergarten-1st, 2nd-3rd and 4th-5th) and one of the seven environmental themes below. Before carrying out an activity, we recommend reading the corresponding section in this guide that matches the theme of the lesson. The themes are: • LOOKING AT LEMURS—CLASSIFICATION AND BIODIVERSITY (PAGE 4) • EXPLORING LEMUR HABITATS (PAGE 10) • INVESTIGATING LEMUR ADAPTATIONS (PAGE 18) • DISCOVERING LEMUR COMMUNITIES—INTER-DEPENDENCE (PAGE 23) • LEARNING ABOUT LEMUR LIFE—LIFE CYCLES AND BEHAVIOR (PAGE 26) • DISCOVERING MADAGASCAR’S PEOPLE AND PLACES (PAGE 33) • MAKING A DIFFERENCE FOR LEMURS (PAGE 40) Lessons can be completed chronologically or independently. Each activity incorporates multiple learning styles and subject areas. -
Polyploidy and the Evolutionary History of Cotton
POLYPLOIDY AND THE EVOLUTIONARY HISTORY OF COTTON Jonathan F. Wendel1 and Richard C. Cronn2 1Department of Botany, Iowa State University, Ames, Iowa 50011, USA 2Pacific Northwest Research Station, USDA Forest Service, 3200 SW Jefferson Way, Corvallis, Oregon 97331, USA I. Introduction II. Taxonomic, Cytogenetic, and Phylogenetic Framework A. Origin and Diversification of the Gossypieae, the Cotton Tribe B. Emergence and Diversification of the Genus Gossypium C. Chromosomal Evolution and the Origin of the Polyploids D. Phylogenetic Relationships and the Temporal Scale of Divergence III. Speciation Mechanisms A. A Fondness for Trans-oceanic Voyages B. A Propensity for Interspecific Gene Exchange IV. Origin of the Allopolyploids A. Time of Formation B. Parentage of the Allopolyploids V. Polyploid Evolution A. Repeated Cycles of Genome Duplication B. Chromosomal Stabilization C. Increased Recombination in Polyploid Gossypium D. A Diverse Array of Genic and Genomic Interactions E. Differential Evolution of Cohabiting Genomes VI. Ecological Consequences of Polyploidization VII. Polyploidy and Fiber VIII. Concluding Remarks References The cotton genus (Gossypium ) includes approximately 50 species distributed in arid to semi-arid regions of the tropic and subtropics. Included are four species that have independently been domesticated for their fiber, two each in Africa–Asia and the Americas. Gossypium species exhibit extraordinary morphological variation, ranging from herbaceous perennials to small trees with a diverse array of reproductive and vegetative -
HABITAT MANAGEMENT PLAN Green Bay and Gravel Island
HABITAT MANAGEMENT PLAN Green Bay and Gravel Island National Wildlife Refuges October 2017 Habitat Management Plans provide long-term guidance for management decisions; set forth goals, objectives, and strategies needed to accomplish refuge purposes; and, identify the Fish and Wildlife Service’s best estimate of future needs. These plans detail program planning levels that are sometimes substantially above current budget allocations and as such, are primarily for Service strategic planning and program prioritization purposes. The plans do not constitute a commitment for staffing increases, operational and maintenance increases, or funding for future land acquisition. The National Wildlife Refuge System, managed by the U.S. Fish and Wildlife Service, is the world's premier system of public lands and waters set aside to conserve America's fish, wildlife, and plants. Since the designation of the first wildlife refuge in 1903, the System has grown to encompass more than 150 million acres, 556 national wildlife refuges and other units of the Refuge System, plus 38 wetland management districts. This page intentionally left blank. Habitat Management Plan for Green Bay and Gravel Island National Wildlife Refuges EXECUTIVE SUMMARY This Habitat Management Plan (HMP) provides vision and specific guidance on enhancing and managing habitat for the resources of concern (ROC) at the refuge. The contributions of the refuge to ecosystem- and landscape-scale wildlife and biodiversity conservation, specifically migratory waterfowl, are incorporated into this HMP. The HMP is intended to provide habitat management direction for the next 15 years. The HMP is also needed to ensure that the refuge continues to conserve habitat for migratory birds in the context of climate change, which affects all units of the National Wildlife Refuge System. -
Towards a Panbiogeography of the Seas
