2.2. Wonderwerk Cave

Total Page:16

File Type:pdf, Size:1020Kb

2.2. Wonderwerk Cave A Micromorphological Approach to Inferring Paleo-Lake System Phases: The Case Study of the Earlier Stone Age at Wonderwerk Cave, South Africa by Ailidh Hathway B.A. (Hons.), Wilfrid Laurier University, 2017 Thesis Submitted in Partial Fulfillment of the Requirements for the Degree of Master of Arts in the Department of Archaeology Faculty of Environment © Ailidh Hathway 2021 SIMON FRASER UNIVERSITY Spring 2021 Copyright in this work rests with the author. Please ensure that any reproduction or re-use is done in accordance with the relevant national copyright legislation. Declaration of Committee Name: Ailidh Hathway Degree: Master of Arts Title: A Micromorphological Approach to Inferring Paleo-Lake System Phases: The Case Study of the Earlier Stone Age at Wonderwerk Cave, South Africa Committee: Chair: Dana Lepofsky Professor, Archaeology Francesco Berna Supervisor Associate Professor, Archaeology George Nicholas Committee Member Professor, Archaeology Paul Goldberg Committee Member Adjunct Faculty, Archaeology Vance T. Holliday Examiner Professor, Anthropology and Geosciences University of Arizona ii Abstract Playa lakes are arid region ephemeral bodies of water that have been found in association with important archaeological sites. These lakes produce distinct sediments in response to changing hydrological and environmental conditions. To provide the means to more effectively study playa lake sediments, I developed an analytical protocol and a model that utilizes micromorphology and grain size distribution analysis of thin sections to identify and interpret paleo-playa lake phases preserved in intact archaeological deposits. To assess the potential of the analytical procedure, I applied it to thin sections collected from Earlier Stone Age deposits at Wonderwerk Cave, South Africa, where a playa lake system existed in proximity to the cave. The results of the study show that sediments produced during different playa lake phases can be distinguished according to a specific set of criteria identifiable through micromorphology and grain size distribution analysis. Keywords: Micromorphology; Playa Lake; Wonderwerk Cave; South Africa; Earlier Stone Age iii Acknowledgements I thank Dr. Francesco Berna for providing me with this opportunity. Thank you for your guidance and expertise as I fumbled my way through this research. I am very appreciative of the hours spent in the lab and the field, passing on your knowledge and mentoring me in the realm of geoarchaeology. I also thank you for the opportunity to work on material from and excavate at a truly remarkable archaeological site. Thank you to Dr. George Nicholas for your guidance, feedback, support, and ultimately making me a better writer. I express my thanks to Dr. Paul Goldberg for offering your expertise and contributions to my thesis. Without your previous work, this research would not have been possible. The Wonderwerk Cave team and Co-Directors, Dr. Michael Chazan, Dr. Liora Kolska Horwitz. Dr. Nathan Jankowski for offering expertise and advice on Lake Mungo sediments. I would not have been able to complete this research without the funding provided by the Social Sciences and Humanities Research Council and the SFU Department of Archaeology. I express my gratitude to Dr. Laure Spake for sharing your skills when it comes to coding in R, but more importantly the endless support, encouragement, and fun times. Thank you to Emily Purcell and Matthea Wiebe for the friendship, support, and encouragement over the last few years. Thank you to Sarah Johnson for the adventures and support. A very special thank you to Candice Koopowitz for your friendship, kindness, guidance, endless laughing, and being my personal tour guide in South Africa. Thank you for being my favourite water gun target. Finally, a special thank you to all my family and friends that encouraged me to keep going when I did not think I could. Without that encouragement I would not have been able to do it. iv Table of Contents Declaration of Committee .................................................................................................. ii Abstract .............................................................................................................................iii Acknowledgements .......................................................................................................... iv Table of Contents .............................................................................................................. v List of Tables ................................................................................................................... viii List of Figures................................................................................................................... ix List of Acronyms .............................................................................................................. xiii Chapter 1. Introduction ................................................................................................ 1 1.1. Research Aims and Objectives ............................................................................... 4 1.2. Thesis Outline ......................................................................................................... 6 Chapter 2. Background ................................................................................................ 8 2.1. Environments and Human Evolution in Africa ......................................................... 8 2.1.1. The Savanna Hypothesis ................................................................................ 8 2.1.2. The Mosaic Environment Hypothesis ............................................................ 