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Fósiles Marinos Del Neógeno De Canarias (Colección De La ULPGC)

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Fósiles Marinos Del Neógeno De Canarias (Colección De La ULPGC) FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC). DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) Autor: Juan Francisco Betancort Lozano Las Palmas de Gran Canaria, 16 de enero de 2012 FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC): DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) D. Juan Luis Gómez Pinchetti Secretario del Departamento de Biología de la Universidad de Las Palmas de Gran Canaria. Certifica: Que el Consejo de Doctores del Departamento en sesion extraordinaria tomó el acuerdo de dar el consentimiento para su tramitación, a la tesis doctoral titulada "FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC). DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA)" presentada por el doctorando Juan Francisco Betancort Lozano y dirigida por el Dr. Joaquín Meco Cabrera. Y para que así conste, y a efectos de lo previsto en el Artº 73.2 del Reglamento de Estudios de Doctorado de esta Universidad, firmo la presente en las Palmas de Gran Canaria, a de Febrero de 2012. 5 FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC): DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) Las Palmas de Gran Canaria, a de Febrero de 2012 Programa de doctorado de Ecología y Gestión de los Recursos Vivos Marinos. Bienio: 2003-2005 Titulo de Tesis: Fósiles marinos del Neógeno de Canarias (Colección de la ULPGC). Dos neotipos, catálogo y nuevas aportaciones (Sistemática, Paleoecología y Paleoclimatología). Tesis Doctoral presentada por D Juan Francisco Betancort Lozano para obtener el grado de Doctor por la Universidad de Las Palmas de Gran Canaria, dirigida por el Dr. Joaquín Meco Cabrera El Director El Doctorando Joaquín Meco Cabrera Juan Fco Betancort Lozano 7 FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC): DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) Agradecimientos Son muchas las personas a las que debo tener en mente es este momento, es mucha la ayuda recibida y muchas las deudas de gratitud contraídas en el desarrollo de este trabajo, debo mencionar a D. Javier Ballester (Xavi), al Dr. Alejandro Lomoschitz Mora-Figueroa, a D. Antonio Cilleros, al Dr. Antonio Juan Gonzalez Ramos, al Dr. Paul Taylor Editor Jefe del Journal of Systematic Palaeontology y la Dra. Consuelo Sendino, ambos del departamento de Paleontología del Museo de Ciencias Naturales de Londres, al Dr. Christoph Mayr del Paläontologisches Museum München y a la Dra. Andrea Kourgli Directora del Abteilung Bibliotheken Naturhistorsches Museum Wien. Finalmente, debo mostrar agradecido al Ministerio de Medio Ambiente, con cuyo apoyo al proyecto Estudio de los Indicadores Paleoclimáticos de Canarias se comezaron estos trabajos. Debo sobre todo agradecer el apoyo, las infinitas correcciones y su paciencia al Dr. Joaquín Meco Cabrera. En la bibliografía y la revisión de los trabajos anteriores, se encuentra el fundamento mismo de este trabajo, sin el apoyo de la Biblioteca Universitaria, muy especialmente del equipo de la Biblioteca de Ciencias del Mar y del Servicio de Acceso al Documento, CIDOC, este trabajo de investigación no hubiera sido posible. Debo estar agradecido a quienes me han ayudado y apoyado; gracias a todos, ellos y ellas saben quienes son y saben lo mucho que les debo. Muchas gracias a todos. 9 FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC): DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) A nana, siempre estarás en mi corazón. 11 FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC): DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC). DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) Contenidos: Resumen 19 Abstract 20 1.- Objetivos y Planteamiento 23 2.- Estudios previos 24 2.1.- Definiciones 24 2.1.1.- Definiciones Taxonómicas y Sistemáticas 25 2.1.1.1.- Términos y estructura de los Braquiópodos 28 2.1.1.2.- Partes de la concha de los moluscos y términos anatómicos de interés 29 2.1.1.3.- Términos y estructura de los Crustáceos Crustáceos Cirrípedos 32 Crustáceos Decápodos 33 2.1.1.4.- Términos y estructura de los Equinodermos 35 2.1.1.5.- Ictiofauna: Anatomía dentaria y otolitos 38 2.1.1.6.- Introducción a las Icnología e Icnofacies 45 2.2.- Estudios anteriores sobre los depósitos de edad neógena de Canarias y su contenido faunistico 49 3.- Materiales y Metodología 57 4. - Fauna del Neógeno de Canarias 63 4.1.