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Revision of Hyalopale (Chrysopetalidae; Phyllodocida; Annelida
Revision of Hyalopale (Chrysopetalidae; Phyllodocida; Annelida) an amphi-Atlantic Hyalopale bispinosa species complex and five new species from reefs of the Caribbean Sea and Indo-Pacific Oceans Watson, Charlotte; Tilic, Ekin; Rouse, Greg W. Published in: Zootaxa DOI: 10.11646/zootaxa.4671.3.2 Publication date: 2019 Document version Publisher's PDF, also known as Version of record Document license: CC BY Citation for published version (APA): Watson, C., Tilic, E., & Rouse, G. W. (2019). Revision of Hyalopale (Chrysopetalidae; Phyllodocida; Annelida): an amphi-Atlantic Hyalopale bispinosa species complex and five new species from reefs of the Caribbean Sea and Indo-Pacific Oceans. Zootaxa, 4671(3), 339-368. https://doi.org/10.11646/zootaxa.4671.3.2 Download date: 27. sep.. 2021 Zootaxa 4671 (3): 339–368 ISSN 1175-5326 (print edition) https://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2019 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4671.3.2 http://zoobank.org/urn:lsid:zoobank.org:pub:99459D5F-3C35-4F7D-9768-D70616676851 Revision of Hyalopale (Chrysopetalidae; Phyllodocida; Annelida): an amphi-Atlantic Hyalopale bispinosa species complex and five new species from reefs of the Caribbean Sea and Indo-Pacific Oceans CHARLOTTE WATSON1, EKIN TILIC2 & GREG W. ROUSE2 1Museum & Art Gallery of the Northern Territory, Box 4646, Darwin, 0801 NT, Australia. E-mail: [email protected] 2Scripps Institution of Oceanography, UC San Diego, La Jolla, CA 92093-0202, USA. E-mail: [email protected] Abstract The formerly monotypic taxon, Hyalopale bispinosa Perkins 1985 (Chrysopetalinae), is comprised of a cryptic species complex from predominantly tropical embayments and island reefs of the Western Atlantic and Indo-Pacific Oceans. -
DEEP SEA LEBANON RESULTS of the 2016 EXPEDITION EXPLORING SUBMARINE CANYONS Towards Deep-Sea Conservation in Lebanon Project
DEEP SEA LEBANON RESULTS OF THE 2016 EXPEDITION EXPLORING SUBMARINE CANYONS Towards Deep-Sea Conservation in Lebanon Project March 2018 DEEP SEA LEBANON RESULTS OF THE 2016 EXPEDITION EXPLORING SUBMARINE CANYONS Towards Deep-Sea Conservation in Lebanon Project Citation: Aguilar, R., García, S., Perry, A.L., Alvarez, H., Blanco, J., Bitar, G. 2018. 2016 Deep-sea Lebanon Expedition: Exploring Submarine Canyons. Oceana, Madrid. 94 p. DOI: 10.31230/osf.io/34cb9 Based on an official request from Lebanon’s Ministry of Environment back in 2013, Oceana has planned and carried out an expedition to survey Lebanese deep-sea canyons and escarpments. Cover: Cerianthus membranaceus © OCEANA All photos are © OCEANA Index 06 Introduction 11 Methods 16 Results 44 Areas 12 Rov surveys 16 Habitat types 44 Tarablus/Batroun 14 Infaunal surveys 16 Coralligenous habitat 44 Jounieh 14 Oceanographic and rhodolith/maërl 45 St. George beds measurements 46 Beirut 19 Sandy bottoms 15 Data analyses 46 Sayniq 15 Collaborations 20 Sandy-muddy bottoms 20 Rocky bottoms 22 Canyon heads 22 Bathyal muds 24 Species 27 Fishes 29 Crustaceans 30 Echinoderms 31 Cnidarians 36 Sponges 38 Molluscs 40 Bryozoans 40 Brachiopods 42 Tunicates 42 Annelids 42 Foraminifera 42 Algae | Deep sea Lebanon OCEANA 47 Human 50 Discussion and 68 Annex 1 85 Annex 2 impacts conclusions 68 Table A1. List of 85 Methodology for 47 Marine litter 51 Main expedition species identified assesing relative 49 Fisheries findings 84 Table A2. List conservation interest of 49 Other observations 52 Key community of threatened types and their species identified survey areas ecological importanc 84 Figure A1. -