10 Temporal and Spatial Mosaics ................................................................................ 10 2.1.3. Lake Margin Settings and Early Hominin Archaeological Sites .................... 12 2.2. Wonderwerk Cave ................................................................................................. 15 2.2.1. Previous Sedimentological and Micromorphological Investigations of the Earlier Stone Age at Wonderwerk Cave ...................................................................... 17 2.2.2. The Potential of Wonderwerk Cave for Testing the Protocol ........................ 18 2.3. Cave Systems ....................................................................................................... 19 2.3.1. Cave Sediments ............................................................................................ 20 2.4. Playa Lakes ........................................................................................................... 20 2.4.1. The Playa Lake Cycle ................................................................................... 21 2.4.2. Playa Lakes in the Geological Record .......................................................... 22 2.5. Lunette Dunes ....................................................................................................... 23 2.5.1. The Formation of Lunette Dunes .................................................................. 24 2.5.2. Clay Aggregates ............................................................................................ 26 2.6. Previous Studies of Playa Lake Sediments in Archaeological Sites ..................... 26 2.6.1. Southern Africa ............................................................................................. 27 2.6.2. The Willandra Lakes Complex and Lake Mungo .......................................... 28 Chapter 3. Methodology ............................................................................................ 33 3.1. Earlier Stone Age Deposits at Wonderwerk Cave ................................................ 33 3.1.1. Lithostratigraphy ............................................................................................ 33 3.2. Modelling Playa Lake Phases Based on Sediment Composition .......................... 37 3.3. Analytical Protocol ................................................................................................. 40 3.3.1. Sedimentology and Soil Micromorphology .................................................... 41 3.3.1.1 Sampling Intact Sediment Blocks ................................................................ 43 Thin Section Preparation ......................................................................................... 43 3.3.1.2 Microfabric Units: Lithostratigraphy at the Microscale ................................. 43 v 3.3.1.3 Micrographs and Subsamples ..................................................................... 44 3.3.1.4 Texture and Sorting ..................................................................................... 45 3.3.1.5 Microstructure .............................................................................................. 46 3.3.1.6 Groundmass: c/f Related Distribution, Coarse Material and Micromass ..... 47 3.3.1.7 Mineral Composition .................................................................................... 48 3.3.1.8 Diagenesis and Pedofeatures ...................................................................... 49 3.3.2. Grain Size Distribution Analysis .................................................................... 52 Particle Size Analysis .............................................................................................. 52 Graphing Grain Size Distribution ............................................................................
Recommended publications
  • The Origin and Early Evolution of Vascular Plant Shoots and Leaves Rstb.Royalsocietypublishing.Org C
    Downloaded from http://rstb.royalsocietypublishing.org/ on January 22, 2018 The origin and early evolution of vascular plant shoots and leaves rstb.royalsocietypublishing.org C. Jill Harrison 1 and Jennifer L. Morris 2 1School of Biological Sciences, and 2School of Earth Sciences, University of Bristol, 24 Tyndall Avenue, Bristol BS8 1TQ, UK Review CJH, 0000-0002-5228-600X; JLM, 0000-0002-7453-3841 Cite this article: Harrison CJ, Morris JL. 2017 The morphology of plant fossils from the Rhynie chert has generated long- standing questions about vascular plant shoot and leaf evolution, for The origin and early evolution of vascular plant instance, which morphologies were ancestral within land plants, when did shoots and leaves. Phil. Trans. R. Soc. B 373 : vascular plants first arise and did leaves have multiple evolutionary origins? 20160496. Recent advances combining insights from molecular phylogeny, palaeobotany http://dx.doi.org/10.1098/rstb.2016.0496 and evo–devo research address these questions and suggest the sequence of morphological innovation during vascular plant shoot and leaf evolution. The evidence pinpoints testable developmental and genetic hypotheses relat- Accepted: 11 August 2017 ing to the origin of branching and indeterminate shoot architectures prior to the evolution of leaves, and demonstrates underestimation of polyphyly in One contribution of 18 to a discussion meeting the evolution of leaves from branching forms in ‘telome theory’ hypotheses issue ‘The Rhynie cherts: our earliest terrestrial of leaf evolution. This review discusses fossil, developmental and genetic ecosystem revisited’. evidence relating to the evolution of vascular plant shoots and leaves in a phylogenetic framework. This article is part of a discussion meeting issue ‘The Rhynie cherts: our Subject Areas: earliest terrestrial ecosystem revisited’.
