Chapter 10 the Oldowan in North Africa Within a Biochronological Framework
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The Moroccan Perspective Jean-Paul Raynal, Fatima-Zohra Sbihi-Alaoui, Denis Geraads, Lionel Magoga, Abderrahim Mohib
The earliest occupation of North-Africa : the Moroccan perspective Jean-Paul Raynal, Fatima-Zohra Sbihi-Alaoui, Denis Geraads, Lionel Magoga, Abderrahim Mohib To cite this version: Jean-Paul Raynal, Fatima-Zohra Sbihi-Alaoui, Denis Geraads, Lionel Magoga, Abderrahim Mohib. The earliest occupation of North-Africa : the Moroccan perspective. Quaternary International, Else- vier, 2001, 75 (1), pp.65-75. halshs-00004433 HAL Id: halshs-00004433 https://halshs.archives-ouvertes.fr/halshs-00004433 Submitted on 3 Aug 2005 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. QI Durban 08/1999, Raynal et al. - p. 1 - THE EARLIEST OCCUPATION OF NORTH-AFRICA : THE MOROCCAN PERSPECTIVE J.P. RAYNAL1, F.Z. SBIHI ALAOUI2, D. GERAADS3, L. MAGOGA4, A. MOHIB5 1 Université de Bordeaux 1, UMR 5808 CNRS, Talence (France) et Mission "Littoral". 2 Institut National des Sciences de l'Archéologie et du Patrimoine, Ministère des Affaires Culturelles, Rabat (Maroc). 3 Musée de l'Homme, UMR 152 CNRS, Paris (France) et Mission "Littoral". 4 Mission "Littoral", Vichy (France). 5 Direction du Patrimoine, Inspection des Monuments et des Sites, Casablanca (Maroc). Abstract : The long sequence at Casablanca covers the last 5.5 Ma. -
First High Resolution Chronostratigraphy for the Early
www.nature.com/scientificreports OPEN First high resolution chronostratigraphy for the early North African Acheulean at Casablanca (Morocco) Rosalia Gallotti1,2,3*, Giovanni Muttoni4, David Lefèvre1,2, Jean‑Philippe Degeai1,2, Denis Geraads5, Andrea Zerboni4, Valérie Andrieu‑Ponel6, Matteo Maron7, Serena Perini4, Mohssine El Graoui8, Séverine Sanz‑Laliberté1,2, Camille Daujeard9, Paul Fernandes3,8,10, Mathieu Rué1,10, Lionel Magoga11, Abderrahim Mohib8,12 & Jean‑Paul Raynal3,8,13 The onset of the Acheulean, marked by the emergence of large cutting tools (LCTs), is considered a major technological advance in the Early Stone Age and a key turning point in human evolution. The Acheulean originated in East Africa at ~ 1.8–1.6 Ma and is reported in South Africa between ~ 1.6 and > 1.0 Ma. The timing of its appearance and development in North Africa have been poorly known due to the near‑absence of well‑dated sites in reliable contexts. The ~ 1 Ma stone artefacts of Tighennif (Algeria) and Thomas Quarry I‑Unit L (ThI‑L) at Casablanca (Morocco) are thus far regarded as documenting the oldest Acheulean in North Africa but whatever the precision of their stratigraphical position, both deserve a better chronology. Here we provide a chronology for ThI‑L, based on new magnetostratigraphic and geochemical data. Added to the existing lithostratigraphy of the Casablanca sequence, these results provide the frst robust chronostratigraphic framework for the early North African Acheulean and frmly establish its emergence in this part of the continent back at least to ~ 1.3 Ma. Over the last two decades, several eforts have been made to refne the chronostratigraphic framework of the Acheulean emergence in Africa. -
The Pleistocene Fauna (Other Than Primates) from Asbole, Lower Awash
