DOCSLIB.ORG

Download PDF (1164K)

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Download PDF (1164K) © 2018 The Japan Mendel Society Cytologia 83(4): 407–414 Karyological Investigation of Sixteen Cirsium Mill. (Asteraceae, Cardueae) Taxa from Turkey Yasar Kiran* Firat University, Science Faculty, Biology Department, Elazig, Turkey Received June 27, 2018; accepted July 18, 2018 Summary The chromosome number and karyotype analysis of sixteen taxa of genus Cirsium Mill. distributed in Turkey were analyzed. This taxa are, Cirsium steriolepis Petr., C. bulgaricum DC., C. polycephalum DC., C. trachylepis Boiss., C. sommieri Petr., C. osseticum (Adams) Petrak, C. leucocephalum (Willd.) Spreng ssp. tenuilobum (K. Koch) Greuter, C. haussknechtii Boiss., C. ekimianum Yildiz & Dirmenci, C. handaniae Yildiz, Dirmenci & Arabacı, C. davisianum Kit Tan & Sorger, C. hypoleucum DC., C. pseudopersonata Boiss & Bal. ssp. pseudopersonata, C. pseudopersonata Boiss. & Bal. ssp. kuznezowianum (Somm. & Lev.) Petrak, C. canum (L.) All., C. obvallatum (Bieb.) Fischer, and nine of them are endemic species (C. steriolepis, C. polycephalum, C. trachylepis, C. sommieri, C. leucocephalum ssp. tenuilobum, C. davisianum, C. handaniae, C. ekimianum and C. pseudopersonata ssp. pseudopersonata). The somatic chromosome number of all studied taxa were deter- meined as 2n=2x=34. Total chromosome lengths varied 68.00 to 147.92 µm among species. Karyotype asymme- try indices other parameters were also calculated. Karyotype analysis indicated that Cirsium taxa generally have median (m and M) chromosomes. In addition, only C. davisianum species has one submedian (sm) chromosome. The chromosome number and karyotype of C. polycephalum, C. handaniae, C. davisianum and C. ekimianum are newly reported. Key words Asteraceae, Cirsium, Karyotype, Chromosome number, Turkey. The genus Cirsium Mill. (thistle) of the family Astera- 102 are either very rare or doubtful (Bures et al. 2004). ceae includes perennial, biennial, and annual members Chromosome studies of the Turkish Cirsium species re- and comprises more than 250 species distributed in Eu- ported that the taxa have 2n=32, 34, 60, and 68 diploid rope North Africa Asia and North and Central America. chromosome numbers (Ozcan et al. 2008, 2011, Yuksel Native species of Cirsium range from sea level to alpine et al. 2013). and occur in many habitats including swamps, mead- Cirsium is a taxonomically complex genus with ows, forests, prairies, sands, screes, steppes, and deserts many hybrids. Several multidisciplinary studies such (Charadze 1963, Davis and Parris 1975, Petrak 1979, as anatomy, palynology, molecular and karyological are Kadereit and Jeffrey 2007). needed to clarify the taxonomy of the genus. Therefore, According to recent studies of the Turkish Cirsium, the karyology of Turkish Cirsium species is currently be- the genus is represented in Turkey by 66 species (78 ing studied in order to clarify their taxonomy and make taxa), of which 27 species (32 taxa) of them are endemic contributions to other multidisciplinary studies on the species, belonging to three sections; Epitrachys DC., genus. Cirsium DC., and Cephalonoplos (Neck.) DC. (Davis and Parris 1975, Davis et al. 1988, Guner et al. 2000, Materials and methods Daskin et al. 2006, Yildiz and Dirmenci 2008, Yildiz et al. 2009a, b, 2011, Arabacı and Dirmenci 2011, Yildiz Plant materials 2012, Yuksel