    [Show full text]
  • Paleoecology and Land-Use of Quaternary Megafauna from Saltville, Virginia Emily Simpson East Tennessee State University
    East Tennessee State University Digital Commons @ East Tennessee State University Electronic Theses and Dissertations Student Works 5-2019 Paleoecology and Land-Use of Quaternary Megafauna from Saltville, Virginia Emily Simpson East Tennessee State University Follow this and additional works at: https://dc.etsu.edu/etd Part of the Paleontology Commons Recommended Citation Simpson, Emily, "Paleoecology and Land-Use of Quaternary Megafauna from Saltville, Virginia" (2019). Electronic Theses and Dissertations. Paper 3590. https://dc.etsu.edu/etd/3590 This Thesis - Open Access is brought to you for free and open access by the Student Works at Digital Commons @ East Tennessee State University. It has been accepted for inclusion in Electronic Theses and Dissertations by an authorized administrator of Digital Commons @ East Tennessee State University. For more information, please contact [email protected]. Paleoecology and Land-Use of Quaternary Megafauna from Saltville, Virginia ________________________________ A thesis presented to the faculty of the Department of Geosciences East Tennessee State University In partial fulfillment of the requirements for the degree Master of Science in Geosciences with a concentration in Paleontology _______________________________ by Emily Michelle Bruff Simpson May 2019 ________________________________ Dr. Chris Widga, Chair Dr. Blaine W. Schubert Dr. Andrew Joyner Key Words: Paleoecology, land-use, grassy balds, stable isotope ecology, Whitetop Mountain ABSTRACT Paleoecology and Land-Use of Quaternary Megafauna from Saltville, Virginia by Emily Michelle Bruff Simpson Land-use, feeding habits, and response to seasonality by Quaternary megaherbivores in Saltville, Virginia, is poorly understood. Stable isotope analyses of serially sampled Bootherium and Equus enamel from Saltville were used to explore seasonally calibrated (δ18O) patterns in megaherbivore diet (δ13C) and land-use (87Sr/86Sr).
    [Show full text]
  • Evolution of the Female Conifer Cone Fossils, Morphology and Phylogenetics
    DEPARTMENT OF BIOLOGICAL AND ENVIRONMENTAL SCIENCES EVOLUTION OF THE FEMALE CONIFER CONE FOSSILS, MORPHOLOGY AND PHYLOGENETICS Daniel Bäck Degree project for Bachelor of Science with a major in Biology BIO602, Biologi: Examensarbete – kandidatexamen, 15 hp First cycle Semester/year: Spring 2020 Supervisor: Åslög Dahl, Department of Biological and Environmental Sciences Examiner: Claes Persson, Department of Biological and Environmental Sciences Front page: Abies koreana (immature seed cones), Gothenburg Botanical Garden, Sweden Table of contents 1 Abstract ............................................................................................................................... 2 2 Introduction ......................................................................................................................... 3 2.1 Brief history of Florin’s research ............................................................................... 3 2.2 Progress in conifer phylogenetics .............................................................................. 4 3 Aims .................................................................................................................................... 4 4 Materials and Methods ........................................................................................................ 4 4.1 Literature: ................................................................................................................... 4 4.2 RStudio: .....................................................................................................................