Geobios 37 (2004) 697–718 http://france.elsevier.com/direct/GEOBIO/ Original article The Pleistocene fauna (other than Primates) from Asbole, lower Awash Valley, Ethiopia, and its environmental and biochronological implications La faune pléistocène (sauf Primates) d’Asbole, basse vallée de l’Awash, Éthiopie: implications environnementales et biochronologiques Denis Geraads a,*, Zeresenay Alemseged b, Denné Reed c, Jonathan Wynn d, Diana C. Roman e a UPR 2147 CNRS, 44, rue de l’Amiral Mouchez, 75014 Paris, France b Institute of Human Origins, PO Box 874101, Arizona State University, Tempe, AZ 85287-4101, USA c Department of Anthropology, SUNY Stony Brook, Stony Brook NY 11794-4364, USA d Department of Geography and Geosciences, Irvine Building, University of St. Andrews, St. Andrews, Fife, Scotland, KY169AL, United Kingdom e Department of Geological Sciences, 1272 University of Oregon, Eugene, OR 97403-1272, USA Received 9 December 2002; accepted 28 May 2003 Abstract The Asbole area in the Lower Awash Valley yielded a diverse fauna of large and small mammals, associated with an Acheulean industry. The most notable forms are a potentially new species of herpestid, a large collection of Kolpochoerus majus, and the earliest known Bos in Africa. Biochronologically, this fauna belongs to the earliest Middle Pleistocene, and is roughly contemporaneous with the Bodo site further south. Paleoenvironmentally, the fauna suggests a mosaic of landscapes among which humid environments, grasslands and forests, are pre- dominant. © 2004 Elsevier SAS. All rights reserved. Résumé La région d’Asbole dans la basse vallée de l’Awash a livré une faune diversifiée de grands et petits Mammifères associée à une industrie acheuléenne. -
Late Quaternary Extinction of Ungulates in Sub-Saharan Africa: a Reductionist's Approach
Late Quaternary Extinction of Ungulates in Sub-Saharan Africa: a Reductionist's Approach Joris Peters Ins/itu/ für Palaeoanatomie, Domestikationsforschung und Geschichte der Tiermedizin, Universität München. Feldmochinger Strasse 7, D-80992 München, Germany AchilIes Gautier Laboratorium VDor Pa/eonlo/agie, Seclie Kwartairpaleo1ltologie en Archeozoölogie, Universiteit Gent, Krijgs/aon 2811S8, B-9000 Gent, Belgium James S. Brink National Museum, P. 0. Box 266. Bloemfontein 9300. Republic of South Africa Wim Haenen Instituut voor Gezandheidsecologie, Katholieke Universiteit Leuven, Capucijnenvoer 35, B-3000 Leuven, Belgium (Received 24 January 1992, revised manuscrip/ accepted 6 November 1992) Comparative osteomorphology and sta ti st ical analysis cf postcranial limb bone measurements cf modern African wildebeest (Collnochaetes), eland (Taura/ragus) and Africa n buffala (Sy" cer"s) have heen applied to reassess the systematic affiliations between these bovids and related extinct Pleistocene forms. The fossil sam pies come from the sites of Elandsfontein (Cape Province) .nd Flarisb.d (Orange Free State) in South Afrie • . On the basis of differenees in skull morphology and size of the appendicular skeleton between fossil and modern blaek wildebeest (ConlJochaeus gnou). the subspecies name anliquus, proposed earlier to designate the Pleistoeene form, ean be retained. The same taxonomie level is accepted for the large Pleistocene e1and, whieh could be named Taurolragus oryx antiquus. The long horned or giant buffa1o, Pelorovis antiquus, can be inc1uded in the polymorphous Syncerus caffer stock and could therefore be called Syncerus caffer antiquus. The ecology of Pleistocene and modern Connochaetes, Taurolragus and Syncerus is discussed. A relationship between herbivore body size and c1imate, as Bergmann's Rule predicts, could not be demonstrated. -
Los Mamíferos Del Plioceno Y Pleistoceno De La Península Ibérica
