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Pennsylvania Amphibians & TTheh e SnappingS n a p p i n g T u r t l e by Andrew L. Shiels

Turtles are members of the Class Reptilia, Testudi- snapping are primarily nocturnal (active at night). nata. They are the oldest living reptiles, having evolved Most turtles are omnivorous—they feed on a wide variety more than 200 million years ago. of foods including insects, fi sh, small , carrion, Turtles have adapted to a variety of environments in- berries, aquatic vegetation and plants. cluding marine, estuarine, freshwater and terrestrial habi- Turtles vary in size from that are several inches in tats. They are found on every continent except Antarctica. length to those that are over 6 feet (sea turtles and giant tor- Although their may differ, all turtles share these toises). The common snapping is the largest Penn- same characteristics: 1) a shell, 2) they lay , and 3) sylvania turtle. Adult snappers can reach a shell length of they build on land to incubate their eggs. 12 inches and typically weigh between 15 and 45 pounds. The most unique feature of a turtle is its shell. The Not only can they reach outstanding weights, but tur- shell’s shape, color and pattern make it a readily used fea- tles also live longer than most other vertebrates. Some box ture for species identifi cation. The shell is divided into turtles have had recorded life spans of longer than a cen- an upper part, the carapace, and a lower part, the plas- tury. Generally, however, most Pennsylvania turtles live 20 tron. Both carapace and plastron are covered with scales to 40 years. called scutes. In addition to using the shell as an identifi er, In Pennsylvania, there are four turtle families compris- the plastron’s shape can be used in sex determination. In ing 12 genera and 15 species and . The families many species male turtles have a concave plastron; female are grouped into snapping turtles; musk and mud turtles; turtles have a fl at plastron. , marsh and box turtles; and softshell turtles Most Pennsylvania turtles are active from April to Oc- Snapping turtles, or “snappers,” are large freshwater tur- tober, spending the remaining portion of the year hiber- tles with a posterior serrated carapace reaching lengths of nating in burrows, in the mud, on the bottom of streams 12 inches The carapace varies from tan, brown or olive to or , or under piles of vegetation. Normal daily ac- black. The plastron is yellowish to tan and greatly reduced tivities include basking in the sun, foraging for food and in size. The head is large with massive jaws. The common resting. The majority of turtles are diurnal (active during snapper has a long tail with three triangular plates. Males daylight hours). However, mud turtles, musk turtles and usually have longer tails than females with the male cloacal opening (anus) located beyond the margin of the top shell. For more information In females, the cloaca does not extend beyond the shell Pennsylvania Amphibians & Reptiles, by Larry L. edge. Weights average in the 15- to 45- Shaffer, is the Commission’s full-color, illustrated pound range. An exceptional snapper book that provides details on Pennsylvania’s found in Wayne County was weighed salamanders, , toads, turtles, liz- by Commission staff in 2006 at over ards and . The 6-inch by 60 pounds. 9-inch, 166-page book includes Snapping turtles have been found each ’ characteris- in almost every kind of freshwater tics, identifi cation, range, in their range. They prefer habitat description, re- still or slow-moving water with production habits, and food soft mud or sand bottoms and requirements. To order this abundant vegetation. Snapping book, see the form in this issue turtles can also be found in on page 62, or order online on the brackish coastal waters. Snap- Commission’s web site, www.fi sh. ping turtles range statewide. state.pa.us. On the main page’s left side, Mating occurs from March to Novem- hold your cursor over “Pubs. & Products” ber. The nesting period for snapping turtles is usu- and click on “The Outdoor Shop.” ally three weeks (mid-May to mid-June). Snapping turtles

46 Pennsylvania Angler & Boater • July-August 2007 www.fi sh.state.pa.us T h e S n a p p i n g TurtleT u r t l e

dig nests in loose sand, loam, railroad beds, plant debris they can be seen fl oating just beneath the water’s surface or sawdust piles. Clutch sizes may vary from six to 100 with only eyes and nostrils protruding. eggs in each , but individual snappers usually lay 25 to Throughout their range, snapping turtles are hunted 50 eggs. The incubation period depends on temperature, for food for personal consumption or the restaurant trade. usually requiring 60 to 90 days. They are often killed because they are mistakenly believed As the name “snapping” turtle suggests, this species to be major predators of young waterfowl. Indeed, snap- uses it powerful jaws to capture prey and defend itself if ping turtles are opportunists and will consume many provoked. However, they will usually attempt to fl ee if types of food, both plant and . However, snapping disturbed. Snapping turtles are one of the most aquatic turtles are important scavengers of dead and decomposing turtle species, spending a majority of their time resting on aquatic animals. Research has shown that 80 percent or the bottom of deep pools or buried in the mud in shallow more of their diet typically consists of aquatic vegetation water with their eyes and nostrils exposed. During the day such as duckweed. Adult snappers can reach a shell length of 12 inches and typically weigh between 15 and 45 pounds.

illustration-Tom Duran Jr.

www.fi sh.state.pa.us Pennsylvania Angler & Boater • July-August July-August 2007 2007 47