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The Academy Is Governed by a Distinguished Group of Eminent The Academy is governed by a distinguished group of eminent scientists who have been appointed Fellows of the California Academy of Sciences in recognition of their notable contributions to one or more of the natural sciences. Nominated by their colleagues and appointed by the Board of Trustees, the Fellows remain members of the Fellowship for life. Below is a complete list of Academy Fellows. David Ackerly Alissa J. Arp Shannon N. Bennett Ecology and Evolutionary Biology Marine Biology Microbiology Bruce M. Alberts John C. Avise William J. Bennetta Molecular and Conservation Biology Genetics and Ecology Science and Public Policy Angel C. Alcala Francisco J. Ayala Robert G. Bergman Marine Biology and Conservation, Genetics Chemistry Herpetology Bruce G. Baldwin Brent Berlin Michele L. Aldrich Botany Anthropology History of Science George E. Ball Giacomo Bernardi Zeresenay Alemseged Entomology Ichthyology Anthropology and Paleoanthropology Richard T. Barber Vladimir A. Beshkov Frank Almeda Marine Biology Herpetology Botany Anthony D. Barnosky Devaki Bhaya Walter Alvarez Paleontology Molecular Biology Geology Reginald H. Barrett Philip E. Bickler Steven C. Anderson Applied Biology, Agricultural and Comparative Physiology, Systematic, Biogeographic and Environmental Science Neuroscience and Anesthesiology Ecological Studies in Herpetology John Barron James L. Bischoff Christopher Andrews Micropaleontology Marine Geology Aquarium Management and Parasitology Bruce Bartholomew Jeffrey M. Black Botany Conservation and Ecology of Water Fred Andrews Fowl Entomology Gibor B. Basri Astronomy and Astrophysics David C. Blackburn Paul H. Arnaud Herpetology Entomology Natalie Batalha Astronomy Elizabeth H. Blackburn E. N. Arnold Molecular Biology Herpetology Aaron M. Bauer Herpetology James A. Blake Jeanne Arnold Invertebrate Zoology Anthropology Steven V. Beckwith Astronomy Page 1 of 11 Academy Fellows Barbara A. Block Robert W. Buddemeier Mark H. Carr Evolutionary, Cellular and Molecular Environmental Studies Marine Ecology Physiology Stephen H. Bullock John Caskey Carol L. Boggs Plant Ecology Geology, Neotectonics Evolutionary Ecology Margaret G. Burke Alan Channing Stephen Bollens Behavioral Ecology and Education Herpetology Biological Oceanography Nathalie A. Cabrol Nona Chiariello Penelope Boston Astrobiology Global Ecology Astrobiology Gregor M. Cailliet Robert L. Christiansen Sarane T. Bowen Ichthyology Geology Genetics Roy L. Caldwell Christopher F. Chyba Rauri C.K. Bowie Invertebrate Zoology Astrobiology Ornithology Douglas G. Capone Frank Cipriano Justin Brashares Biological Oceanography, Marine Ecology and Evolutionary Biology Ecology and Wildlife Conservation Microbiology David A. Clague John I. Brauman Ring Cardé Volcanology Chemistry Entomology William A. Clemens Thomas S. Briggs James R. Carey Paleontology Entomology Entomology Lloyd S. Cluff Winslow R. Briggs Sherwin Carlquist Geology Botany Botany Kenneth Coale Adriana Briscoe Sandra J. Carlson Oceanography Evolutionary Biology Paleontology William P. Cochlan Michael E. Brown James T. Carlton Biological Oceanography Astronomy Marine Biodiversity and Conservation Anne C. Cohen Robert D. Brown, Jr. Edward J. Carpenter Invertebrate Systematics Geology Biological Oceanography and Marine Microbiology Daniel M. Cohen Thomas D. Bruns Ichthyology Mycology James M. Carpenter Entomology Sarah Cohen Richard C. Brusca Marine Genetics Marine Biology and Conservation Kent E. Carpenter Marine Biology and Ichthyology Page 2 of 11 Academy Fellows James P. Collman Marymeghan Daly Carmen Domingo Chemistry Animal Systematics and Ecology, Science Education Cnidaria Robert K. Colwell Robert V. Dowell Zoology Thomas F. Daniel Entomology Botany Lynn Cominsky John T. Doyen Cosmology Bruno David Entomology Paleontology, Marine Biology, Margaret W. Conkey Evolution James A. Doyle Anthropology Botany Todd E. Dawson Bruce Conklin Physiological Plant Ecology William T. Doyle Bioinformatics, Genomics and Botany Molecular Pharmacology Donald J. DePaolo Geosciences Frank Drake Edward F. Connor Astronomy Ecology Ruth DeFries Landscape Ecology Robert C. Drewes T. J. Conomos Herpetology Marine Geology Joseph DeRisi Molecular Biology, Genomics, Richard Dugdale Claudia Corti Parasitology, Virology, Oceanography Herpetology Computational Biology John P. Dumbacher Lawrence Cory David Des Marais Ornithology Zoology Geology and Astrobiology Thomas O. Duncan Daniel P. Costa Dennis E. Desjardin Botany Vertebrate Ecology Mycology Thomas Dunne Daniel E. Crocker Susan P. Desmond-Hellmann Geomorphology and Hydrology Ecology and Physiology Oncology and Biotechnology William H. Durham Donald Croll Marian C. Diamond Human Biology Ecology of Marine Predators, Island Anatomy Ecology