94 investigación Los mamíferos del Plioceno y Pleistoceno de la Península Ibérica Bienvenido Martínez-Navarro | IPHES, Institut Català de Paleoecologia Humana i Evolució Social, Àrea de Prehistòria, Universitat Rovira i Virgili (URV), ICREA Sergio Ros-Montoya, María-Patrocinio Espigares | Dpto. de Ecología y Geología, Facultad de Ciencias, Universidad de Málaga Joan Madurell-Malapeira | Institut Català de Paleontologia Miquel Crusafont Paul Palmqvist | Dpto. de Ecología y Geología, Facultad de Ciencias, Universidad de Málaga URL de la contribución <www.iaph.es/revistaph/index.php/revistaph/article/view/4203> RESUMEN España es el país con mayor número de yacimientos bien conservados, cantidad y calidad de fósiles de todo el continente europeo. Aquí se describe el patrimonio paleobiológico correspondiente al Plio-Pleistoceno (últimos 5,3 millones de años) registrado en los principales yacimientos con presencia de fósiles de grandes mamíferos de la Península Ibérica. Ningún otro país de nuestro entorno al norte del Mediterráneo ofrece mayores posibilidades para el estudio y disfrute de los registros paleontológicos del Plioceno y Pleistoceno. En este contexto destacan las principales cuencas sedimentarias como las de Besalú-Bañolas, Vallés, Calatayud-Teruel, o ya en el sur las cuencas intrabéticas, donde merece especial interés la de Baza y Guadix, con localidades emblemáticas como Baza 1 para el Plioceno, y como los yacimientos de Orce (Fuente Nueva 1 y 3, Venta Micena o Barranco León, entre otros) para el Pleistoceno inferior. También se hace referencia a otro tipo de yacimientos, como son los maares volcánicos pliocénicos del Camp dels Ninots en Cataluña, o el de las Higueruelas en la Mancha, a las terrazas fósiles de los grandes ríos peninsulares, así como a los extraordinarios registros kársticos, donde destaca el de Atapuerca en Burgos. -
Bovidae (Mammalia) from the Lower Pliocene of Chad Denis Geraads, Cécile Blondel, Hassan Mackaye, Andossa Likius, Patrick Vignaud, Michel Brunet
Bovidae (Mammalia) from the lower Pliocene of Chad Denis Geraads, Cécile Blondel, Hassan Mackaye, Andossa Likius, Patrick Vignaud, Michel Brunet To cite this version: Denis Geraads, Cécile Blondel, Hassan Mackaye, Andossa Likius, Patrick Vignaud, et al.. Bovidae (Mammalia) from the lower Pliocene of Chad. Journal of Vertebrate Paleontology, Society of Verte- brate Paleontology, 2009, 29 (3), pp.923-933. 10.1671/039.029.0311. halshs-00433315 HAL Id: halshs-00433315 https://halshs.archives-ouvertes.fr/halshs-00433315 Submitted on 18 Nov 2009 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. Author manuscript, published in "Journal of Vertebrate Palaeontology 29, 3 (2009) 923-933" Bovidae (Mammalia) from the lower Pliocene of Chad DENIS GERAADS, *,1 CECILE BLONDEL, 2 HASSANE TAISSO MACKAYE, 3 ANDOSSA LIKIUS, 4 PATRICK VIGNAUD, 5 and MICHEL BRUNET 6 1 CNRS UPR 2147, 44 rue de l'Amiral Mouchez, 75014 Paris, France, [email protected]; 2 IPHEP, CNRS UMR 6046, Université de Poitiers, 40 Avenue du Recteur Pineau, 86022 Poitiers cedex, France, [email protected]; -
Dental Remains of Early Bison from the Tatrot Formation of the Upper Siwaliks, Pakistan
