The Academy is governed by a distinguished group of eminent scientists who have been appointed Fellows of the California Academy of Sciences in recognition of their notable contributions to one or more of the natural sciences. Nominated by their colleagues and appointed by the Board of Trustees, the Fellows remain members of the Fellowship for life.
Below is a complete list of Academy Fellows.
David Ackerly Alissa J. Arp Shannon N. Bennett Ecology and Evolutionary Biology Marine Biology Microbiology
Bruce M. Alberts John C. Avise William J. Bennetta Molecular and Conservation Biology Genetics and Ecology Science and Public Policy
Angel C. Alcala Francisco J. Ayala Robert G. Bergman Marine Biology and Conservation, Genetics Chemistry Herpetology Bruce G. Baldwin Brent Berlin Michele L. Aldrich Botany Anthropology History of Science George E. Ball Giacomo Bernardi Zeresenay Alemseged Entomology Ichthyology Anthropology and Paleoanthropology Richard T. Barber Vladimir A. Beshkov Frank Almeda Marine Biology Herpetology Botany Anthony D. Barnosky Devaki Bhaya Walter Alvarez Paleontology Molecular Biology Geology Reginald H. Barrett Philip E. Bickler Steven C. Anderson Applied Biology, Agricultural and Comparative Physiology, Systematic, Biogeographic and Environmental Science Neuroscience and Anesthesiology Ecological Studies in Herpetology John Barron James L. Bischoff Christopher Andrews Micropaleontology Marine Geology Aquarium Management and Parasitology Bruce Bartholomew Jeffrey M. Black Botany Conservation and Ecology of Water Fred Andrews Fowl Entomology Gibor B. Basri Astronomy and Astrophysics David C. Blackburn Paul H. Arnaud Herpetology Entomology Natalie Batalha Astronomy Elizabeth H. Blackburn E. N. Arnold Molecular Biology Herpetology Aaron M. Bauer Herpetology James A. Blake Jeanne Arnold Invertebrate Zoology Anthropology Steven V. Beckwith Astronomy
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Academy Fellows
Barbara A. Block Robert W. Buddemeier Mark H. Carr Evolutionary, Cellular and Molecular Environmental Studies Marine Ecology Physiology Stephen H. Bullock John Caskey Carol L. Boggs Plant Ecology Geology, Neotectonics Evolutionary Ecology Margaret G. Burke Alan Channing Stephen Bollens Behavioral Ecology and Education Herpetology Biological Oceanography Nathalie A. Cabrol Nona Chiariello Penelope Boston Astrobiology Global Ecology Astrobiology Gregor M. Cailliet Robert L. Christiansen Sarane T. Bowen Ichthyology Geology Genetics Roy L. Caldwell Christopher F. Chyba Rauri C.K. Bowie Invertebrate Zoology Astrobiology Ornithology Douglas G. Capone Frank Cipriano Justin Brashares Biological Oceanography, Marine Ecology and Evolutionary Biology Ecology and Wildlife Conservation Microbiology David A. Clague John I. Brauman Ring Cardé Volcanology Chemistry Entomology William A. Clemens Thomas S. Briggs James R. Carey Paleontology Entomology Entomology Lloyd S. Cluff Winslow R. Briggs Sherwin Carlquist Geology Botany Botany Kenneth Coale Adriana Briscoe Sandra J. Carlson Oceanography Evolutionary Biology Paleontology William P. Cochlan Michael E. Brown James T. Carlton Biological Oceanography Astronomy Marine Biodiversity and Conservation Anne C. Cohen Robert D. Brown, Jr. Edward J. Carpenter Invertebrate Systematics Geology Biological Oceanography and Marine Microbiology Daniel M. Cohen Thomas D. Bruns Ichthyology Mycology James M. Carpenter Entomology Sarah Cohen Richard C. Brusca Marine Genetics Marine Biology and Conservation Kent E. Carpenter Marine Biology and Ichthyology
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Academy Fellows
