New Species, Combinations, and Records in Rubiaceae from the La Selva Biological Station, Costa Rica Charlotte M

Total Page:16

File Type:pdf, Size:1020Kb

New Species, Combinations, and Records in Rubiaceae from the La Selva Biological Station, Costa Rica Charlotte M Selbyana 12: 134-140 1991] TAYI NEW SPECIES, COMBINATIONS, AND RECORDS IN RUBIACEAE FROM THE LA SELVA BIOLOGICAL STATION, COSTA RICA CHARLOTTE M. TAYLORS BARRY E. HAMMEL Missouri Botanical Garden, P.O. Box 299, St. Louis, Missouri 63166-0299 WILLIAM C. BURGER Department of Botany, Field Museum of Natural History, Chicago, Illinois 60605 ABSTRACT. The new species Coussarea nigrescens C. M. Taylor & Hammel, Coussarea psychotrioides C. M. Taylor & Hammel, and Hillia grayumii C. M. Taylor are described. The new combinations Chione sylvicola (Standley) W. Burger, Coussarea hondensis (Standley) C. M. Taylor & W. Burger, Psychotria camponutans (Dwyer & Hayden) Hammel, Psychotria elata (Swartz) Hammel, and Psychotria guapilensis (Standley) Hammel are made. The species Borojoa panamensis Dwyer, Faramea parvibractea Steyermark, and Lasianthus panamensis (Dwyer) Robbrecht are reported from Costa Rica for the first time. The La Selva Biological Station is situated in near the junction of the Rios Puerto Viejo and wet forest in the Caribbean lowlands of north- Sarapiqui, 100m, 5 Nov 1980,5. Hammel 10383 eastern Costa Rica (Frankie et al., 1974; Hammel (DUKE!, holotype). & Grayum, 1982; Wilbur, 1986). The following Puberulent trees to 7 m tall, drying hlack. new species descriptions and nomenclatural Leaves decussate; blades elliptic, 8-20.5 cm long, combinations result from the study of the Ru- 3.5-9 cm wide, acute to slightly acuminate at biaceae for the Vascular Flora of the La Selva apex, cuneate at base, thinly chartaceous; sec- Biological Station (Wilbur, 1986), and the con- ondary veins 9-11 pairs, without domatia; pet- current study of this family in Costa Rica for the ioles 10-35 mm long; stipules interpetiolarly and Flora Costaricensis and the Manual Flora of Cos- intrapetiolarly connate, persistent, triangular, 1.5- ta Rica. Keys, descriptions, and illustrations of 2 mm long, shallowly bilobed. Inflorescences ter- the stipules for all species of Rubiaceae found at minal, corymbiform to pyramidal, 1-2 cm long the La Selva Station will be presented in the and wide; peduncles ca. 2 cm long; bracts tri- Vascular Flora. Several other new species of Ru- angular to ovate, 1-3 cm long, caducous; flowers biaceae known from the La Selva Biological Sta- sessile in glomerules of 3-7, bisexual, 5-merous, tion will be described by Dwyer and Lorence in calyx minutely puberulent, the limb 0.5 mm long, the Flora Mesoamericana. broadly and shallowly lobed; corolla salverform, white, glabrous, the tube 22-24 mm long, the FIGURE 1. A, Coussarea ni; New Species lobes lingulate, 6-8 mm long. Fruit globose to ellipsoid, ca. 1 cm in diameter, black; pyrenes Coussarea nigrescens C. M. Taylor & Hammel, 1-2, globose to ellipsoid or planoconvex, PHENOLOGY. Collected in fl sp. nov. FIGURE 1. smooth. and September to November Arbor ad 7 m alta, puberula, exsiccata nigra. ADDITIONAL SPECIMENS EXAMINED. Costa Rica. HERE- in February. Folia petiolata oppositaque; laminis ellipticis 8- DIA: Canton de Sarapiqui, 2 km S of Magsaysay Penal Colony, N. Garwood et al. 1129 (CR). LIMON: Canton HABITAT AND DISTRIBUTION. 