Annelida, Amphinomidae) in the Mediterranean Sea with an Updated Revision of the Alien Mediterranean Amphinomids
A peer-reviewed open-access journal ZooKeys 337: 19–33 (2013)On the occurrence of the firewormEurythoe complanata complex... 19 doi: 10.3897/zookeys.337.5811 RESEARCH ARTICLE www.zookeys.org Launched to accelerate biodiversity research On the occurrence of the fireworm Eurythoe complanata complex (Annelida, Amphinomidae) in the Mediterranean Sea with an updated revision of the alien Mediterranean amphinomids Andrés Arias1, Rômulo Barroso2,3, Nuria Anadón1, Paulo C. Paiva4 1 Departamento de Biología de Organismos y Sistemas (Zoología), Universidad de Oviedo, Oviedo 33071, Spain 2 Pontifícia Universidade Católica do Rio de Janeiro , Rio de Janeiro, Brazil 3 Museu de Zoologia da Unicamp, Campinas, SP, Brazil 4 Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro (UFRJ) , Rio de Janeiro, RJ, Brasil Corresponding author: Andrés Arias ([email protected]) Academic editor: C. Glasby | Received 17 June 2013 | Accepted 19 September 2013 | Published 30 September 2013 Citation: Arias A, Barroso R, Anadón N, Paiva PC (2013) On the occurrence of the fireworm Eurythoe complanata complex (Annelida, Amphinomidae) in the Mediterranean Sea with an updated revision of the alien Mediterranean amphinomids. ZooKeys 337: 19–33. doi: 10.3897/zookeys.337.5811 Abstract The presence of two species within the Eurythoe complanata complex in the Mediterranean Sea is reported, as well as their geographical distributions. One species, Eurythoe laevisetis, occurs in the eastern and cen- tral Mediterranean, likely constituting the first historical introduction to the Mediterranean Sea and the other, Eurythoe complanata, in both eastern and Levantine basins. Brief notes on their taxonomy are also provided and their potential pathways for introduction to the Mediterranean are discussed. -
Zootaxa, Loandalia (Polychaeta: Pilargidae)
Zootaxa 1119: 59–68 (2006) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA 1119 Copyright © 2006 Magnolia Press ISSN 1175-5334 (online edition) New species of Loandalia (Polychaeta: Pilargidae) from Queensland, Australia SHONA MARKS1 & SCOTT HOCKNULL2 1 S. A. Marks. CSIRO Marine Research, PO Box 120, Cleveland QLD 4163. [email protected]. 2 S. A. Hocknull. Queensland Museum, 122 Gerler Rd Hendra QLD 4711. [email protected] Abstract Two new species of Loandalia are described from Queensland, Australia. Loandalia fredrayorum sp. nov. is described from Moreton Bay, south eastern Queensland and is distinguished from all other species of Loandalia by the presence of singular palpostyles; uniramous parapodia at chaetiger 1; an emergent notopodial spine at chaetiger 9; neurochaetae numbering 20–24; ventral cirri begin on chaetiger 7 and the pygidium with two lateral papillae-like anal cirri. Loandalia gladstonensis sp. nov. is described from Gladstone Harbour, central eastern Queensland and is distinguished from all other species of Loandalia by the presence of bifid palpostyles; chaetiger 1 uniramous with remaining chaetigers biramous; an emergent notopodial spine from chaetiger 7–8; ventral cirri present from chaetiger 5 and neurochaetae numbering 5–6. Key words: Loandalia fredrayorum sp. nov., Loandalia gladstonensis sp.nov., Pilargidae, Queensland, Australia, new species, systematics. Introduction Saint-Joseph (1899) established the Pilargidae for the new species Pilargis verrucosa Saint-Joseph. Prior to this, pilargids had been placed in several different families including the Syllidae, Hesionidae and Polynoidae (Licher & Westheide 1994). Recent cladistic analyses of the Phyllodocida firmly recognise Pilargidae as a distinct clade (Glasby 1993; Pleijel & Dahlgren 1998). -