and Conservation David M. Eakin Janis Dickinson Physics Dale P. Cruikshank Behavioral Ecology, Conservation Planetary Astronomy Biology Sylvia A. Earle Botany Gretchen C. Daily Rodolfo Dirzo Conservation Biology Ecology and Conservation Biology Anne H. Ehrlich Conservation Biology Howell V. Daly Daniel Doak Entomology Evolutionary Ecology Page 3 of 11 Academy Fellows Paul R. Ehrlich Marcus W. Feldman Inez Fung Conservation Biology Mathematical Biology Atmosphere-Biosphere Interactions Emil R. Eisenhardt Carl J. Ferraris Jon S. Galehouse Conservation Ichthyology Geology Paul Ekman Peggy L. Fiedler Newell T. Garfield Comparative Psychology Botany Physical Oceanography Charles Elachi Christopher B. Field Carlos Garza Earth, Ocean and Atmosphere Global Ecology Molecular Ecology, Conservation Science from Space Genetics Alexei Filippenko Peter M. Elias Astronomy Jacques A. Gauthier Medicine Herpetology Paul V. Fine Thomas S. Elias Ecology and Evolutionary Biology Jonathan B. Geller Botany Marine Ecology and Marine Brian Fisher Molecular Ecology David Epel Entomology Cell Biology and Developmental Wayne M. Getz Biology Jonathan A. Foley Entomology Global Sustainability Don C. Erman Andrea Ghez Fisheries Ecology Susan L. Forsburg Astrophysics Molecular Biology, Cancer Biology Terry L. Erwin and DNA Michael T. Ghiselin Entomology Invertebrate Zoology Michael S. Foster William N. Eschmeyer Marine Biology Gregory S. Gilbert Ichthyology Microbial Ecology Laurel R. Fox James A. Estes Ecology Rosemary G. Gillespie Marine Ecology Entomology Andrew Fraknoi Sandra Faber Astronomy Gonzalo Giribet Astronomy Entomology Laurence G. Frank Gregory C. Farrington Mammalogy Alexander N. Glazer Chemistry Molecular and Conservation Biology Peter W. Fritsch Daphne G. Fautin Botany Stephen E. Glickman Invertebrate Zoology Psychology Robert J. Full Lewis J. Feldman Zoology Lynda J. Goff Botany Phycology Page 4 of 11 Academy Fellows David M. Golden Claes Gustafsson Colin O. Hermans Chemistry Molecular Biology and Biotechnology Invertebrate Zoology Charles R. Goldman Elizabeth Hadly Donald Heyneman Zoology Mammalogy, Vertebrate Evolution, Invertebrate Zoology, Tropical Climate Change Medicine, Epidemiology and Michael A. Goldman Parasitology Medical Genetics and Chromosomal John E. Hafernik Evolution Entomology Carole S. Hickman Paleontology Edgardo D. Gomez Benjamin Halpern Marine Biology, Conservation Marine Ecology Tessa M. Hill Earth Sciences Deborah M. Gordon David Halpern Ecology Physical Oceanography Stanley Hillman Environmental Physiology of George C. Gorman Alexander Harcourt Amphibians and Herpetology Evolutionary Biology Primatology Frederick G. Hochberg, Jr. Terrence M. Gosliner Susan P. Harrison Invertebrate Zoology Malacology Conservation Biology Tori M. Hoehler Michael H. Graham Christopher Harrold Astrobiology and Biochemistry Marine Ecology Marine Biology, Conservation John P. Holdren Harry W. Greene John Harte Energy and Natural Resources Herpetology Environmental Studies Karen Holl H. Gary Greene James T. Harvey Restoration Ecology Marine Geology Marine Biology Gustavo Hormiga David W. Greenfield Christine A. Hastorf Entomology Ichthyology Anthropology Kim M. Howell Gary B. Griggs David Haussler Herpetology Geology and Oceanology Computer Science Sarah B. Hrdy Charles E. Griswold Tyrone B. Hayes Anthropology Entomology Herpetology James C. Ingle Richard K. Grosberg Michael S. Hedrick Paleontology Evolution and Invertebrate Biology Environmental Physiology and Herpetology B. L. Ingram Karen Grove Geology Ecology Carl E. Heiles Astronomy Page 5 of 11 Academy Fellows Michael E. Irwin Patrick V. Kirch John D. Lattin Entomology Anthropology Entomology Lynne A. Isbell David H. Kistner James V. Lawry, Jr. Evolutionary Anthropology Entomology Anatomy Tomio Iwamoto Richard G. Klein Burney J. Le Boeuf Ichthyology Paleoanthropology Biology Nina G. Jablonski Paul L. L. Koch Richard E. Leakey Anthropology Paleobiology and Geobiology Anthropology Rodney M. Jackson Patrick Kociolek Walter S. Leal Conservation Biology Diatom Studies Chemical Ecology and Entomology Matthew J. James Mimi A. Koehl Gretchen LeBuhn Paleontology and History of Science Invertebrate Zoology Evolutionary Ecology David L. Jameson Walter D. Koenig Welton Lee Zoology Avian Ecology Invertebrate Zoology William A. Jensen Claire Kremen Robert Leidy Botany Conservation Biology and Wetland Specialist, Aquatic Ecology, Entomology Fish Biology Donald C. Johanson Anthropology Leonard V. Kuhi Margaret S. Leinen Astronomy Oceanography, Climate Engineering Michael N. Josselyn Ecology and Phycology Eileen A. Lacey Janet L. Leonard Behavioral Ecology and Mammalogy Neurobiology and Behavior David H. Kavanaugh Entomology Watson M. Laetsch William A. Lester Botany Physical Chemistry James C. Kelley Oceanography
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