14 x 3.5-6 cm; stipulis persistentibus, truncatis de Limon, Cerro Muchilla, Fila Matona, 9°46'50"N and moist forest at 20-850 rr ad parum bilobatis 1.5-2 mm longis. Inflores- 83°05'30"W, R. Robles & A. Chacon 2667 (CR). and the Osa Peninsula of Co: centia rotundata usque pyramidalis 1-2 x 1-2 PUNTARENAS: Canton de Osa, Reserva Forestal Golfo cm, pedunculo ca. 2 cm longo, flores sessilis Dulce, Rio Tigre, 8°31'40"N 83°26'30"W, B. Hammel RELATIONSHIPS. This new si glomerulatis, calyce 0.5 mm longo, late lobulato; et al. 17851 (CR); Parque Nacional Corcovado, Rio rated from other Costa Ricai corolla tubulosa, ubique glabra, tubo 22-24 mm Sirena, 8°28'N 83°35'W, C. Kernan 1328 (CR), 8°31'N sarea by its leaves, which ha- longo, lobis 6-8 mm longis. 83°31 'W, R. Liesner 3084 (CR). PUNTARENAS-SAN JOSE when fresh, and turn black whe BORDER REGION: Tarrazu, faldas de Cerro Nara cerca color is alluded to in the specifi TYPE. Costa Rica, Heredia. Finca La Selva, the Esquipulas, 9°29'N 84°03'W, J. Gomez Laurito et al. the flowers are distylous or mi OTS field station near Puerto Viejo de Sarapiqui, 11594 (CR, DUKE, MO, usj). known. 134 1991] TAYLOR ET AL.: NEW RUBIACEAE OF LA SELVA 135 tural History, Ol Hammel, Coussarea psychotrioides bed. The new combinations Chione n 1. Taylor & W. Burger, Psychotria lamrnel, and Psychotria guapilensis , Faramea parvibractea Steyermark, 3 sta Rica for the first time. ;ion of the Rios Puerto Viejo and I m, 5 Nov 1980, B. Hammel 10383 . trees to 7 m tall, drying black. ate; blades elliptic, 8-20.5 cm long, ie, acute to slightly acuminate at : at base, thinly chartaceous; sec- 9-11 pairs, without domatia; pet- im long; stipules interpetiolarly and f connate, persistent, triangular, 1 . 5- lallowly bilobed. Inflorescences ter- ibiform to pyramidal, 1-2 cm long iduncles ca. 2 cm long; bracts tri- ate, 1-3 cm long, caducous; flowers nerules of 3-7, bisexual, 5-merous, .y pubemlent, the limb 0.5 mm long, ihallowly lobed; corolla salverform, >us, the tube 22-24 mm long, the FIGURE 1. A, Coussarea nigrescens, habit with inflorescence. B, flower. From Hammel 10383 (DUKE). te, 6-8 mm long. Fruit globose to 1 cm in diameter, black; pyrenes ;e to ellipsoid or planoconvex, PHENOLOGY. Collected in flower in April, July Coussarea psychotrioides C. M. Taylor & Ham- and September to November and in young fruit mel, sp. nov. FIGURE 2. in February. 'ECIMENS EXAMINED. Costa Rica. HERE- Arbor usque ad 12 m alta, glabrescens. Folia e Sarapiqui, 2 km S of Magsaysay Penal HABITAT AND DISTRIBUTION. Infrequent in wet irwood et al. 1129 (CR). LIMON: Canton petiolata oppositaque; laminis ellipticis, 8-18 x ;rro Muchilla, Fila Matona, 9°46'50"N and moist forest at 20-850 m. Caribbean slopes 2.5-6.5 cm; stipulis persistentibus, truncatis ad R. Robles & A. Chacon 2667 (CR). and the Osa Peninsula of Costa Rica. parum bilobatis, 0.5-1 mm longis. Inflorescentia Canton de Osa, Reserva Forestal Golfo RELATIONSHIPS. This new species can be sepa- rotundata usque pyramidalis, 3-7 x 5-8 cm; pe- gre, 8°31'40"N 83°26'30"W, B. Hammel rated from other Costa Rican species of Cous- dunculo 1-3 cm longo. Flores sessilis glomeru- (CR); Parque Nacional Corcovado, Rio sarea by its leaves, which have a velvety sheen latis; calyce 0.3 mm longo, subtruncato; corolla si 83°35'W, C. Kernan 1328 (CR), 8°31'N hypocrateriformi, ubique glabra, tubo 12-15 mm 'Jesner 3084 (CR). PUNTARENAS-SAN JOSE when fresh, and turn black when dried. This dried color is alluded to in the specific epithet. Whether longo, lobis 10-12 mm longis. Fructus ellipsoi- ON: Tarrazu, faldas de Cerro Nara cerca deus, 1.5-2 cm longus. '°29'N 84°03'W, /. Gomez Laurito et al. the flowers are distylous or monomorphic is un- UKE, MO, usj). known. TYPE. Costa Rica, Heredia. Finca La Selva, the.