Biological Journal of the Linnean Society, 2005, 84, 675-723. Towards a panbiogeography of the seas MICHAEL HEADS* Biology Department, University of the South Pacific, PO Box 1168, Suva, Fiji Islands Received 6 October 2003; accepted for publication 28 June 2004 A contrast is drawn between the concept of spéciation favoured in the Darwin-Wallace biogeographic paradigm (founder dispersal from a centre of origin) and in panbiogeography (vicariance or allopatry). Ordinary ecological dis persal is distinguished from founder dispersal. A survey of recent literature indicates that ideas on many aspects of marine biology are converging on a panbiogeographic view. Panbiogeographic conclusions supported in recent work include the following observations: fossils give minimum ages for groups and most taxa are considerably older than their earliest known fossil; Pacific/Atlantic divergence calibrations based on the rise of the Isthmus of Panama at 3 Ma are flawed; for these two reasons most molecular clock calibrations for marine groups are also flawed; the means of dispersal of taxa do not correlate with their actual distributions; populations of marine species may be closed systems because of self-recruitment; most marine taxa show at least some degree of vicariant differentiation and vicariance is surprisingly common among what were previously assumed to be uniform, widespread taxa; man grove and seagrass biogeography and migration patterns in marine taxa are best explained by vicariance; the Indian Ocean and the Pacific Ocean represent major -
The Biogeography of Large Islands, Or How Does the Size of the Ecological Theater Affect the Evolutionary Play
The biogeography of large islands, or how does the size of the ecological theater affect the evolutionary play Egbert Giles Leigh, Annette Hladik, Claude Marcel Hladik, Alison Jolly To cite this version: Egbert Giles Leigh, Annette Hladik, Claude Marcel Hladik, Alison Jolly. The biogeography of large islands, or how does the size of the ecological theater affect the evolutionary play. Revue d’Ecologie, Terre et Vie, Société nationale de protection de la nature, 2007, 62, pp.105-168. hal-00283373 HAL Id: hal-00283373 https://hal.archives-ouvertes.fr/hal-00283373 Submitted on 14 Dec 2010 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. THE BIOGEOGRAPHY OF LARGE ISLANDS, OR HOW DOES THE SIZE OF THE ECOLOGICAL THEATER AFFECT THE EVOLUTIONARY PLAY? Egbert Giles LEIGH, Jr.1, Annette HLADIK2, Claude Marcel HLADIK2 & Alison JOLLY3 RÉSUMÉ. — La biogéographie des grandes îles, ou comment la taille de la scène écologique infl uence- t-elle le jeu de l’évolution ? — Nous présentons une approche comparative des particularités de l’évolution dans des milieux insulaires de différentes surfaces, allant de la taille de l’île de La Réunion à celle de l’Amé- rique du Sud au Pliocène. -
Human Translocation As an Alternative Hypothesis to Explain the Presence of Giant Tortoises on Remote Islands in the Southwestern Indian Ocean
See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/298072054 Human translocation as an alternative hypothesis to explain the presence of giant tortoises on remote islands in the Southwestern Indian Ocean ARTICLE in JOURNAL OF BIOGEOGRAPHY · MARCH 2016 Impact Factor: 4.59 · DOI: 10.1111/jbi.12751 READS 63 3 AUTHORS: Lucienne Wilmé Patrick Waeber Missouri Botanical Garden ETH Zurich 50 PUBLICATIONS 599 CITATIONS 37 PUBLICATIONS 113 CITATIONS SEE PROFILE SEE PROFILE Jörg U. Ganzhorn University of Hamburg 208 PUBLICATIONS 5,425 CITATIONS SEE PROFILE All in-text references underlined in blue are linked to publications on ResearchGate, Available from: Lucienne Wilmé letting you access and read them immediately. Retrieved on: 18 March 2016 Journal of Biogeography (J. Biogeogr.) (2016) PERSPECTIVE Human translocation as an alternative hypothesis to explain the presence of giant tortoises on remote islands in the south-western Indian Ocean Lucienne Wilme1,2,*, Patrick O. Waeber3 and Joerg U. Ganzhorn4 1School of Agronomy, Water and Forest ABSTRACT Department, University of Antananarivo, Giant tortoises are known from several remote islands in the Indian Ocean Madagascar, 2Missouri Botanical Garden, (IO). Our present understanding of ocean circulation patterns, the age of the Madagascar Research & Conservation Program, Madagascar, 3Forest Management islands, and the life history traits of giant tortoises makes it difficult to com- and Development, Department of prehend how these animals arrived -
Lemurs, Little and Large