Neanthes Limnicola Class: Polychaeta, Errantia
Phylum: Annelida Neanthes limnicola Class: Polychaeta, Errantia Order: Phyllodocida, Nereidiformia A mussel worm Family: Nereididae, Nereidinae Taxonomy: Depending on the author, Ne- wider than long, with a longitudinal depression anthes is currently considered a separate or (Fig. 2b). subspecies to the genus Nereis (Hilbig Trunk: Very thick segments that are 1997). Nereis sensu stricto differs from the wider than they are long, gently tapers to pos- genus Neanthes because the latter genus terior (Fig. 1). includes species with spinigerous notosetae Posterior: Pygidium bears two, styli- only. Furthermore, N. limnicola has most form ventrolateral anal cirri that are as long as recently been included in the genus (or sub- last seven segments (Fig. 1) (Hartman 1938). genus) Hediste due to the neuropodial setal Parapodia: The first two setigers are unira- morphology (Sato 1999; Bakken and Wilson mous. All other parapodia are biramous 2005; Tusuji and Sato 2012). However, re- (Nereididae, Blake and Ruff 2007) where both production is markedly different in N. limni- notopodia and neuropodia have acicular lobes cola than other Hediste species (Sato 1999). and each lobe bears 1–3 additional, medial Thus, synonyms of Neanthes limnicola in- and triangular lobes (above and below), called clude Nereis limnicola (which was synony- ligules (Blake and Ruff 2007) (Figs. 1, 5). The mized with Neanthes lighti in 1959 (Smith)), notopodial ligule is always smaller than the Nereis (Neanthes) limnicola, Nereis neuropodial one. The parapodial lobes are (Hediste) limnicola and Hediste limnicola. conical and not leaf-like or globular as in the The predominating name in current local in- family Phyllodocidae. (A parapodium should tertidal guides (e.g. -
OREGON ESTUARINE INVERTEBRATES an Illustrated Guide to the Common and Important Invertebrate Animals
OREGON ESTUARINE INVERTEBRATES An Illustrated Guide to the Common and Important Invertebrate Animals By Paul Rudy, Jr. Lynn Hay Rudy Oregon Institute of Marine Biology University of Oregon Charleston, Oregon 97420 Contract No. 79-111 Project Officer Jay F. Watson U.S. Fish and Wildlife Service 500 N.E. Multnomah Street Portland, Oregon 97232 Performed for National Coastal Ecosystems Team Office of Biological Services Fish and Wildlife Service U.S. Department of Interior Washington, D.C. 20240 Table of Contents Introduction CNIDARIA Hydrozoa Aequorea aequorea ................................................................ 6 Obelia longissima .................................................................. 8 Polyorchis penicillatus 10 Tubularia crocea ................................................................. 12 Anthozoa Anthopleura artemisia ................................. 14 Anthopleura elegantissima .................................................. 16 Haliplanella luciae .................................................................. 18 Nematostella vectensis ......................................................... 20 Metridium senile .................................................................... 22 NEMERTEA Amphiporus imparispinosus ................................................ 24 Carinoma mutabilis ................................................................ 26 Cerebratulus californiensis .................................................. 28 Lineus ruber ......................................................................... -
The Namanereidinae (Polychaeta: Nereididae). Part 1, Taxonomy and Phylogeny
