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Maquetación 1 EARLY CRETACEOUS ORNITHOMIMOSAURS (DINOSAURIA: COELUROSAURIA) FROM AFRICA PAUL C. SERENO Department of Organismal Biology and Anatomy and Committee on Evolutionary Biology, University of Chicago, 1027 East 57th Street, Chicago, Illinois, 60637, U.S.A. Submitted: August 5 th , 2017 - Accepted: October 23 rd , 2017 - Published online: November 1 st , 2017 To cite this article: Paul C. Sereno (2017). Early Cretaceous ornithomimosaurs (Dinosauria: Coelurosauria) from Africa. Ameghiniana 54: 576–616. To link to this article: http://dx.doi.org/ 10.5710/AMGH.23.10.2017.3155 PLEASE SCROLL DOWN FOR ARTICLE Also appearing in this issue: Two new taxa unveil the A new ornithomimosaur taxon Murusraptor had a brain morphology previously unrecognized diversity from the Early Cretaceous of Niger similar to tyrannosaurids but of Coelophysidae in the Late Triassic and new anatomical data on neurosensorial capabilities of South America. Nqwebasaurus from South Africa. resembling that of allosauroids. ISSN 0002-7014 GONDWANAN PERSPECTIVES AMEGHINIANA - 2017 - Volume 54 (5): 576 – 616 EARLY CRETACEOUS ORNITHOMIMOSAURS (DINOSAURIA: COELUROSAURIA) FROM AFRICA PAUL C. SERENO Department of Organismal Biology and Anatomy and Committee on Evolutionary Biology, University of Chicago, 1027 East 57th Street, Chicago, Illinois, 60637, U.S.A. [email protected] Abstract . A new genus and species of ornithomimosaur, Afromimus tenerensis , is described based on a fragmentary skeleton from the Lower Cretaceous (Aptian–Albian) El Rhaz Formation of Niger. The holotype and only known individual preserves caudal vertebrae, chevrons and por - tions of the right hind limb. Derived ornithomimosaurian features include the broad, peanut-shaped articular surfaces of mid caudal centra, parasagittal fossae on mid caudal centra for reception of the postzygapophyses of the preceding vertebra, and a raised, subtriangular platform on the ventral aspect of the pedal phalanges. New information is given for, and comparisons made to, Nqwebasaurus thwazi from southern Africa, the oldest and most basal ornithomimosaur. Unlike other coelurosaurian clades that have expansive radiations on northern landmasses, the oldest ornithomimosaur and now another basal form are known from a southern landmass, Africa . Key words. Theropoda. Ornithomimosauria. Ornithomimid. Paleobiogeography. Gondwana. Afromimus . Nqwebasaurus . Resumen. ORNITHOMIMOSAURIOS DEL CRETÁCICO TEMPRANO (DINOSAURIA: COELUROSAURIA) DE ÁFRICA. Un nuevo género y especie de ornithomimosaurio, Afromimus tenerensis , es descrito en base a un esqueleto fragmentario de la Formación El Rhaz del Cretácico Temprano (Aptiano–Albiano) de Nigeria. El holotipo y único individuo conocido preserva vértebras caudales, chevrones, y portiones del miembro posterior derecho. Las características derivadas de ornithomimosaurios incluyen superficie articular de caudales medias en forma de maní, fosas parasa - gitales en centros de caudales medias para la recepción de postzygapófisis de la vértebra precedente, y una plataforma subtriangular elevada en la cara ventral de la falanges. La nueva información se da, y se compara con, Nqwebasaurus thwazi del sur de África, el ornitomimosaurio más antiguo y más basal. En contraposición a otros clados coelurosauriios que poseen una radiación expansiva en las masas continentales del he - misferio norte, el ornitomimosaurio más antiguo y ahora otra forma basal provienen de una masa continental del sur, África . Palabras clave. Theropoda. Ornithomimosauria. Ornitomímido. Oaleobiogeografía. Gondwana. Afromimus . Nqwebasaurus . ORNITHOMIMOSAURS are best known from Late Cretaceous postcrania from Valanginian-to-Berriasian-age rocks in deposits on northern landmasses. These well known genera southern England (Allain et al. , 2014), Nedcolbertia from include Gallimimus (Osmólska et al. , 1972), Garudimimus Berriasian-age rocks in Utah (Kirkland et al. , 1998; Brown - (Barsbold, 1981; Kobayashi and Barsbold, 2005a), Deino - stein, 2017), and Kinnareemimus from ?Berriasian-age rocks cheirus (Osmólska and Roniewicz, 1970; Lee et al. , 2014), in Thailand (Buffetaut et al. , 2009). Together, these wide - and Sinornithomimus (Kobayashi and Lü, 2003) from Asia and spread, fragmentary materials suggest that Early Creta - Ornithomimus (Sternberg, 1933) and Struthiomimus (Osborn, ceous strata on Laurasia will eventually reveal important 1917) from Laramidia. Similarly, older ornithomimosaurs of early stages in the evolution of Ornithomimosauria. late Early Cretaceous age (Berriasian to Aptian) also are Other taxa from Laurasian landmasses have been des- found only on northern landmassses. The most complete cribed recently as ornithomimosaurs, although their mor - of these include Harpymimus (Barsbold and