Recommended publications
  • 2011 Vol. 14, Issue 3
    Department of Botany & the U.S. National Herbarium The Plant Press New Series - Vol. 14 - No. 3 July-September 2011 Island Explorations and Evolutionary Investigations By Vinita Gowda or over a century the Caribbean eastward after the Aves Ridge was formed On joining the graduate program region, held between North and to the West. Although the Lesser Antilles at The George Washington University FSouth America, has been an active is commonly referred to as a volcani- in Washington, D.C., in the Fall of area of research for people with interests cally active chain of islands, not all of the 2002, I decided to investigate adapta- in island biogeography, character evolu- Lesser Antilles is volcanic. Based on geo- tion in plant-pollinator interactions tion, speciation, as well as geology. Most logical origin and elevation all the islands using a ‘multi-island’ comparative research have invoked both dispersal and of the Lesser Antilles can be divided into approach using the Caribbean Heliconia- vicariance processes to explain the distri- two groups: a) Limestone Caribbees (outer hummingbird interactions as the study bution of the local flora and fauna, while arc: calcareous islands with a low relief, system. Since I was interested in under- ecological interactions such as niche dating to middle Eocene to Pleistocene), standing factors that could influence partitioning and ecological adaptations and b) Volcanic Caribbees (inner arc: plant-pollinator mutualistic interactions have been used to explain the diversity young volcanic islands with strong relief, between the geographically distinct within the Caribbean region. One of dating back to late Miocene). islands, I chose three strategic islands of the biggest challenges in understanding the Lesser Antilles: St.
    [Show full text]
  • Roy Emile Gereau, Born 5 December L947, Rock Island, Illinois, U.S.A
    CURRICULUM VITAE Personal: Roy Emile Gereau, born 5 December l947, Rock Island, Illinois, U.S.A. Education and Degrees: B.A., mathematics and French, University of Iowa, l969 (graduation with Highest Distinction); B.S., forestry, Michigan Technological University, l975 (graduation with Highest Distinction); M.S., biological sciences, Michigan Technological University, l978 (graduation with Highest Distinction); enrolled in Ph.D. program at Michigan State University, 1978, successfully completed all course work, 1981. Professional and Scholastic Positions: Teaching Assistant, Department of Mathematical and Computer Sciences, Michigan Technological University, l975-l977; Instructor, Department of Mathematical and Computer Sciences, Michigan Technological University, l977-l978; Instructor, Department of Biological Sciences, Michigan Technological University, summer terms l977 and l978 (vascular plant taxonomy); Teaching Assistant, Department of Botany and Plant Pathology, Michigan State University, spring terms l979 and l98l; Herbarium Assistant, Beal-Darlington Herbarium, Department of Botany and Plant Pathology, Michigan State University, l978-l983; Curatorial Assistant, Missouri Botanical Garden, St. Louis, Missouri, l983-2005; Assistant Curator, Missouri Botanical Garden, St. Louis, Missouri, April 2005-present. Editorial Positions: Consulting Editor, Novon; Member of Editorial Committee, Annals of the Missouri Botanical Garden; Member of Reviewer Panel, African Journal of Ecology Awards: Sigma Xi Grant-in-Aid of Research, l979; National
    [Show full text]
  • Glycosides in the Rubiaceae*