© Copyright Australian Museum, 1999 Records of the Australian Museum, Supplement 25 (1999). ISBN 0-7313-8856-9 The Namanereidinae (Polychaeta: Nereididae). Part 1, Taxonomy and Phylogeny CHRISTOPHER J. GLASBY National Institute for Water & Atmospheric Research, PO Box 14-901, Kilbirnie, Wellington, New Zealand [email protected] ABSTRACT. A cladistic analysis and taxonomic revision of the Namanereidinae (Nereididae: Polychaeta) is presented. The cladistic analysis utilising 39 morphological characters (76 apomorphic states) yielded 10,000 minimal-length trees and a highly unresolved Strict Consensus tree. However, monophyly of the Namanereidinae is supported and two clades are identified: Namalycastis containing 18 species and Namanereis containing 15 species. The monospecific genus Lycastoides, represented by L. alticola Johnson, is too poorly known to be included in the analysis. Classification of the subfamily is modified to reflect the phylogeny. Thus, Namalycastis includes large-bodied species having four pairs of tentacular cirri; autapomorphies include the presence of short, subconical antennae and enlarged, flattened and leaf-like posterior cirrophores. Namanereis includes smaller-bodied species having three or four pairs of tentacular cirri; autapomorphies include the absence of dorsal cirrophores, absence of notosetae and a tripartite pygidium. Cryptonereis Gibbs, Lycastella Feuerborn, Lycastilla Solís-Weiss & Espinasa and Lycastopsis Augener become junior synonyms of Namanereis. Thirty-six species are described, including seven new species of Namalycastis (N. arista n.sp., N. borealis n.sp., N. elobeyensis n.sp., N. intermedia n.sp., N. macroplatis n.sp., N. multiseta n.sp., N. nicoleae n.sp.), four new species of Namanereis (N. minuta n.sp., N. serratis n.sp., N. stocki n.sp., N. -
Polychaete Worms Definitions and Keys to the Orders, Families and Genera
THE POLYCHAETE WORMS DEFINITIONS AND KEYS TO THE ORDERS, FAMILIES AND GENERA THE POLYCHAETE WORMS Definitions and Keys to the Orders, Families and Genera By Kristian Fauchald NATURAL HISTORY MUSEUM OF LOS ANGELES COUNTY In Conjunction With THE ALLAN HANCOCK FOUNDATION UNIVERSITY OF SOUTHERN CALIFORNIA Science Series 28 February 3, 1977 TABLE OF CONTENTS PREFACE vii ACKNOWLEDGMENTS ix INTRODUCTION 1 CHARACTERS USED TO DEFINE HIGHER TAXA 2 CLASSIFICATION OF POLYCHAETES 7 ORDERS OF POLYCHAETES 9 KEY TO FAMILIES 9 ORDER ORBINIIDA 14 ORDER CTENODRILIDA 19 ORDER PSAMMODRILIDA 20 ORDER COSSURIDA 21 ORDER SPIONIDA 21 ORDER CAPITELLIDA 31 ORDER OPHELIIDA 41 ORDER PHYLLODOCIDA 45 ORDER AMPHINOMIDA 100 ORDER SPINTHERIDA 103 ORDER EUNICIDA 104 ORDER STERNASPIDA 114 ORDER OWENIIDA 114 ORDER FLABELLIGERIDA 115 ORDER FAUVELIOPSIDA 117 ORDER TEREBELLIDA 118 ORDER SABELLIDA 135 FIVE "ARCHIANNELIDAN" FAMILIES 152 GLOSSARY 156 LITERATURE CITED 161 INDEX 180 Preface THE STUDY of polychaetes used to be a leisurely I apologize to my fellow polychaete workers for occupation, practised calmly and slowly, and introducing a complex superstructure in a group which the presence of these worms hardly ever pene- so far has been remarkably innocent of such frills. A trated the consciousness of any but the small group great number of very sound partial schemes have been of invertebrate zoologists and phylogenetlcists inter- suggested from time to time. These have been only ested in annulated creatures. This is hardly the case partially considered. The discussion is complex enough any longer. without the inclusion of speculations as to how each Studies of marine benthos have demonstrated that author would have completed his or her scheme, pro- these animals may be wholly dominant both in num- vided that he or she had had the evidence and inclina- bers of species and in numbers of specimens. -