Lesson Description LEMURS, LITTLE Students practice measuring and learn that lemurs come in many sizes by measuring the length of Bitika the mouse lemur AND LARGE and other lemur species that she encounters on her evening adventure. The discussion focuses on lemur biodiversity (and 2-3rd grade also island gigantism and dwarfism) and the risks and benefits Language Arts, Science. Math of being big and small. Students will know that lemurs come in a variety of sizes and be able to use different tools ● Sets of marked lemur measurement rope and and units to measure them. identification, made using: ● Lemur Fact Cards ● Thick yarn or twine (15 feet long, one per group of ● Measure the length of different lemur species students plus one for the teacher) ● Describe why lemurs are so diverse in size ● Toilet paper, paper towel roll, or similar tube ● Measure lengths using a variety of tools ● Paper clips (8 per group of students ) ● Compare various units of measurement ● Colored tape (to mark lengths on rope) ● Describe why island species are sometimes very ● Masking tape to tape the ends of the large or very small lemur measuring rope to the floor ● Understand complexity of conservation issues ● Menabe-Antanimena Ako Poster i LOOKING AT LEMURS SETUP . Print out one set of Lemur Fact Cards for each rope. Cut out each card and hole-punch the corner. Place a paper clip through the hole. The paper clip will be used to attach the card at the distance representing the length of each lemur depicted. Cut one 15 foot length of yarn for each student group. -
Comparative Thinking in Biology
Comparative Thinking in Biology Adrian Currie This material has been published in the Cambridge University Press series Elements in the Philosophy of Biology, edited by Grant Ramsey and Michael Ruse. This version is free to view and download for personal use only. Not for re-distribution, re-sale or use in derivative works. © Adrian Currie. Acknowledgements I’m grateful to Marta Halina, Sabina Leonelli, Alison McConwell, Aaron Novick, Trevor Pearce, Russell Powell, William Wong and two anonymous referees for extremely useful and kind feedback on draft material. Also thanks to Grant Ramsey and Michael Ruse. Ideas from this element were presented to Exeter’s Cognition and Culture reading group, at Philosophy of Biology at Dolphin Beach 13 and at the Recent Trends in Philosophy of Biology conference in Bilkent; thanks to the audiences there. Many thanks to Kimberly Brumble for the wonderful illustrations. Some of the research for this Element was funded by the Templeton World Charity Foundation. Dedicated to the incomparable Kate. Table of Contents Cats versus Dogs............................................................................................................................... 1 1. Comparative Thinking .......................................................................................................... 14 1.1 Comparative Concepts .................................................................................................... 14 1.2 Two Kinds of Inference ................................................................................................... -
Did Lemurs Have Sweepstake Tickets? an Exploration of Simpson's Model for the Colonization of Madagascar by Mammals
Journal of Biogeography (J. Biogeogr.) (2006) 33, 221–235 ORIGINAL Did lemurs have sweepstake tickets? An ARTICLE exploration of Simpson’s model for the colonization of Madagascar by mammals J. Stankiewicz1, C. Thiart1,2, J. C. Masters3,4* and M. J. de Wit1 Departments of 1Geological Sciences and ABSTRACT 2Statistical Sciences, African Earth Observatory Aim To investigate the validity of Simpson’s model of sweepstakes dispersal, Network (AEON) and University of Cape Town, Rondebosch, 3Natal Museum, particularly as it applies to the colonization of Madagascar by African mammals. Pietermaritzburg and 4School of Biological & We chose lemurs as a classic case. Conservation Sciences, University of KwaZulu- Location The East African coast, the Mozambique Channel and Madagascar. Natal, Scottsville, South Africa Methods First, we investigated the assumptions underlying Simpson’s statistical model as it relates to dispersal events. Second, we modelled the fate of a natural raft carrying one or several migrating mammals under a range of environmental conditions: in the absence of winds or currents, in the presence of winds and currents, and with and without a sail. Finally, we investigated the possibility of an animal being transported across the Mozambique Channel by an extreme climatic event like a tornado or a cyclone. Results Our investigations show that Simpson’s assumptions are consistently violated when applied to scenarios of over-water dispersal by mammals. We suggest that a simple binomial probability model is an inappropriate basis for extrapolating the likelihood of dispersal events. One possible alternative is to use a geometric probability model. Our estimates of current and wind trajectories show that the most likely fate for a raft emerging from an estuary on the east coast of Africa is to follow the Mozambique current and become beached back on the African coast. -
Does Relaxed Predation Drive Phenotypic Divergence Among Insular Populations?