    The occurrence of asperulosidic glycosides in the Rubiaceae* P. Kooiman Laboratorium voor Algemene en Technische Biologie Technische Hogeschool, Delft. SUMMARY Some properties of the new iridoid compounds Galium glucoside and Gardenia glucoside are described. Galium glucoside and asperuloside occurin many species belongingto the Rubioideae (sensu Bremekamp); they were not found in other subfamilies of the Rubiaceae. Gardenia glucoside occurs in several species ofthe tribe Gardenieae (subfamily Ixoroideae). The distribution of the asperulosidic glucosides in the Rubiaceae corresponds with the classi- fication proposed by Bremekamp, although there are some exceptions (Hamelieae, Opercu- laria and Pomax, possibly the Gaertnereae). To a somewhat less degreethe system proposedby Verdcourt is supported. 1. INTRODUCTION Apart from the classification arrived at by Bremekamp (1966) the only other modern system of the Rubiaceae was proposed by Verdcourt (1958); both au- thors considered their classifications tentative. The have several fea- as systems tures in common, but deviate in some points. The main differences are in the po- sition ofthe Urophylloideae sensu Bremekamp, which are included in the subfa- mily Rubioideaeby Verdcourt, and in the relationship between the Cinchonoideae the Ixoroideae and (both sensu Bremekamp) which are united in the subfamily Cinchonoideae by Verdcourt. Both systems diverge widely and principally from all older classifications which appeared to become more and more unsatis- factory as the number of described species increased. In 1954 Briggs & Nicholes reported on the presence or absence of the iridoid glucoside asperuloside (1) in most species of Coprosma and in many other Rubiaceae. The reaction they used for the detection of asperuloside is now known to be not specific for this glucoside; it detects in addition some struc- turally and most probably biogenetically related glycosides.
    [Show full text]
  • Common Epiphytes and Lithophytes of BELIZE 1 Bruce K
    Common Epiphytes and Lithophytes of BELIZE 1 Bruce K. Holst, Sally Chambers, Elizabeth Gandy & Marilynn Shelley1 David Amaya, Ella Baron, Marvin Paredes, Pascual Garcia & Sayuri Tzul2 1Marie Selby Botanical Gardens, 2 Ian Anderson’s Caves Branch Botanical Garden © Marie Selby Bot. Gard. ([email protected]), Ian Anderson’s Caves Branch Bot. Gard. ([email protected]). Photos by David Amaya (DA), Ella Baron (EB), Sally Chambers (SC), Wade Coller (WC), Pascual Garcia (PG), Elizabeth Gandy (EG), Bruce Holst (BH), Elma Kay (EK), Elizabeth Mallory (EM), Jan Meerman (JM), Marvin Paredes (MP), Dan Perales (DP), Phil Nelson (PN), David Troxell (DT) Support from the Marie Selby Botanical Gardens, Ian Anderson’s Caves Branch Jungle Lodge, and many more listed in the Acknowledgments [fieldguides.fieldmuseum.org] [1179] version 1 11/2019 TABLE OF CONTENTS long the eastern slopes of the Andes and in Brazil’s Atlantic P. 1 ............. Epiphyte Overview Forest biome. In these places where conditions are favorable, epiphytes account for up to half of the total vascular plant P. 2 .............. Epiphyte Adaptive Strategies species. Worldwide, epiphytes account for nearly 10 percent P. 3 ............. Overview of major epiphytic plant families of all vascular plant species. Epiphytism (the ability to grow P. 6 .............. Lesser known epiphytic plant families as an epiphyte) has arisen many times in the plant kingdom P. 7 ............. Common epiphytic plant families and species around the world. (Pteridophytes, p. 7; Araceae, p. 9; Bromeliaceae, p. In Belize, epiphytes are represented by 34 vascular plant 11; Cactaceae, p. 15; p. Gesneriaceae, p. 17; Orchida- families which grow abundantly in many shrublands and for- ceae, p.