Khan et al., The Journal of Animal & Plant Sciences, 21(4): 2011, Page: J.86 Anim.2-867Plant Sci. 21(4):2011 ISSN: 1018-7081 DENTAL REMAINS OF EARLY BISON FROM THE TATROT FORMATION OF THE UPPER SIWALIKS, PAKISTAN M. A. Khan, S. Nasim, T. Ikram*, A. Ghafoor* and M. Akhtar*** Zoology Department, GC University, Faisalabad, Pakistan *Government College for Woman, Farooq Colony, Sargodha, Punjab, Pakistan **Zoology Department, Quaid-e-Azam Campus, Punjab University, Lahore, Pakistan ABSTRACT Dental fossil remains assigned to cf. Bison sivalensis are described and discussed. The recovered assemblages comprising 3 upper molars and one lower premolar reflect the morphological features of the genus Bison. The discovered material comes from the late Pliocene continental deposits of the Tatrot village (Tatrot Formation, Upper Siwaliks, northern Pakistan) dated approximately from 3.3 to 2.6 Ma. A new finding for the site documents the dentition of the early bison. Key words: Vertebrates, Mammals, Bovidae, Bovini, Bison, Siwaliks. INTRODUCTION The fossiliferous deposits of the Tatrot Formation outcropping in the area consist of pale pinkish- The studied specimens came from the deposits orange brown clays, brownish grey siltstones and shale, nearby Tatrot village, Jhelum district, northern Pakistan and greenish grey fine to medium grained sandstones (Fig. 1). The outcrops belong to the Tatrot Formation of intercalated with dark grey conglomerates (Khan et al., the Upper Siwaliks (Shah, 1980; Johnson et al., 1982). 2010). Hussain et al. (1992) and Barry et al. (2002) dated The Upper Siwaliks fluvial sequence of the Indian the lower boundary of the Tatrot Formation between 3.5- subcontinent is one of the most continuous of its age, 3.3 or 3.4-3.2 My, corresponding to the lower part of the spanning in time from the Late Pliocene up to the Middle Gauss magnetochron, whereas Kumaravel et al. -
The Evolution of Animal Weapons
The Evolution of Animal Weapons Douglas J. Emlen Division of Biological Sciences, The University of Montana, Missoula, Montana 59812; email: [email protected] Annu. Rev. Ecol. Evol. Syst. 2008. 39:387-413 Key Words First published online as a Review in Advance on animal diversity, sexual selection, male competition, horns, antlers, tusks September 2, 2008 The Annual Review of Ecology, Evolution, and Abstract Systematics is online at ecolsys.annualreviews.org Males in many species invest substantially in structures that are used in com- This article's doi: bat with rivals over access to females. These weapons can attain extreme 10.1146/annurev.ecolsys.39.110707.173 502 proportions and have diversified in form repeatedly. I review empirical lit- Copyright © 2008 by Annual Reviews. erature on the function and evolution of sexually selected weapons to clarify All rights reserved important unanswered questions for future research. Despite their many 1543-592X/08/1201-0387$20.00 shapes and sizes, and the multitude of habitats within which they function, animal weapons share many properties: They evolve when males are able to defend spatially restricted critical resources, they are typically the most variable morphological structures of these species, and this variation hon- estly reflects among-individual differences in body size or quality. What is not clear is how, or why, these weapons diverge in form. The potential for male competition to drive rapid divergence in weapon morphology remains one of the most exciting and understudied topics in sexual selection research today. 3*7 INTRODUCTION Sexual selection is credited with the evolution of nature's most extravagant structures, and these include showy male adornments that are attractive to females (ornaments) and an arsenal of outgrowths that function in male-male combat (weapons) (Darwin 1871). -
The Large Mammal Fauna of the Oldowayan Sites of Melka-Kunturé, Ethiopia. Denis Geraads, Vera Eisenmann, Germaine Petter