Annelida: Polychaeta) from the NE Atlantic, with Some Further Records of Related Species
European Journal of Taxonomy 539: 1–21 ISSN 2118-9773 https://doi.org/10.5852/ejt.2019.539 www.europeanjournaloftaxonomy.eu 2019 · Ravara A. et al. This work is licensed under a Creative Commons Attribution Licence (CC BY 4.0). Research article urn:lsid:zoobank.org:pub:17F463A6-5663-4E82-8FD4-759ACD25D2F2 Description of a new genus and species of Chrysopetalidae (Annelida: Polychaeta) from the NE Atlantic, with some further records of related species Ascensão RAVARA 1,*, M. Teresa AGUADO 2, Clara F. RODRIGUES 3, Luciana GÉNIO 4 & Marina R. CUNHA 5 1,3,4,5 Departamento de Biologia & CESAM, Universidade de Aveiro, 3810-193 Aveiro, Portugal. 2 Animal Evolution and Biodiversity, Johann-Friedrich-Blumenbach Institute for Zoology and Anthropology, Georg-August-Universität, 37073 Göttingen, Germany. 2 Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Departamento de Biología, Universidad Autó noma de Madrid, Cantoblanco, 28049 Madrid, Spain. * Corresponding author: [email protected] 2 Email: [email protected] 3 Email: [email protected] 4 Email: [email protected] 5 Email: [email protected] 1 urn:lsid:zoobank.org:author:677F8AB4-7FD3-483A-A047-C4BD5A6A449D 2 urn:lsid:zoobank.org:author:F9D27435-8C88-4785-A980-A3B07C41DD22 3 urn:lsid:zoobank.org:author:D54DAA7A-BE73-4E37-B5A1-760517AF1BA5 4 urn:lsid:zoobank.org:author:3E6080EC-1249-459C-B368-D621FB641D47 5 urn:lsid:zoobank.org:author:553A98B5-0AE0-424F-9ED5-EC50F129519C Abstract. Five chrysopetalid species are reported from samples collected at bathyal depths in three NE Atlantic regions: the Bay of Biscay, the Horseshoe Abyssal Plain and the Gulf of Cadiz. -
Oxygen, Ecology, and the Cambrian Radiation of Animals
Oxygen, Ecology, and the Cambrian Radiation of Animals The Harvard community has made this article openly available. Please share how this access benefits you. Your story matters Citation Sperling, Erik A., Christina A. Frieder, Akkur V. Raman, Peter R. Girguis, Lisa A. Levin, and Andrew H. Knoll. 2013. Oxygen, Ecology, and the Cambrian Radiation of Animals. Proceedings of the National Academy of Sciences 110, no. 33: 13446–13451. Published Version doi:10.1073/pnas.1312778110 Citable link http://nrs.harvard.edu/urn-3:HUL.InstRepos:12336338 Terms of Use This article was downloaded from Harvard University’s DASH repository, and is made available under the terms and conditions applicable to Other Posted Material, as set forth at http:// nrs.harvard.edu/urn-3:HUL.InstRepos:dash.current.terms-of- use#LAA Oxygen, ecology, and the Cambrian radiation of animals Erik A. Sperlinga,1, Christina A. Friederb, Akkur V. Ramanc, Peter R. Girguisd, Lisa A. Levinb, a,d, 2 Andrew H. Knoll Affiliations: a Department of Earth and Planetary Sciences, Harvard University, Cambridge, MA, 02138 b Scripps Institution of Oceanography, University of California San Diego, La Jolla, CA, 92093- 0218 c Marine Biological Laboratory, Department of Zoology, Andhra University, Waltair, Visakhapatnam – 530003 d Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA, 02138 1 Correspondence to: [email protected] 2 Correspondence to: [email protected] PHYSICAL SCIENCES: Earth, Atmospheric and Planetary Sciences BIOLOGICAL SCIENCES: Evolution Abstract: 154 words Main Text: 2,746 words Number of Figures: 2 Number of Tables: 1 Running Title: Oxygen, ecology, and the Cambrian radiation Keywords: oxygen, ecology, predation, Cambrian radiation The Proterozoic-Cambrian transition records the appearance of essentially all animal body plans (phyla), yet to date no single hypothesis adequately explains both the timing of the event and the evident increase in diversity and disparity. -