    [Show full text]
  • Curriculum Vitae
    CURRICULUM VITAE ORCID ID: 0000-0003-0186-6546 Gar W. Rothwell Edwin and Ruth Kennedy Distinguished Professor Emeritus Department of Environmental and Plant Biology Porter Hall 401E T: 740 593 1129 Ohio University F: 740 593 1130 Athens, OH 45701 E: [email protected] also Courtesy Professor Department of Botany and PlantPathology Oregon State University T: 541 737- 5252 Corvallis, OR 97331 E: [email protected] Education Ph.D.,1973 University of Alberta (Botany) M.S., 1969 University of Illinois, Chicago (Biology) B.A., 1966 Central Washington University (Biology) Academic Awards and Honors 2018 International Organisation of Palaeobotany lifetime Honorary Membership 2014 Fellow of the Paleontological Society 2009 Distinguished Fellow of the Botanical Society of America 2004 Ohio University Distinguished Professor 2002 Michael A. Cichan Award, Botanical Society of America 1999-2004 Ohio University Presidential Research Scholar in Biomedical and Life Sciences 1993 Edgar T. Wherry Award, Botanical Society of America 1991-1992 Outstanding Graduate Faculty Award, Ohio University 1982-1983 Chairman, Paleobotanical Section, Botanical Society of America 1972-1973 University of Alberta Dissertation Fellow 1971 Paleobotanical (Isabel Cookson) Award, Botanical Society of America Positions Held 2011-present Courtesy Professor of Botany and Plant Pathology, Oregon State University 2008-2009 Visiting Senior Researcher, University of Alberta 2004-present Edwin and Ruth Kennedy Distinguished Professor of Environmental and Plant Biology, Ohio
    [Show full text]
  • Transformative Paleobotany
    Chapter 6 Lower Permian Flora of the Sanzenbacher Ranch, Clay County, Texas William A. DiMichele1, Robert W. Hook2, Hans Kerp3, Carol L. Hotton1,4, Cindy V. Looy5 and Dan S. Chaney1 1NMNH Smithsonian Institution, Washington, DC, United States; 2The University of Texas at Austin, Austin, TX, United States; 3Westfälische Wilhelms-Universität Münster, Münster, Germany; 4National Institutes of Health, Bethesda, MD, United States; 5University of California Berkeley, Berkeley, CA, United States 1. INTRODUCTION 1985; Broutin, 1986; Popa, 1999; Steyer et al., 2000; Wagner and Mayoral, 2007; Bercovici and Broutin, 2008; Since 1989, field parties supported by the U.S. National Barthel, 2009; Wagner and Álvarez-Vázquez, 2010; Museum of Natural History have obtained large collections Barthel and Brauner, 2015). Furthermore, because this of mainly Permian plant fossils from north central Texas. locality was collected on three occasions over a time period This work was undertaken to study known localities and to of 50 years and by different parties, comparative analysis of find new fossiliferous deposits that would contribute to a the Sanzenbacher collections provides a basis for assessing better understanding of floral and paleoenvironmental sites that have comparable histories. changes within the region during the early Permian. From the outset, the effort was interdisciplinary and grew, through the contributions of nearly 20 paleobotanists, 2. GEOLOGY palynologists, invertebrate and vertebrate paleontologists, Clay County is the only county in the Permo-Carboniferous and sedimentary geologists of several subdisciplines, to be outcrop belt of north central Texas that lacks marine rocks. quite comprehensive. Our reporting of results, however, has These alluvial sediments accumulated east of a broad been influenced by unexpected developments, including the coastal plain that bordered the Eastern Shelf of the Midland discovery of new plant-fossil assemblages in areas once Basin.
    [Show full text]
  • Phylogenetic Analyses of Cretaceous Fossils Related to Chloranthaceae and Their Evolutionary Implications
    UC Davis UC Davis Previously Published Works Title Phylogenetic Analyses of Cretaceous Fossils Related to Chloranthaceae and their Evolutionary Implications Permalink https://escholarship.org/uc/item/0d58r5r0 Journal Botanical Review, 84(2) ISSN 0006-8101 Authors Doyle, JA Endress, PK Publication Date 2018-06-01 DOI 10.1007/s12229-018-9197-6 Peer reviewed eScholarship.org Powered by the California Digital Library University of California Phylogenetic Analyses of Cretaceous Fossils Related to Chloranthaceae and their Evolutionary Implications James A. Doyle & Peter K. Endress The Botanical Review ISSN 0006-8101 Volume 84 Number 2 Bot. Rev. (2018) 84:156-202 DOI 10.1007/s12229-018-9197-6 1 23 Your article is protected by copyright and all rights are held exclusively by The New York Botanical Garden. This e-offprint is for personal use only and shall not be self- archived in electronic repositories. If you wish to self-archive your article, please use the accepted manuscript version for posting on your own website. You may further deposit the accepted manuscript version in any repository, provided it is only made publicly available 12 months after official publication or later and provided acknowledgement is given to the original source of publication and a link is inserted to the published article on Springer's website. The link must be accompanied by the following text: "The final publication is available at link.springer.com”. 1 23 Author's personal copy Bot. Rev. (2018) 84:156–202 https://doi.org/10.1007/s12229-018-9197-6 Phylogenetic Analyses of Cretaceous Fossils Related to Chloranthaceae and their Evolutionary Implications James A.