James P. Collman Marymeghan Daly Carmen Domingo Chemistry Animal Systematics and Ecology, Science Education Cnidaria Robert K. Colwell Robert V. Dowell Zoology Thomas F. Daniel Entomology Botany Lynn Cominsky John T. Doyen Cosmology Bruno David Entomology Paleontology, Marine Biology, Margaret W. Conkey Evolution James A. Doyle Anthropology Botany Todd E. Dawson Bruce Conklin Physiological Plant Ecology William T. Doyle Bioinformatics, Genomics and Botany Molecular Pharmacology Donald J. DePaolo Geosciences Frank Drake Edward F. Connor Astronomy Ecology Ruth DeFries Landscape Ecology Robert C. Drewes T. J. Conomos Herpetology Marine Geology Joseph DeRisi Molecular Biology, Genomics, Richard Dugdale Claudia Corti Parasitology, Virology, Oceanography Herpetology Computational Biology John P. Dumbacher Lawrence Cory David Des Marais Ornithology Zoology Geology and Astrobiology Thomas O. Duncan Daniel P. Costa Dennis E. Desjardin Botany Vertebrate Ecology Mycology Thomas Dunne Daniel E. Crocker Susan P. Desmond-Hellmann Geomorphology and Hydrology Ecology and Physiology Oncology and Biotechnology William H. Durham Donald Croll Marian C. Diamond Human Biology Ecology of Marine Predators, Island Anatomy Ecology and Conservation David M. Eakin Janis Dickinson Physics Dale P. Cruikshank Behavioral Ecology, Conservation Planetary Astronomy Biology Sylvia A. Earle Botany Gretchen C. Daily Rodolfo Dirzo Conservation Biology Ecology and Conservation Biology Anne H. Ehrlich Conservation Biology Howell V. Daly Daniel Doak Entomology Evolutionary Ecology
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Academy Fellows
Paul R. Ehrlich Marcus W. Feldman Inez Fung Conservation Biology Mathematical Biology Atmosphere-Biosphere Interactions
Emil R. Eisenhardt Carl J. Ferraris Jon S. Galehouse Conservation Ichthyology Geology
Paul Ekman Peggy L. Fiedler Newell T. Garfield Comparative Psychology Botany Physical Oceanography
Charles Elachi Christopher B. Field Carlos Garza Earth, Ocean and Atmosphere Global Ecology Molecular Ecology, Conservation Science from Space Genetics Alexei Filippenko Peter M. Elias Astronomy Jacques A. Gauthier Medicine Herpetology Paul V. Fine Thomas S. Elias Ecology and Evolutionary Biology Jonathan B. Geller Botany Marine Ecology and Marine Brian Fisher Molecular Ecology David Epel Entomology Cell Biology and Developmental Wayne M. Getz Biology Jonathan A. Foley Entomology Global Sustainability Don C. Erman Andrea Ghez Fisheries Ecology Susan L. Forsburg Astrophysics Molecular Biology, Cancer Biology Terry L. Erwin and DNA Michael T. Ghiselin Entomology Invertebrate Zoology Michael S. Foster William N. Eschmeyer Marine Biology Gregory S. Gilbert Ichthyology Microbial Ecology Laurel R. Fox James A. Estes Ecology Rosemary G. Gillespie Marine Ecology Entomology Andrew Fraknoi Sandra Faber Astronomy Gonzalo Giribet Astronomy Entomology Laurence G. Frank Gregory C. Farrington Mammalogy Alexander N. Glazer Chemistry Molecular and Conservation Biology Peter W. Fritsch Daphne G. Fautin Botany Stephen E. Glickman Invertebrate Zoology Psychology Robert J. Full Lewis J. Feldman Zoology Lynda J. Goff Botany Phycology
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Academy Fellows
David M. Golden Claes Gustafsson Colin O. Hermans Chemistry Molecular Biology and Biotechnology Invertebrate Zoology