Perle, 1984; phology and affinity remain poorly established. These Kobayashi and Barsbold, 2005b) and Shenzhosaurus (Ji et al. , include Hexing from ?Berriasian rocks in northern China (Jin 2003) from Asia, and Pelecanimimus (Perz-Moreno et al., et al. , 2012) and Lepidocheirosaurus from Late Jurassic 1994) and the Angeac ornithomimosaur (Allain et al. , 2014) (?Tithonian) rocks in Transbaikal Siberia (Alifanov and from western Europe. Saveliev, 2015). The partial holotypic skeleton of Hexing is Other fragmentary Early Cretaceous ornithomimosaurs very poorly preserved with some portions of the skeleton, from Laurasian rocks include Valdoraptor and other isolated such as the manus, exhibiting morphology and phalangeal 576 AMGHB2-0002-7014/12$00.00+.50 SERENO: EARLY AFRICAN ORNITHOMIMOSAURS formula unlike that of any ornithomimosaur. These same terms are used here as opposed to veterinarian directional deposits, it should be noted, yielded the well-preserved (e.g. , “rostral”) and anatomical ( e.g. , “alular digit”) terms used or nithomimosaur Shenzhousaurus (Ji et al. , 2003). The re - within the crown group Aves (see Wilson, 2006). Regarding mains of Lepidocheirosaurus are extremely fragmentary, the taxonomic terminology, Ornithomimosauria and Ornithomi - holotype limited to portions of the left manus and referred midae, are used as defined by Sereno (2005a,b), the former material to a few isolated caudal vertebrae. Its identifica - a stem-based taxon including Nqwebasaurus thwazi and all tion as an ornithomimosaur and the diagnosis of the species species closer to Ornithomimus edmontonicus than a range are not based on synapomorphies and autapomorphies, of other coelurosaurian clades (Tab. 2). Ornithomimidae is re spectively, and so the affinity and basis for this taxon re - also a stem-based taxon circumscribing the best known Late main problematic. Cretaceous species, while excluding the more basal Early Southern “ornithomimosaurs” reassigned . Initial reports of Cretaceous species Shenzhousaurus orientalis, Harpymimus or nithomimosaurs from southern landmasses have been okladnikovi, Pelecanimimis polyodon , and Garudimimus bre - reassigned to other theropod clades or questioned as or - vipes (Tab. 2). nithomimosaurs. The Late Jurassic Elaphosaurus bambergi Ornithomimoidea Marsh 1890, following traditional use from Tanzania, formerly interpreted as an ornithomimosaur of suffixes in Linnean taxonomy, should circumscribe a clade (Galton, 1982), has been reevaluated as an abelisauroid more inclusive than Ornithomimidae but less inclusive (noasaurid) theropod (Holtz, 1994; Rauhut and Caranno, than Ornithomimosauria, a suprafamilial taxon. As is ap - 2016). The Early Cretaceous (Aptian–Albian) Timimus her - par ent below, Nqwebasaurus thwazi is decisively more prim - mani from the southern coast of Australia (Rich and Rich, itive in many skeletal features than other species of Early 1994) was based on isolated femora more likely referable Cretaceous age ( e.g. , Shenzhousaurus oreintalis , Harpymimus to Dromaeosauridae (Agnolin et al. , 2010). okladnikovi and Pelecanimimis polyodon ). The taxon Ornitho - Nqwebasaurus , southern basal ornithomimosaur . The south - mimoidea, thus, could be a heuristic stem-based clade to ern African genus Nqwebasaurus (de Klerk et al. , 2000; encompass all ornithomimosaurs more advanced than Choiniere et al. , 2012) is known from a single partial skele - Nqwebasaurus thwazi . To that end, a definitional revision is ton discovered in Early Cretaceous (Berriasian–Valanginian) proposed here for Ornithomimoidea Marsh 1890 (Tab. 2), a rocks of southern Africa. Nqwebasaurus , first recognized as taxon initially defined by Sereno (1999a, 1999b) to encom - an ornithomimosaur by Sereno (2001), stands as the only pass ornithomimids and alvarezsaurids, prior to the dis - accepted representative of this clade from a southern cov ery of basal taxa within each subgroup that have land mass—and the oldest ornithomimosaur on record. undermined their monophyly). A new southern ornithomimosaur . Here I describe a second Fossil preservation . The holotype and only known specimen ornithomimosaur from Africa of slightly younger age than of Afromimus tenerensis (MNBH GAD112; Figs. 1–10) was Nqwebasaurus . The holotypic and only known specimen weathering on a nearly horizontal erosional surface and may was discovered in 1997 in the Ténéré Desert (Gadoufaoua) have been considerably more complete when originally of Niger in the Lower Cretaceous (Aptian–Albian) El Rhaz buried. No other vertebrate fossils were preserved at the For mation. Further details, in addition, are given for Nqwe - type locality, and no other specimens pertaining to an or - basaurus to supplement previous descriptions (de Klerk et nithomimosaur
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