doi: 10.1111/jeb.12421 Does relaxed predation drive phenotypic divergence among insular populations? A. RUNEMARK*, M. BRYDEGAARD† &E.I.SVENSSON* *Evolutionary Ecology Unit, Department of Biology, Lund University, Lund, Sweden †Atomic Physics Division, Department of Physics, Lund University, Lund, Sweden Keywords: Abstract antipredator defence; The evolution of striking phenotypes on islands is a well-known phenome- body size; non, and there has been a long-standing debate on the patterns of body size coloration; evolution on islands. The ecological causes driving divergence in insular crypsis; populations are, however, poorly understood. Reduced predator fauna is lizards; expected to lower escape propensity, increase body size and relax selection Podarcis; for crypsis in small-bodied, insular prey species. Here, we investigated population divergence; whether escape behaviour, body size and dorsal coloration have diverged as variance. predicted under predation release in spatially replicated islet and mainland populations of the lizard species Podarcis gaigeae. We show that islet lizards escape approaching observers at shorter distances and are larger than main- land lizards. Additionally, we found evidence for larger between-population variation in body size among the islet populations than mainland popu- lations. Moreover, islet populations are significantly more divergent in dorsal coloration and match their respective habitats poorer than mainland lizards. These results strongly suggest that predation release on islets has driven population divergence in phenotypic and behavioural traits and that selective release has affected both trait means and variances. Relaxed preda- tion pressure is therefore likely to be one of the major ecological factors driving body size divergence on these islands. adjacent mainland localities. -
The Island Rule and Its Application to Multiple Plant Traits
The island rule and its application to multiple plant traits Annemieke Lona Hedi Hendriks A thesis submitted to the Victoria University of Wellington in partial fulfilment of the requirements for the degree of Master of Science in Ecology and Biodiversity Victoria University of Wellington, New Zealand 2019 ii “The larger the island of knowledge, the longer the shoreline of wonder” Ralph W. Sockman. iii iv General Abstract Aim The Island Rule refers to a continuum of body size changes where large mainland species evolve to become smaller and small species evolve to become larger on islands. Previous work focuses almost solely on animals, with virtually no previous tests of its predictions on plants. I tested for (1) reduced floral size diversity on islands, a logical corollary of the island rule and (2) evidence of the Island Rule in plant stature, leaf size and petiole length. Location Small islands surrounding New Zealand; Antipodes, Auckland, Bounty, Campbell, Chatham, Kermadec, Lord Howe, Macquarie, Norfolk, Snares, Stewart and the Three Kings. Methods I compared the morphology of 65 island endemics and their closest ‘mainland’ relative. Species pairs were identified. Differences between archipelagos located at various latitudes were also assessed. Results Floral sizes were reduced on islands relative to the ‘mainland’, consistent with predictions of the Island Rule. Plant stature, leaf size and petiole length conformed to the Island Rule, with smaller plants increasing in size, and larger plants decreasing in size. Main conclusions Results indicate that the conceptual umbrella of the Island Rule can be expanded to plants, accelerating understanding of how plant traits evolve on isolated islands.