Charles R. Goldman Elizabeth Hadly Donald Heyneman Zoology Mammalogy, Vertebrate Evolution, Invertebrate Zoology, Tropical Climate Change Medicine, Epidemiology and Michael A. Goldman Parasitology Medical Genetics and Chromosomal John E. Hafernik Evolution Entomology Carole S. Hickman Paleontology Edgardo D. Gomez Benjamin Halpern Marine Biology, Conservation Marine Ecology Tessa M. Hill Earth Sciences Deborah M. Gordon David Halpern Ecology Physical Oceanography Stanley Hillman Environmental Physiology of George C. Gorman Alexander Harcourt Amphibians and Herpetology Evolutionary Biology Primatology Frederick G. Hochberg, Jr. Terrence M. Gosliner Susan P. Harrison Invertebrate Zoology Malacology Conservation Biology Tori M. Hoehler Michael H. Graham Christopher Harrold Astrobiology and Biochemistry Marine Ecology Marine Biology, Conservation John P. Holdren Harry W. Greene John Harte Energy and Natural Resources Herpetology Environmental Studies Karen Holl H. Gary Greene James T. Harvey Restoration Ecology Marine Geology Marine Biology Gustavo Hormiga David W. Greenfield Christine A. Hastorf Entomology Ichthyology Anthropology Kim M. Howell Gary B. Griggs David Haussler Herpetology Geology and Oceanology Computer Science Sarah B. Hrdy Charles E. Griswold Tyrone B. Hayes Anthropology Entomology Herpetology James C. Ingle Richard K. Grosberg Michael S. Hedrick Paleontology Evolution and Invertebrate Biology Environmental Physiology and Herpetology B. L. Ingram Karen Grove Geology Ecology Carl E. Heiles Astronomy
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Academy Fellows
Michael E. Irwin Patrick V. Kirch John D. Lattin Entomology Anthropology Entomology
Lynne A. Isbell David H. Kistner James V. Lawry, Jr. Evolutionary Anthropology Entomology Anatomy
Tomio Iwamoto Richard G. Klein Burney J. Le Boeuf Ichthyology Paleoanthropology Biology
Nina G. Jablonski Paul L. L. Koch Richard E. Leakey Anthropology Paleobiology and Geobiology Anthropology
Rodney M. Jackson Patrick Kociolek Walter S. Leal Conservation Biology Diatom Studies Chemical Ecology and Entomology
Matthew J. James Mimi A. Koehl Gretchen LeBuhn Paleontology and History of Science Invertebrate Zoology Evolutionary Ecology
David L. Jameson Walter D. Koenig Welton Lee Zoology Avian Ecology Invertebrate Zoology
William A. Jensen Claire Kremen Robert Leidy Botany Conservation Biology and Wetland Specialist, Aquatic Ecology, Entomology Fish Biology Donald C. Johanson Anthropology Leonard V. Kuhi Margaret S. Leinen Astronomy Oceanography, Climate Engineering Michael N. Josselyn Ecology and Phycology Eileen A. Lacey Janet L. Leonard Behavioral Ecology and Mammalogy Neurobiology and Behavior David H. Kavanaugh Entomology Watson M. Laetsch William A. Lester Botany Physical Chemistry James C. Kelley Oceanography Michael D. Lagios Alan E. Leviton Ichthyology Herpetology Donald Kennedy Biological Sciences Robert S. Lane Matthew R. Lewin Entomology Emergency Medicine William H. Kimbel Anthropology Leo F. Laporte Lowell N. Lewis Geology Botany Wim Kimmerer Estuarine Ecology Ralph J. Larson Wilfredo Licuanan Ichthyology Coral reef ecology
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Academy Fellows
Paul Licht York T. Mandra Fiorenza Micheli Herpetology Paleontology Marine Ecology and Conservation
William Z. Lidicker, Jr. Marc S. Mangel Kathy Ann Miller Mammalogy Behavior and Ecology Marine Algal Systematics
Kent G. Lightfoot Geoffrey Marcy Katharine Milton Archaeology and Anthropology Astrophysics Anthropology
David R. Lindberg Charles R. Marshall Brent D. Mishler Malacology Paleontology Botany
Jere H. Lipps Pamela A. Matson Richard L. Moe Paleontology Environmental Studies Phycology
Werner J. Loher Claire Max John P. Monteverdi Entomology Astronomy, Adaptive Optics Meteorology
Sharon R. Long Daniel McCleese Richard J. Mooi Plant and Microbial Symbiosis Space Science Invertebrate Zoology