    [Show full text]
  • A 23 Kyrs Record of Environmental Change from Cherangani Hills, Kenya
    Georgia State University ScholarWorks @ Georgia State University Geosciences Theses Department of Geosciences 5-10-2019 A 23 Kyrs Record Of Environmental Change From Cherangani Hills, Kenya Vicky Cheruiyot Follow this and additional works at: https://scholarworks.gsu.edu/geosciences_theses Recommended Citation Cheruiyot, Vicky, "A 23 Kyrs Record Of Environmental Change From Cherangani Hills, Kenya." Thesis, Georgia State University, 2019. https://scholarworks.gsu.edu/geosciences_theses/131 This Thesis is brought to you for free and open access by the Department of Geosciences at ScholarWorks @ Georgia State University. It has been accepted for inclusion in Geosciences Theses by an authorized administrator of ScholarWorks @ Georgia State University. For more information, please contact [email protected]. A 23 KYRS RECORD OF ENVIRONMENTAL CHANGE FROM CHERANGANI HILLS, KENYA by Vicky Cheruiyot Under supervision of Lawrence Kiage, PhD. ABSTRACT The sensitivity of East Africa’s environment to anthropogenic and natural climatic changes is poorly understood. Therefore, there is need to incorporate continuous high-resolution records to provide insights into the timings of climatic signals to reconstruct East Africa’s paleoenvironment. This study presents sediment core data (KAP-01) from Kapkanyar Swamp, Cherangani Hills, based on fungal spores, Loss-On-Ignition (LOI) and microscopic charcoal analyses. The climate during the Last Glacial Maximum (LGM) was generally cold and dry with intermittent presence of moist environmental conditions evidenced by the minimal presence of coprophilous taxa, a well-known taxon used to signify presence herbivores, representing diminished fauna. Presence of charcoal in the LGM reflects probable natural fires suggesting dry environmental conditions. The deglaciation period, on the other hand, was relatively colder and drier compared to any other period in the last ~23 Kyrs with scanty preservation fungal spores.
    [Show full text]
  • Two Examples from the Late Cretaceous of Western North America
    Paleobotanical Estimates of Climate in Deep Time: Two Examples From the Late Cretaceous of Western North America Garland Upchurch1, Ann Marie Prue1, Joan Parrott1, Emilio Estrada-Ruiz2, and Dori Contreras3 1 Texas State University, San Marcos, TX 2 National Polytechnic Institute of Mexico, Mexico City, Mexico 3 University of California, Berkeley, CA Introduction • Paleobotany provides important information on paleoclimate. – Shallow geologic time • Closest living relatives of fossil plant species • Paleoclimate inferred from modern species distributions – Deep geologic time • Relationships with extant species and genera more distant. • Adaptive characteristics of plants – Physiognomy • Paleoclimate inferred from relation between plant adaptive characteristics and modern climate Physiognomy • Physiognomy: Adaptive features of plant structure – Environment: Evolved multiple times – Life form (PFT) – Organs – Tissue structure Tropical leaf (left) vs. • Leaves of dicot flowering plants temperate zone leaf – Margin, shape, size • Woods – Presence/absence of annual rings – Dicot woods: Amount of soft tissue, water conduit diameter, other features Tropical (balsa) wood Physiognomic Methods • Leaves – Simple Linear Regression • Leaf Margin Analysis (LMA) – Mean Annual Temperature (MAT) • Leaf Area Analysis (LAA) Left to right: – Mean Annual Precipitation (MAP) Tropical, desert, and – Multivariate ordination temperate leaves • Climate Leaf Analysis Multivariate Program (CLAMP) • Mean annual and seasonal temperatures, precipitation – Multiple regression
    [Show full text]
  • Fossil Rodents from Bone Cave at the Koanaka Hills Locality