The Large Mammal Fauna of the Oldowayan sites of Melka-Kunturé, Ethiopia. Denis Geraads, Vera Eisenmann, Germaine Petter To cite this version: Denis Geraads, Vera Eisenmann, Germaine Petter. The Large Mammal Fauna of the Oldowayan sites of Melka-Kunturé, Ethiopia.. Istituto Italiano di Preistoria e Protoistoria. Studies on the early palaeolithic site of Melka Kunturé, Ethiopia, pp.169-192, 2004. halshs-00004899 HAL Id: halshs-00004899 https://halshs.archives-ouvertes.fr/halshs-00004899 Submitted on 10 Oct 2005 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. THE LARGE MAMMAL FAUNA OF THE OLDOWAYAN SITES OF MELKA-KUNTURÉ, ETHIOPIA Denis GERAADS *, Véra EISENMANN ** and Germaine PETTER ** * UPR 2147 CNRS - 44 rue de l'Amiral Mouchez - 75014 PARIS -FRANCE tel: 33 (0)1 43 13 56 21 fax 33 (0)1 43 13 56 30 [email protected] ** UMR 8569 CNRS - Institut de Paléontologie, 8 rue Buffon, 75005 PARIS - FRANCE RESUME La majeure partie des restes fauniques de Melka-Kunturé provient du site de Garba IV. Le site plus ancien de Gomboré I est moins riche, tandis que celui de Karré n'a livré que quelques fragments. -
Mohamed Sahnouni & Jan
stone age institute publication series Series Editors Kathy Schick and Nicholas Toth Stone Age Institute Gosport, Indiana and Indiana University, Bloomington, Indiana Number 1. THE OLDOWAN: Case Studies into the Earliest Stone Age Nicholas Toth and Kathy Schick, editors Number 2. BREATHING LIFE INTO FOSSILS: Taphonomic Studies in Honor of C.K. (Bob) Brain Travis Rayne Pickering, Kathy Schick, and Nicholas Toth, editors Number 3. THE CUTTING EDGE: New Approaches to the Archaeology of Human Origins Kathy Schick, and Nicholas Toth, editors Number 4. THE HUMAN BRAIN EVOLVING: Paleoneurological Studies in Honor of Ralph L. Holloway Douglas Broadfield, Michael Yuan, Kathy Schick and Nicholas Toth, editors STONE AGE INSTITUTE PUBLICATION SERIES NUMBER 3 Series Editors Kathy Schick and Nicholas Toth the cutting edge: New Approaches to the Archaeology of Human Origins Editors Kathy Schick Stone Age Institute & Indiana University Nicholas Toth Stone Age Institute & Indiana University Stone Age Institute Press · www.stoneageinstitute.org 1392 W. Dittemore Road · Gosport, IN 47433 COVER CAPTIONS AND CREDITS Top: Homo habilis Utilizing Stone Tools. Painting by artist-naturalist Jay H. Matternes. Copyright 1995, Jay H. Matternes. Inspired by a prehistoric scenario by K. Schick and N. Toth in Making Silent Stones Speak: Human Origins and the Dawn of Technology (1993), Simon and Schuster, New York. Pp.147-149. Lower right: Whole fl ake of trachyte lava from the 2.6 million-year-old site of Gona EG-10, Ethiopia. Reported by S. Semaw (2006), “The Oldest Stone Artifacts from Gona (2.6-2.5 Ma), Afar, Ethiopia: Implications for Understanding the Earliest Stages of Knapping” in The Oldowan: Case Studies into the Earliest Stone Age, eds. -
Pan-African Genetic Structure in the African Buffalo (Syncerus Caffer): Investigating Intraspecific Divergence