    [Show full text]
  • Survey of Rubiaceae in the Golfo Dulce Area, Costa Rica
    ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Stapfia Jahr/Year: 2008 Band/Volume: 0088 Autor(en)/Author(s): Will Sabine, Kiehn Michael Artikel/Article: Survey of Rubiaceae in the Golfo Dulce area, Costa Rica: new species, combinations and name changes since 2001 111-120 © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at Survey of Rubiaceae in the Golfo Dulce area, Costa Rica: new species, combinations and name changes since 2001 Investigaciones en Rubiaceae en el área de Golfo Dulce, Costa Rica: nuevas especies, combinaciones y cambios de nombre desde 2001 S abine W ILL &Michael K IEHN Abstract: Since the publication of the last Rubiaceae treatment of the Golfo Dulce area in southern Costa Rica (WILL et al. 2001a), several new taxa have been described and changes in generic and species level within the family have been made. The new species, combinations and name changes are presented here and a new checklist of the Rubiaceae for the region is provided. Key words: Costa Rica, Golfo Dulce, Rubiaceae, checklist. Resumen: Desde la última publicación de Rubiaceas del área del Golfo Dulce en la zona Sur de Costa Rica (WILL et al. 2001a), nuevos taxa han sido descritos y también se han realizado cambios a nivel génerico y específico dentro de la familia. En esta re- visión se presentan las nuevas especies, combinaciones y cambios de nombre, así como la lista actualizada de las Rubiaceas de la región. Palabras clave: Costa Rica, Golfo Dulce, Rubiaceae, lista actualizada. Introduction green forests and wet rainforests, up to dry forests and desert environments.
    [Show full text]
  • Floral Fragrances. a Day in the Life of a Linalool Molecule
    Plant Species Biology (1999) 14, 95Ð120 NEW PERSPECTIVES IN POLLINATION BIOLOGY: FLORAL FRAGRANCES A day in the life of a linalool molecule: Chemical communication in a plant-pollinator system. Part 1: Linalool biosynthesis in ßowering plants ROBERT A. RAGUSO* and ERAN PICHERSKY *Department of Biological Sciences, University of South Carolina, Columbia, SC 29208, USA. Department of Biology, University of Michigan, Ann Arbor, MI 48109Ð1048, USA Abstract The monoterpene alcohol, linalool, is present in the ßoral fragrance of diverse plant fami- lies and is attractive to a broad spectrum of pollinators, herbivores and parasitoids. Floral emission of linalool has evolved de novo in the fragrant, moth-pollinated annual Clarkia breweri (Gray) Greene (Onagraceae) through a combination of up-regulation and ectopic expression of its biosynthetic enzyme, linalool synthase (LIS), in conjunction with allo- metric size increases in all ßoral organs. Linalool synthase activity and linalool emissions are 1000-fold lower in a sibling species, C. concinna (Fischer & Meyer) Greene, that is diurnally pollinated. Linalool synthase expression is spatially and temporally regulated during C. breweri ßower development, immediately precedes free linalool emission and is absent from nonßoral tissues. Its activity is highest in the style, but most of the linalool product appears to be converted to the pyranoid and furanoid linalool oxides. The LIS structural gene is a member of the terpene synthase gene family, sharing sequence iden- tity with two discrete classes, represented by limonene synthase (LMS) and copalyl pyrophosphate synthase (CPS). Genetic crosses between C. breweri and C. concinna indi- cate that strong linalool emission segregates as a dominant mendelian trait, whereas the inheritance of linalool oxide formation is more complex, suggesting epistatic biosynthetic pathway interactions.