    FOSSIL RODENTS FROM BONE CAVE AT THE KOANAKA HILLS LOCALITY, BOTSWANA _____________ A Thesis Presented to The Faculty of the Department of Biological Sciences Sam Houston State University _____________ In Partial Fulfillment of the Requirements for the Degree of Master of Science _____________ by Zachary W. Pierce May, 2020 FOSSIL RODENTS FROM BONE CAVE AT THE KOANAKA HILLS LOCALITY, BOTSWANA by Zachary W. Pierce ______________ APPROVED: Patrick J. Lewis, PhD Thesis Director Monte L. Thies, PhD Committee Member Jeffrey R. Wozniak, PhD Committee Member John B. Pascarella, PhD Dean, College of Science and Engineering Technology DEDICATION This work is dedicated to my mom and dad, Drs. Maria and Robert Pierce. None of this is possible without the hard work and perseverance exhibited throughout their lives, and I am eternally grateful for this sacrifice. This work is also dedicated to my wife Lillian Pierce who provided constant love and support during this process which gave me the perseverance and motivation to finish. iii ABSTRACT Pierce, Zachary W, Fossil rodents from Bone Cave at the Koanaka Hills locality, Botswana. Master of Science (Biology), May, 2020, Sam Houston State University, Huntsville, Texas. In this study I analyze a Middle Pleistocene rodent fauna from Bone Cave locality, Koanaka Hills, northwestern Botswana and attempt to reconstruct the paleoenvironment of the surrounding area. Only a few Pliocene and Pleistocene fossil localities exist between eastern and southern Africa, and the fossil rodents collected from within the Koanaka Hills partially fills this significant geographic and temporal gap in the paleontological record of Africa. Rodent remains from owl accumulations are frequently found in the fossil record and used to reconstruct the paleoenvironment Similarly, prey remains from owl accumulations are used to reconstruct modern community composition.
    [Show full text]
  • Tracking Late-Quaternary Extinctions in Interior Alaska Using Megaherbivore Bone Remains and Dung Fungal Spores
    Type of Article: Research Article Title: Tracking late-Quaternary extinctions in interior Alaska using megaherbivore bone remains and dung fungal spores Authors: Keziah J. Conroy1,2, Ambroise G. Baker1,3 *, Vivienne J. Jones1*, Maarten van Hardenbroek4,5, Emma J. Hopla4, Robert Collier4, Adrian M. Lister 6, Mary E. Edwards4 1 ECRC, Department of Geography, University College London, Gower Street, London, WC1E 6BT 2 Leverhulme Centre for Human Evolutionary Studies, Henry Wellcome Building, Fitzwilliam Street, Cambridge, CB2 1QH 3 School of Health and Life Science, Teesside University, Middlesbrough, TS1 3BX 4 Geography and Environment, University of Southampton, Southampton, SO17 1BJ 5 Geography Politics and Sociology, Newcastle University, Newcastle upon Tyne, NE1 7RU 6 Department of Earth Sciences, Natural History Museum, London, SW7 5BD * Corresponding authors: Dr Baker: [email protected] +44 (0)164273 8208 Prof Jones: [email protected] +44 (0)20 7679 0555 1 ABSTRACT One major challenge in the study of late-Quaternary extinctions (LQEs) is providing better estimates of past megafauna abundance. To show how megaherbivore population size varied before and after the last extinctions in interior Alaska, we use both a database of radiocarbon-dated bone remains (spanning 25-0 ka) and spores of the obligate dung fungus, Sporormiella, recovered from radiocarbon-dated lake- sediment cores (spanning 17-0 ka). Bone fossils show that the last stage of LQEs in the region occurred at about 13 ka ago, but the number of megaherbivore bones remains high into the Holocene. Sporormiella abundance also remains high into the Holocene and does not decrease with major vegetation changes recorded by arboreal pollen percentages.