- Descripción de los niveles fósilíferos 63 4.2- Neotipos 79 13 FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC): DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) Trivia canariensis Rothpletz y Simonelli 1890 80 Marginella augustiformis Rothpletz y Simonelli 1890 82 4.3.- Otra fauna de interés. 85 Brachiopoda Terebratula sinuosa Brocchi 1814 90 Lacazella mediterranea Risso 1826 91 Mollusca, Gastropoda Haliotis volhynica Eichwald 1829 94 Diodora italica (Defrance 1820) 95 Fissurella cf. costicillatissima Sacco 1897 96 Patella ambrogii Lecointre 1952 97 Diloma patulum Brocchi 1814 98 Nerita emiliana Mayer 1872 99 Turritella subangulata Brocchi 1814 101 Petaloconchus intortus Lamarck 1818 102 Lemintina arenaria Linné 1766 103 Tenagodes anguinus Linné 1766 105 Cerithium taurinium Bellardi & Michelotti 1841 106 Cerithium vulgatum Bruguière 1789 106 Cerithium europaeum Mayer 1878 107 Scala (Fuscoscala) turtonis (Turton 1819) 108 Cheila cf. equestris Linné,1758 109 Rothpletzia rudista Simonelli 1890 110 Capulus cf. ungaricus Linné, 1758 111 Strombus coronatus Defrance 1827 112 Natica (Euspira) helicina Brocchi 1814 114 Trivia avellana Sowerby 1823 116 Zonaria elongata Brocchi 1814 117 Malea cf. orbiculata (Brocchi, 1814) 118 Cassis cf. mamillaris Grateloup 1833 119 Bursa scrobilator Linné, 1758 120 Gyrineum (Aspa) marginatum Martin 1777 120 Hexaplex rudis Borson 1821 122 Hexaplex trunculus Linné 1758 124 14 FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC): DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) Nucella plessisi Lecointre 1952 125 Columbella mercatoria Linné 1758 125 Nassa atlantica Mayer 1860 126 Hinia (Hinia) porrecta Bellardi, 1878 127 Olivella stricta Bellardi 1882 128 Solarium carocollatum Lamarck 1822 129 Ancilla glandiformis Lamarck 1822 130 Genota (Pseudotoma) praecedens (Bellardi 1877) 131 Conus mercati Brocchi 1814 132 Conus pelagicus Brocchi 1814 133 Conus virginalis Brocchi 1814 134 Conus brocchii Bronn 1831 134 Conus (Lautoconus) mediterraneus Hwass in Bruguiere, 1792 135 Gadinia garnoti Payraudeau 1826 136 Mollusca, Bivalvia Arca tetragona Poli 1795 137 Barbatia barbata Linné 1758 138 Barbatia clathrata Defrance 1816 140 Glycymeris insolita Mayer 1868 141 Glycymeris bimaculata Poli, 1795 142 Glycymeris violacescens Lamarck 1819 143 Pectunculus pilosus Linné 1767 146 Isognomon soldanii Deshayes 1836 147 Chlamys varia (Linné 1789) 149 Chlamys multistriata Poli 1795 150 Chlamys latissima Brocchi 1814 152 Chlamys opercularis Linné 1789 154 Chlamys (Manupecten) pesfelis Linné 1758 156 Hinnites ercolanianus Cocconi 1873 157 Pecten benedictus Lamarck 1819 159 Spondylus gaederopus Linné 1758 160 Anomia ephippium Linné 1758 162 Lima lima Linné 1758 163 Lopha hyotis Linné 1758 165 15 FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC): DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) Gryphaea virleti Deshayes 1832 166 Lucina leonina Basterot 1825 169 Ctena eburnea Gmelin 1788 170 Ctena decussata Da Costa 1843 170 Linga columbella Lamarck 1818 171 Loripes (Loripes) lacteus Linné 1758 173 Cyclocardia (Scalaricardita) scalaris Sowerby 1825 174 Cardita ajar Bruguière 1792 175 Cardita calyculata Linné 1758 175 Chama gryphoides Linné 1758 177 Chama gryphina Lamarck 1819 181 Chama laminosa Mollet 1854 183 Astarte (Astarte) omalii Jonkaire 1823 183 Acanthocardia tuberculata (Linné 1758) 184 Cardium clodiense Renieri 1804 185 Tellina compressa Brocchi 1814 186 Tellina cf. planata Linné 1758 187 Donax venustus Poli 1795 188 Venus multilamella Lamarck 189 Irus irus Linné, 1758 191 Callista chione Linné, 1758 192 Clausinella fasciata Da Costa 1778 193 Mollusca, Scaphopoda Fustiaria rubescens Deshayes, 1825 195 Arthropoda, Crustacea, Cirripedia Balanus concavus Bronn 1831 196 Balanus perforatus Bruguière 1789 197 Balanus spongicola Brown 1827 198 Chelonobia testudinaria Linnè 1767 199 Tetraclita cf. rubescens Darwin, 1854 200 16 FÓSILES MARINOS DEL NEÓGENO DE CANARIAS (COLECCIÓN DE LA ULPGC): DOS NEOTIPOS, CATÁLOGO Y NUEVAS APORTACIONES (SISTEMÁTICA, PALEOECOLOGÍA Y PALEOCLIMATOLOGÍA) Arthropoda, Crustacea, Decapoda Callianassa matsoni Rathbun 1935 202 Callianassa xavii n. sp. 204 Callianassa sp.1 206 Callianassa sp. 2 206 Callianassa sp. 3 206 Upogebia sp. 207 Scylla michelini Forskall, 1775 208 Cancer mecoi n. sp. 209 Eriphia sp. 215 Eriphia ajachensis n.sp. 216 Maja sp 219 Equinodermata Cidaris desmoulinsi Sismonda 1842 220 Clypeaster aegyptiacus Wright 1861 222 Clypeaster altus Leslie in Klein 1778 223 Clypeaster colloti Lambert1913 228 Rotula rumphii Klein 1734 229 Brissus unicolor (Leske, 1778) 231 Pisces Carcharodon megalodon Agassiz (1843) 234 Carcharodon carcharias Linné
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