    [Show full text]
  • A SURVEY of the SYSTEMATIC WOOD ANATOMY of the RUBIACEAE by Steven Jansen1, Elmar Robbrecht2, Hans Beeckman3 & Erik Smets1
    IAWA Journal, Vol. 23 (1), 2002: 1–67 A SURVEY OF THE SYSTEMATIC WOOD ANATOMY OF THE RUBIACEAE by Steven Jansen1, Elmar Robbrecht2, Hans Beeckman3 & Erik Smets1 SUMMARY Recent insight in the phylogeny of the Rubiaceae, mainly based on macromolecular data, agrees better with wood anatomical diversity patterns than previous subdivisions of the family. The two main types of secondary xylem that occur in Rubiaceae show general consistency in their distribution within clades. Wood anatomical characters, espe- cially the fibre type and axial parenchyma distribution, have indeed good taxonomic value in the family. Nevertheless, the application of wood anatomical data in Rubiaceae is more useful in confirming or negating already proposed relationships rather than postulating new affinities for problematic taxa. The wood characterised by fibre-tracheids (type I) is most common, while type II with septate libriform fibres is restricted to some tribes in all three subfamilies. Mineral inclusions in wood also provide valuable information with respect to systematic re- lationships. Key words: Rubiaceae, systematic wood anatomy, classification, phylo- geny, mineral inclusions INTRODUCTION The systematic wood anatomy of the Rubiaceae has recently been investigated by us and has already resulted in contributions on several subgroups of the family (Jansen et al. 1996, 1997a, b, 1999, 2001; Lens et al. 2000). The present contribution aims to extend the wood anatomical observations to the entire family, surveying the second- ary xylem of all woody tribes on the basis of literature data and original observations. Although Koek-Noorman contributed a series of wood anatomical studies to the Rubiaceae in the 1970ʼs, there are two principal reasons to present a new and com- prehensive overview on the wood anatomical variation.
    [Show full text]
  • Historical Biogeography of the Predominantly Neotropical
    Historical Biogeography of the Predominantly Neotropical Subfamily Cinchonoideae (Rubiaceae): Into or Out of America? Author(s): Ulrika Manns, Niklas Wikström, Charlotte M. Taylor, Birgitta Bremer Reviewed work(s): Source: International Journal of Plant Sciences, Vol. 173, No. 3 (March 2012), pp. 261-286 Published by: The University of Chicago Press Stable URL: http://www.jstor.org/stable/10.1086/663971 . Accessed: 02/05/2012 04:59 Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at . http://www.jstor.org/page/info/about/policies/terms.jsp JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact [email protected]. The University of Chicago Press is collaborating with JSTOR to digitize, preserve and extend access to International Journal of Plant Sciences. http://www.jstor.org Int. J. Plant Sci. 173(3):261–289. 2012. Ó 2012 by The University of Chicago. All rights reserved. 1058-5893/2012/17303-0004$15.00 DOI: 10.1086/663971 HISTORICAL BIOGEOGRAPHY OF THE PREDOMINANTLY NEOTROPICAL SUBFAMILY CINCHONOIDEAE (RUBIACEAE): INTO OR OUT OF AMERICA? Ulrika Manns,1,* Niklas Wikstro¨m,* Charlotte M. Taylor,y and Birgitta Bremer* *Bergius Foundation, Royal Swedish Academy of Sciences and Department of Botany, Stockholm University, SE-10691 Stockholm, Sweden; and yMissouri Botanical Garden, P.O. Box 299, Saint Louis, Missouri 63166, U.S.A.