Jerold Lowenstein John E. McCosker Harold A. Mooney Medicine Aquatic Biology Environmental Science
Margaret D. Lowman Alistair W. McCrone Eldridge Moores Ecology Geology Geology
Jane Lubchenco Jimmy A. McGuire Michael J. Moratto Marine Biology Herpetology, Vertebrate Biology and Anthropology Evolutionary Biology Bruce E. Lyon Steve Morgan Behavioral Ecology Henry M. McHenry Behavioral Ecology, Biological Biological Anthropology Oceanography James M. Lyons Plant Physiology Monica Medina Craig C. Moritz Evolutionary Genomics and Systems Evolutionary and Conservation James P. Mackey Biology Genetics Herpetology Gopinathan Menon David D. Morrison David Maddison Biochemistry Planetary Astronomy Entomology, Informatics, Computer Programming and Software Adina Merenlender Carol W. Mosher Development Conservation Biology Chemistry
Rocco Mancinelli Elisabeth Micheli Peter B. Moyle Astrobiology Eco-Hydrology Ichthyology
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Academy Fellows
Richard A. Muller Stephen R. Palumbi Lewis Petrinovich Astrophysics Molecular Evolution and Ornithology Conservation Biology Michael A. Murphy Samuel George Philander Paleontology and Geology Theodore Papenfuss Ocean-Atmosphere Interactions Herpetology Michael W. Nachman Theodore W. Pietsch Mammalogy and Evolutionary Lynne R. Parenti Ichthyology Genetics Ichthyology Carl B. Pilcher Donald J. Nevins Ingrid Parker Astrobiology, Astronomy, Planetary Botany Ecology Science
Andrew T. Newberry Sue Taylor Parker Edwin P. Pister Invertebrate Biology Anthropology Conservation Biology
Frederic H. Nichols Thomas Parker Karl S. Pister Invertebrate Zoology Botany Civil Engineering, Education
Karina Nielsen Dennis R. Parnell Norman I. Platnick Marine Ecology Botany Entomology
Thomas M. Niesen Robert Patterson Grant Pogson Invertebrate Biology Botany Molecular Population Genetics and Ecological Genetics Daniel H. Norris James L. Patton Botany Mammalogy George O. Poinar, Jr. Entomology Peter Norvig John S. Pearse Artificial Intelligence and Computer Invertebrate Zoology Katherine Pollard Science Evolution and Computational Biology Vicki B. Pearse Amos M. Nur Invertebrate Zoology Carolyn Porco Geology Planetary Science Yvonne Pendleton Patrick M. O’Grady Astrophysics Donald C. Potts Evolutionary Biology Invertebrate Zoology David Perlman Richard E. Packard Journalism Jerry A. Powell Physics Entomology Saul Perlmutter Kevin Padian Astrophysics Thomas M. Powell Paleontology Oceanography, Limnology and Raymond Pestrong Aquatic Biology Robert E. Page, Jr. Geology Entomology
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Academy Fellows
Mary E. Power Rollin C. Richmond Eric J. Routman Ecology of Riverine Systems Evolutionary Genetics Herpetology
Joel R. Primack John W. Rick Rodolfo Ruibal Astronomy and Physics Archaeology and Paleoecology Herpetology
Wojciech J. Pulawski Neil J. Risch Bruce Runnegar Entomology Human Evolutionary Genetics and Geology and Paleontology Epidemiology Alexander H. Purcell Robert Sapolsky Entomology Bruce H. Robison Neuroendocrinology Deep-Sea Biology Peter R. Raimondi Jose Sarukhan Marine Ecology Luiz A. Rocha Botany Ichthyology Katherine Ralls Russell L. Schweickart Conservation Biology and Peter U. Rodda Astrobiology Mammalogy Geology Judith G. Scotchmoor Janet A. Randall George K. Roderick Science Education and Outreach Mammalogy Population Genetics and Evolutionary Biology Eugenie C. Scott Galen B. Rathbun Anthropology Mammalogy Peter Rodman Anthropology Ravinder N. M. Sehgal Peter H. Raven Avian