Eteone Lighti Order: Phyllodocida, Phyllodociformia
Phylum: Annelida Class: Polychaeta, Errantia Eteone lighti Order: Phyllodocida, Phyllodociformia A paddleworm Family: Phyllodocidae, Eteoninae Taxonomy: The genus Eteone was revised tentacular cirri (Fig. 2). into three genera (Eteone, Hypereteone, Parapodia: Uniramous with neuropodia only. Mysta) by Wilson (1988) based on anal cirri All but first segment with a flat triangular dor- morphology and the presence and location sal cirrus, about as wide as long (Fig. 3), of proboscis papillae. Thus, E. lighti is these become longer and narrower posterior- sometimes referred to as Hypereteone lighti. ly. The ventral cirrus has a broad base taper- While the presence of three major groups ing to a blunt tip and is shorter than the neu- are apparent, splitting Eteone into these ropodial lobe (Fig. 3). Note: parapodium genera has not been recognized by most should be examined in side view to check for authors and E. lighti is the name most com- flatness, inflatedness, etc. monly seen (Pleijel 1991; Blake 1992; Blake Setae (chaetae): Setae are compound 1997; Blake and Ruff 2007). (Phyllodocidae, Blake 1975b) and consist of long, fine, colorless spinigers (Hartman 1968) Description (Figs. 4a,b). Size: Individuals to 30 mm in length and 1– Eyes/Eyespots: Two eyespots are present 1.5 mm in width (Hartman 1968). A 25-mm on posterior third of the prostomium (Fig. 2a). long Coos Bay specimen weighed 0.17 g Anterior Appendages: Prostomium bears (wet weight, Baker et al. 1970). two pairs of short, conical antennae and ap- Color: Pale or white, deep yellow dorsal pendages on the first segment include two transverse stripes (Hartman 1968) and dor- pairs of short and slender tentacular cirri sal cirri with deep yellow tips. -
Of Polychaetes (Annelida: Polychaeta) from the Atlantic and Mediterranean Coasts of the Iberian Peninsula: an Annotated Checklist E
López and Richter Helgol Mar Res (2017) 71:19 DOI 10.1186/s10152-017-0499-6 Helgoland Marine Research ORIGINAL ARTICLE Open Access Non‑indigenous species (NIS) of polychaetes (Annelida: Polychaeta) from the Atlantic and Mediterranean coasts of the Iberian Peninsula: an annotated checklist E. López* and A. Richter Abstract This study provides an updated catalogue of non-indigenous species (NIS) of polychaetes reported from the conti- nental coasts of the Iberian Peninsula based on the available literature. A list of 23 introduced species were regarded as established and other 11 were reported as casual, with 11 established and nine casual NIS in the Atlantic coast of the studied area and 14 established species and seven casual ones in the Mediterranean side. The most frequent way of transport was shipping (ballast water or hull fouling), which according to literature likely accounted for the intro- ductions of 14 established species and for the presence of another casual one. To a much lesser extent aquaculture (three established and two casual species) and bait importation (one established species) were also recorded, but for a large number of species the translocation pathway was unknown. About 25% of the reported NIS originated in the Warm Western Atlantic region, followed by the Tropical Indo West-Pacifc region (18%) and the Warm Eastern Atlantic (12%). In the Mediterranean coast of the Iberian Peninsula, nearly all the reported NIS originated from warm or tropi- cal regions, but less than half of the species recorded from the Atlantic side were native of these areas. The efects of these introductions in native marine fauna are largely unknown, except for one species (Ficopomatus enigmaticus) which was reported to cause serious environmental impacts.