    [Show full text]
  • Beltrami Island Land Utilization Project Comprehensive Conservation Management Plan Draft, 2012
    Beltrami Island Land Utilization Project Comprehensive Conservation Management Plan Draft, 2012 1 Beltrami Island Land Utilization Project Comprehensive Conservation Management Plan Draft, 2011 Minnesota Department of Natural Resources U.S. Fish and Wildlife Service 2 Acknowledgements This plan is a product of the Minnesota Department of Natural Resources. Some sections of this plan, particularly where it references other plans, borrows language directly from those plans, including other Minnesota DNR documents, the Red Lake Bands’ Wolf Management Plan, and various Watershed District and watershed basin plans. Much of the material presented in Chapter 3 on soils, peatlands, and vegetation communities was contributed and written by Scott Zager of Wildlands Inc., 2009 Maryknoll Avenue North, Maplewood, MN. The section on insect resources was provided by Kyle Johnson, University of Wisconsin, Madison. Materials, guidance and edits were provided by members of the planning Leadership Team (see Appendix E) and other DNR staff including Jana Albers, Wes Bailey, Heidi Cyr, Margaret Dexter, Katie Haws, Steve Heutmaker, Chris Kotter-Weir, Mike Kovacovich, Mike Larson, Cindy Leuth, Mike Magner, Becky Marty, Cynthia Osmundson, Dave Radford,, Chris Scharenbroich, Dave Schiller, Dennis Simon, Phil Talmadge, and Dave Thomas. Pete Takash set up the project website and assisted with production and distribution of news press releases. Lead author: Michael R. North. Cover art work by Ross Hier, copyright 2011. “We and our families have enjoyed the Beltrami Forest area for many years. Its become an important part of our lives. My grandparents took us hunting, fishing, trapping, berrying all our lives, and it continues [for] our grandchildren and great grandchildren.
    [Show full text]
  • An Updated Phylogenetic Classification of the Flowering Plants Robert F
    Aliso: A Journal of Systematic and Evolutionary Botany Volume 13 | Issue 2 Article 8 1992 An Updated Phylogenetic Classification of the Flowering Plants Robert F. Thorne Rancho Santa Ana Botanic Garden Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons Recommended Citation Thorne, Robert F. (1992) "An Updated Phylogenetic Classification of the Flowering Plants," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 13: Iss. 2, Article 8. Available at: http://scholarship.claremont.edu/aliso/vol13/iss2/8 ALISO ALISO 13(2), 1992, pp. 365-389 ~Amer. Acad. AN UPDATED PHYLOGENETIC CLASSIFICATION OF THE FLOWERING PLANTS : Amer. Acad. I ~r. Acad. Arts ROBERT F. THORNE Mem. Amer. Rancho Santa Ana Botanic Garden Claremont, California 91711 ABSTRACT This update of my classification of the flowering plants, or Angiospermae, is based upon about 800 pertinent books, monographs, and other botanical papers published since my last synopsis appeared in the Nordic Journal of Science in 1983. Also I have narrowed my family- and ordinal-gap concepts to bring acceptance of family and ordinal limits more in line with those of current taxonomists. This new information and the shift in my phylogenetic philosophy have caused significant changes in my interpretation of relationships and numbers and content of taxa. Also the ending "-anae" has been accepted for superorders in place in the traditional but inappropriate" -iflorae." A new phyletic "shrub" replaces earlier versions, and attempts to indicate relationships among the superorders, orders, and suborders. One table includes a statistical summary of flowering-plant taxa: ca. 235,000 species of 12,615 genera, 440 families, and 711 subfamilies and undivided families in 28 superorders, 70 orders, and 7 5 suborders of Angiospermae.