Pan-African Genetic Structure in the African Buffalo (Syncerus caffer): Investigating Intraspecific Divergence Nathalie Smitz1*,Ce´cile Berthouly2, Daniel Corne´lis2, Rasmus Heller3, Pim Van Hooft4, Philippe Chardonnet5, Alexandre Caron2,6,7, Herbert Prins8, Bettine Jansen van Vuuren9, Hans De Iongh10, Johan Michaux1,11 1 Departement of Life Sciences- Conservation Genetics, University of Lie`ge, Lie`ge, Belgium, 2 Centre de Coope´ration Internationale en Recherche Agronomique pour le De´veloppement (CIRAD), Campus International de Baillarguet, Montferrier-le-Lez, France, 3 Department of Biology- Bioinformatics, University of Copenhagen, Copenhagen, Denmark, 4 Resource Ecology Group, Wageningen University, Wageningen, The Netherlands, 5 International Foundation for the Conservation of Wildlife (IGF), Paris, France, 6 Department Environment and Societies- Centre de Coope´ration Internationale en Recherche Agronomique pour le De´veloppement (CIRAD), University of Zimbabwe, Harare, Zimbabwe, 7 Department of Zoology and Entomology- Mammal Research Institute, University of Pretoria, Pretoria, South Africa, 8 Tropical Nature Conservation and Vertebrate Ecology Group, Wageningen University, Wageningen, The Netherlands, 9 Department of Zoology- Centre for Invasion Biology, University of Johannesburg, Johannesburg, South Africa, 10 Institute of Environmental Sciences, Leiden University, Leiden, The Netherlands, 11 Centre de Biologie et de Gestion des Populations (CBGP), Campus International de Baillarguet, Montferrier-le-Lez, France Abstract The African buffalo (Syncerus caffer) exhibits extreme morphological variability, which has led to controversies about the validity and taxonomic status of the various recognized subspecies. The present study aims to clarify these by inferring the pan-African spatial distribution of genetic diversity, using a comprehensive set of mitochondrial D-loop sequences from across the entire range of the species. -
Bovini: Bovidae: Mammalia) Lower Case from the Pinjor Formation of Pakistan
Pakistan J. Zool., vol. 43(4), pp. 615-621, 2011. Proamphibos kashmiricus (Bovini: Bovidae: Mammalia) Lower Case From the Pinjor Formation of Pakistan Muhammad Akbar Khan1* and Muhammad Akhtar2 1Department of Zoology, Government College University, Faisalabad, Pakistan 2Palaeontology Laboratory, Department of Zoology, University of the Punjab, Lahore-54590, Pakistan Abstract. - Newly discovered fossil material of Proamphibos kashmiricus from the Plio-Pleistocene of the Pinjor Formation (2.6 – 0.6 Ma) in the Upper Siwaliks (Pakistan) is reported and described here. The new specimen is one opisthocranium of a hornless skull attributed to a female individual of P. kashmiricus coming from the Plio- Pleistocene deposits of Rohtas (2.2 Ma) in the Jhelum district, northern Pakistan. The opisthocranium is the largest female individual described so far for this species and the morphology of the present specimen allow examining some skull anatomical features of P. kashmiricus. The validity of the genus Bucapra is also discussed and the synonymy with Proamphibos is proposed. Keywords: Proamphibos, Pinjor Formation, Upper Siwaliks, Plio-Pleistocene, Bovine. INTRODUCTION (Rutimeyer, 1878; Pilgrim, 1937, 1939; Hooijer, 1958; Nanda, 2008). The unique characteristic of the Pinjor fauna, The late Pliocene and early Pleistocene is its similarity to the living faunas as it is the deposits of the Upper Siwaliks have produced youngest fauna of the Siwaliks and largely several fossil bovine taxa including Bison, Bubalus, consistent with an open grassland environment. Proamphibos, Leptobos, Bos, Hemibos and Regarding the bovid composition of the fauna, the Bucapra. This is a time of important radiation in forerunners of Boselaphus tragocamelus, Bubalus Bovini (Nanda, 1997, 2002; Masini 1989; Corvinus arnae, Bison bison, Bos taurus, Bibos banteng and and Rimal, 2001; Opdyke et al., 1979; Hussain et Antilope cervicapra are present in the youngest al., 1992; Pilgrim, 1937, 1939; Colbert, 1935) and sediments of the Siwaliks.