    [Show full text]
  • Alexandru Mihail Florian Tomescu, Ph.D. Curriculum Vitae
    Alexandru Mihail Florian Tomescu, Ph.D. Curriculum Vitae Department of Biological Sciences 1 Harpst Street Humboldt State University Arcata, California 95521-8299 USA ORCID: 0000-0002-2351-5002 Phone: (+1) 707-826-3229 (office) / Fax: (+1) 707-826-3201 [email protected] Tomescu Lab Group web page: http://mihaibotany.online/#welcome Humboldt State University faculty web page: http://www.humboldt.edu/~biosci/faculty/tomescu.htm Academic Background Ph.D., Biological Sciences – Environmental and Plant Biology, Ohio University, Athens, Ohio. 2004. Advisor: Dr. Gar W. Rothwell. Late Ordovician - Early Silurian terrestrial biotas of Virginia, Ohio, and Pennsylvania: an investigation into the early colonization of land M.S., Geology (Paleobotany and Palynology), University of Bucharest, Romania. 1993. Advisor: Dr. Ovidiu Dragastan. The palynology of Pliocene coal-bearing deposits of Oltenia (SW Romania) Professional Experience and Appointments 2016-present – Professor, Department of Biological Sciences, Humboldt State University, Arcata CA. 2011-2016 – Associate Professor, Department of Biological Sciences, Humboldt State University, Arcata CA. 2005-2011 – Assistant Professor, Department of Biological Sciences, Humboldt State University, Arcata CA. 2005-2006 – Visiting Scholar (summer), Department of Geological Sciences, Indiana University, Bloomington IN. 1999-2004 – Graduate Teaching Assistant, Department of Environmental and Plant Biology, Ohio University, Athens OH. 1999-2003 – Scientific Imaging Facility technician, Ohio University, Athens OH. 1999 – Archeological field camp supervisor for the National Center for Pluridisciplinary Research, National History Museum of Romania. 1997-1999 – Research team leader, National Center for Pluridisciplinary Research, National History Museum of Romania. 1997-1998 – Adjunct lecturer, Department of Geology and Geophysics, University of Bucharest (Romania). 1996-1997 – Archeological field camp manager for the National Center for Pluridisciplinary Research, National History Museum of Romania.
    [Show full text]
  • Sphenopsid and Fern Remains from the Upper Triassic of Krasiejów (Sw Poland)
    Annales Societatis Geologorum Poloniae (2019), vol. 89: 307 – 316 doi: https://doi.org/10.14241/asgp.2019.13 SPHENOPSID AND FERN REMAINS FROM THE UPPER TRIASSIC OF KRASIEJÓW (SW POLAND) Grzegorz PACYNA Jagiellonian University in Kraków, Faculty of Biology, Institute of Botany, Department of Taxonomy, Phytogeography and Palaeobotany, Gronostajowa 3, 30-387 Kraków, Poland; e-mail: [email protected]; ORCID identifier 0000-0003-4365-3549 Pacyna, G., 2019. Sphenopsid and fern remains from the Upper Triassic of Krasiejów (SW Poland). Annales Soci- etatis Geologorum Poloniae, 89: 307 – 316. Abstract: Hydrophilic elements, such as sphenopsids and ferns, are rare in the flora of the Krasiejów site and they are poorly preserved. One sphenopsid species, Neocalamites merianii, was recognised. It is preserved as impres- sions, flattened casts, moulds and isolated leaves. So far, only one small fern specimen, determined asSphenopteris sp., has been found. These elements probably were transported to the site of deposition and did not grow where they were buried. Key words: Fossil plants, Neocalamites, Sphenopteris, Germanic Basin. Manuscript received 21 February 2019, accepted 11 August 2019 INTRODUCTION Recent discoveries of Upper Triassic outcrops in Silesia jer, 2005; Barycka; 2007; Konietzko-Meier and Wawro, (southwestern Poland) have yielded not only rich vertebrate 2007; Konietzko-Meier and Klein, 2013; Konietzko-Mei- material, but also new plant fossils (Dzik and Sulej, 2007; er and Sander, 2013; Konietzko-Meier et al., 2013, 2014; Pacyna, 2014). Plant macrofossils rarely are found in the Gruntmejer et al., 2016; Fortuny et al., 2017; Antczak and Upper Triassic of Poland (Reymanówna, 1986; Pacyna, Bodzioch, 2018b; Konietzko-Meier et al., 2018; Teschner 2014; Pacyna et al., 2017; Kustatscher et al., 2018) and et al., 2018), a gliding archosauromorph (Dzik and Sulej, hence are poorly known; such discoveries open new oppor- 2016), phytosaur (Butler et al., 2014), a rauisuchian (Sulej, tunities for study of them.
    [Show full text]