Parasitology, Ecology of Botany Barbara Rogoff Infectious Disease Developmental Psychology David C. F. Rentz Erwin Seibel Entomology Peter Roopnarine Oceanography Invertebrate Paleontology Vincent H. Resh H. Bradley Shaffer Aquatic Biology Terry L. Root Herpetology Climate Change Sensitivity of Don R. Reynolds Vertebrates Arthur M. Shapiro Botany Zoology Barry Roth Wynetka A. Reynolds Malacology and Paleontology Beth Shapiro Biochemistry Ancient DNA Lynn J. Rothschild Mark Richards Astrobiology and Evolutionary Howard S. Shellhammer Geophysics Protistology Mammalogy
Peter J. Richerson Joan E. Roughgarden James R. Shevock Cultural Evolution Herpetology Botany
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Academy Fellows
Seth Shostak Michael E. Soule Carol Tang Astronomy Evolutionary and Conservation Paleontology Biology Neil H. Shubin Kimberly Tanner Evolutionary Biology Wayne P. Sousa Science Education Marine Ecology Petra Sierwald Jill C. Tarter Entomology Chelsea Specht Astronomy Plant Genetics and Genomics Mary W. Silver Catherine A. Tauber Biological Oceanography Howard J. Spero Insect Ecology, Behavior and Paleoceanography Physiology W. Brian Simison Genomics Gregory S. Spicer Leighton R. Taylor Invertebrate Zoology Ichthyology Ellen L. Simms Botany Maureen L. Stanton John W. Taylor Population Biology Mycology Barry R. Sinervo Ecology Calvin H. Stevens Richard R. Tenaza Geology Mammalogy Leonard S. Sklar Geology Jonathon H. Stillman Robbin W. Thorp Invertebrate Physiology and Entomology Doris H. Sloan Molecular Biology Geology Ignacio Tinoco Scott W. Stine Chemistry Alan R. Smith Geography Botany Regina Tomlinson Sharon Y. Strauss Invertebrate Zoology Edmund H. Smith Evolutionary Ecology Invertebrate Zoology O. Brian Toon John L. Strother Atmospheric Science Edward L. Smith Botany Entomology Jonathan D. Trent Raymond Sullivan Astrobiology, Marine Biology and Thomas B. Smith Geology Microbiology Ecology James R. Sweeney Brian E. Tucker George Somero Botany Seismology, Seismic Safety, Geology Evolutionary Biochemistry Frank Talamantes Ronald D. Vale Victoria L. Sork Endocrinology Cell Biology, Public Science Education Evolutionary Biology Frank H. Talbot James W. Valentine Marine Biology Paleontology
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Academy Fellows
Blaire Van Valkenburgh Robert Wayne James C. Zachos Vertebrate Paleontology Molecular Systematics Paleoclimatology, Marine Stratigraphy and Paloeoceanography Geerat J. Vermeij William (Jack) Welch Evolutionary Biology Astronomy Frank G. Zalom Entomology Peter M. Vitousek Christian M. Wemmer Ecology Conservation Biology Richard N. Zare Chemistry Vance T. Vredenburg Cherie L. Wetzel Herpetology Botany Erika S. Zavaleta Community and Ecosystem Ecology, Peter C. Wainwright Lisa E. White Conservation Biology Evolution and Ecology Geology and Paleontology Ermi Zhao David B. Wake Timothy D. White Herpetology Herpetology Anthropology Adrienne L. Zihlman Marvalee H. Wake Frances Wilkerson Anthropology Zoology Biological Oceanography Irving Zucker Diana H. Wall Mark Wilkinson Biology Soil Biology Herpetology
Robert C. Walter Kipling W. Will Geology Entomology
Peter D. Ward Gary C. Williams Geology Invertebrate Zoology
Philip S. Ward R. Sanders Williams Ant Systematics Medicine, Cell Biology and Genetics
Marius S. Wasbauer Stanley C. Williams Entomology Entomology
Dorothy Washburn Susan L. Williams Anthropology Marine Ecology
Kenneth E. Watt Terrie Williams Zoology Comparative Physiology
Ward B. Watt Dawn Wright Evolutionary Biology Oceanography
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