    [Show full text]
  • Amazon Plant Diversity Revealed by a Taxonomically Verified Species List
    Amazon plant diversity revealed by a taxonomically verified species list Domingos Cardosoa,1,2, Tiina Särkinenb,1, Sara Alexanderc, André M. Amorimd, Volker Bittriche, Marcela Celisf,g, Douglas C. Dalyh, Pedro Fiaschii, Vicki A. Funkc, Leandro L. Giacominj, Renato Goldenbergk, Gustavo Heidenl, João Igancim, Carol L. Kelloffc, Sandra Knappn, Haroldo Cavalcante de Limao, Anderson F. P. Machadop, Rubens Manoel dos Santosq, Renato Mello-Silvar, Fabián A. Michelangelih, John Mitchellh, Peter Moonlightb, Pedro Luís Rodrigues de Moraess, Scott A. Morih, Teonildes Sacramento Nunesp, Terry D. Penningtont, José Rubens Piranir, Ghillean T. Prancet, Luciano Paganucci de Queirozp, Alessandro Rapinip, Ricarda Riinau, Carlos Alberto Vargas Rinconv, Nádia Roquea, Gustavo Shimizuw, Marcos Sobralx, João Renato Stehmanny, Warren D. Stevensz, Charlotte M. Taylorz, Marcelo Trovóaa, Cássio van den Bergp, Henk van der Werffz, Pedro Lage Vianabb, Charles E. Zartmancc, and Rafaela Campostrini Forzzao aNational Institute of Science and Technology in Interdisciplinary and Transdisciplinary Studies in Ecology and Evolution (INCT IN-TREE), Instituto de Biologia, Universidade Federal da Bahia, 40170-115 Salvador, BA, Brazil; bRoyal Botanic Garden Edinburgh, Edinburgh EH5 3LR, United Kingdom; cNational Museum of Natural History, Smithsonian Institution, Washington, DC 20560-0163; dDepartamento de Ciências Biológicas, Universidade Estadual de Santa Cruz, 45662-900 Ilhéus, BA, Brazil; ePrivate address, 13083-290 Campinas, SP, Brazil; fDepartamento de Química y Biología,
    [Show full text]
  • University of Minneso
    / I ,J UNIVERSITY OF MINNESO ./ L ': BULLETIN NO. 43. DIVISION OF ENTOMOLOGY. --- --, Decer.n.ber, 1995. INSECTS IN.JURIOUS IN 1895. ST. ANTHONY PARIC, RAMSEY COUNTY, MINNESOTA. ·' H.\RinSON & S~IITH, PRINTERS, MINNEAPOLIS. UNIVERSITY OF MINNESOTA. BOARD OF REGENTS. The HON. JOHN S. PILLSBURY, MINNEAPOLIS, Regent for Li,fe The HON. GREENLEAF CLARK, M.A., ST. PAUL,­ 1900 The HON. CUSHMAN K. DAVIS, M. A., ST. PAUL, - 1900 The HON. WM. H. YALE, WINONA, 1896 The HON. JOEL P. HEATWOLE, NORTHFIELD, 1897 The HON. L. S. SWENSON, ALBERT LEA, 1897 The HON. ALPHONSO BARTO, ST. CLOUD, 1896 The HON. WM. M. LIGGETT, BENSON, 1897 The HON. S. M. OWEN, MINNEAPOLIS, 1901 The HON. STEPHEN MAHONEY, B. A., MINNEAPOLIS, 1901 The HON. DAVID M. CLOUGH, MINNEAPOLIS, Ex-Officio The Governor of the State. The HON. W. W. PENDERGAST, M. A., HUTCHINSON, - Ex-Officio The State Superintendent of Public Instruction. CYRUS NORTHROP, LL. D., MINNEAPOLIS, - - Ex-Ojficio The President of the University. TIIE AGIUCULTURAL COJ\Il\IITTEE. The HON. WILLIAM M. LIGGETT, Chairman. The HON. J. S. PILLSBURY. The RON. 8. M. OWEN. The HON. W. W. PENDERGAST. OFFICERS OF THE STA'.fION: WM. M. LIGGETT, - Chairman. WILLET M. HAYS, B. S. A., Vice Chairman and Agriculturist. SAMUEL B. GREEN, B. S., - Horticulturist. OTTO LUGGER, Ph.D., - Entomologist and Botanist. HARRY SNYDER, B. S., Chemist. T. L. H1ECKER, - Dairy Husbandry. M. H. REYNOLDS, M. D , V. M., - Veterinarian. TROS. SHAW, Animal Husbandry. T. A. HOVERSTAD, Assistant in Agriculture, at Crookston . .T. A. VYE. - Secretary. ANDREW BOSS, Farm Foreman. If@" The Bulletins of this Station are mailed free to all residents of the state who make application for them.
    [Show full text]