et al. 2013). In Turkey, this 66 species oc- All plant specimens were collected from natural cur, mainly distributed in east and north-east Anatolia, habitats during the fruiting season in 2006 and 2007 as with the main center of diversity between Trabzon and a part of a taxonomic revision of the genus Cirsium in Artvin provinces (Davis 1971). Turkey. The examined taxa, localities and voucher num- Approximately 69% of species of the genus Cirsium bers are given in Table 1. The specimens were identified are diploid 2n=34, 10% are tetraploid 2n=68, and 5% according to the pertinent reference works given in the are 2n=30 or 2n=32, while several chromosome num- literature (Davis and Parris 1975, Davis et al. 1988, bers such as 2n=16, 18, 20, 22, 24, 26, 28, 26, 51, and Guner et al. 2000, Yildiz et al. 2009a, Yildiz 2012). The voucher specimens were deposited at the Balıkesir and * Corresponding author, e-mail: ykiran@firat.edu.tr Inönü University Herbarium, Turkey. DOI: 10.1508/cytologia.83.407 408 Y. Kiran Cytologia 83(4) Table 1. Localities and voucher numbers of the studied Cirsium taxa. Taxa Locality Collecter (voucher number) C. steriolepis A1 Tekirdag: Inecik, Ormanli-Guzel Village, Quercus area, 500–600 m, B. Yildiz & T. Dirmenci (BY 16507) 01.08.2007 C. bulgaricum B6 Yozgat: Akdag Mine, Olukozu Village, 1 km west, 1200 m, 11.08.2006 B.Yildiz & T. Dirmenci (BY 16232) C. polycephalum A1 Kırklareli: Saray, between Beyceler-Sinekli, 3–4 km, Quercus area, 170 m, B. Yildiz & T. Dirmenci (BY 16530) 02.08.2007 C. trachylepis A8 Rize: Ikizdere, Bas Village, Cicekli plateau, stony and meadow area, B. Yildiz & T. Arabacı (BY 16663) 2100–2300 m, 15.09.2007 C. sommieri B8 Bingol: Between Karlıova-Bingol, 15 km, 1767 m, 11.08.2007 T. Dirmenci & T. Arabacı (TD 3515) C. osseticum A7 Trabzon: Between Caykara, Karacakaya (Karacam) Village-Soganlı Gorge, B. Yildiz (BY 16755) 1900–2000 m, 21.09.2007 C. leucocephalum ssp. tenuilobum A8 Erzurum: Askale, Kop Mountain, steppe, 1750 m, 21.09.2007 B. Yildiz (BY 16746) C. haussknechtii B7 Malatya: Akcadag, Dedeyazı Village, Canakci location, 1450 m, 10.08.2007 T. Dirmenci & T. Arabacı (TD 3503) C. ekimianum A8 Erzurum: Askale, Kop Mountain, steppe, 1750 m, 21.09.2007 B. Yildiz (BY 16745) C. handaniae A9 Ardahan: Between Turkozu-Posof 3 km, 1500 m, 19.09.2007 B. Yildiz & T. Arabacı (BY 16731) C. davisianum B8 Erzurum: Askale, Kop Mountain, stony area, 2450 m, 04.08.2008 B. Yildiz (BY 16850) C. hypoleucum A2 Bursa: Between Keles-Inegöl, 15 km, Pinus forest, 1300 m, 21.08.2007 B. Yildiz, T. Arabacı (BY 16593) C. pseudopersonata ssp. pseudopersonata B7 Bayburt: Soganlı Mountain, steppe, 2200 m, 20.09.2007 B. Yildiz (BY 16754) C. pseudopersonata ssp. kuznezowianum A9 Artvin: Between Borcka-Macahel 34 km, forest openings, 1450 m, 16.09. B. Yildiz & T. Arabacı (BY 16682) 2007 C. canum A3 Bolu: Between Abant Lake-Akcaalan Village, water edge, 1250 m, 22.08.2007 B. Yildiz & T. Arabacı (BY 16610) C. obvallatum A8 Artvin: Ardanuc, Bilbilan Plateau, 2500 m, 19.09.2007 B. Yildiz & T. Arabacı (BY 16724) Chromosome analysis asymmetry for karyotypes (A) (Huziwara 1962, Arano Approximately 100 seeds belong to 10 plant speci- 1963, Greilhuber and Speta 1976, Watanabe et al. 1999, mens were used for each taxon. Thirty roots of each spe- Venora et al. 2002). While the intrachromosomal asym- cies were used from seeds germinated on a moist filter metry index (A1) and the interchromosomal asymmetry paper in petri dishes at 25°C. The actively growing root index (A2) was calculated according to the formula pro- tips were pretreated with 0.05% colchicine for 3–3.5 h at posed by Romero Zarco (1986). Also, karyotype sym- room temperature. The material was fixed with Carnoy metry nomenclature was determined based on Stebbins' fixative (1 : 3 glacial acetic acid–absolute ethanol) for classification (Stebbins 1971). at least 24 h at 4°C, hydrolyzed in 1 M HCl at 60°C for 15 min, then rinsed in tap water for 3–5 min, stained in Results Feulgen staining or reaction for 1 h, and mounted in 45% acetic acid (Elci 1982). Digital microphotographs from C. steirolepis Petrak is an endemic species and in at least seven well-spread metaphase plates were taken montane woods (Davis and Parris 1975). The somatic using an Olympus BX51 microscope and were recorded chromosome number of this species was found to be with an Olympus Camedia C-4000 digital camera. 2n=2x=34. The chromosome length ranges 5.00 to The chromosome numbers of the species were iden- 8.93 µm and the total karyotype length was 113.9 µm. It tified and checked using the Flora of Turkey (Davis consists of 17 m chromosomes. Arm ratios are 1.07–1.43 and Parris 1975) and supplements (Davis et al. 1988, and relative lengths are 4.38–7.84% (Table 2). Somatic Guner et al. 2000). Also, the online chromosome num- metaphase chromosomes and idiograms are given in ber databases, Index to Plant Chromosome Numbers Figs. 1a and 2a. (http://www.tropicos.org/Project/IPCN) and Index to C. bulgaricum DC. İs growing in Pinus vegatation by Chromosome Numbers in Asteraceae (http://www.lib. a roadside, in Abies area, siliceous substrate, between kobe-u.ac.jp/infolib/meta_pub/G0000003asteraceaesre- altitudes of 1200 and 1800 m asl (Davis and Parris sult-en) were checked. 1975). The somatic chromosome number was found to The number of somatic chromosomes, chromosome be 2n=2x=34. The chromosome length ranges 3.60 to length range, haploid chromosome length, arm ratio, 6.40 µm and the total karyotype length was 83.80 µm. It relative length, and karyotype formula were deter- consists of 1M and 16 m chromosomes. Arm ratios are mined. Karyotype formula was determined based on 1.00–1.50 and relative lengths are 4.29–7.63% (Table centromere position (Levan et al. 1964). The ideograms 2). Somatic metaphase chromosomes and idiograms are of these taxa were arranged in decreasing lengths ac- given in Figs. 1b and 2b. cording to the metaphase chromosome size (Martin C. polycephalum DC. is an endemic species that et al. 2009). Karyotype asymmetry was determined by grows on dry places by roadsides, in fields, dry hills, using various techniques including total form percent- open places, near sea level, at altitudes of 50 m (Da- age (TF%), percentage karyotype asymmetry index vis and Parris 1975). The seeds were collected from (AsK%), index of karyotype symmetry (Syi), index of Kırklareli Province (Table 1). The somatic chromosome chromosome size resemblance (Rec) and the degree of number of this species was found to be 2n=2x=34.
Load more

Recommended publications
  • Thistles of Colorado
    Thistles of Colorado About This Guide Identification and Management Guide Many individuals, organizations and agencies from throughout the state (acknowledgements on inside back cover) contributed ideas, content, photos, plant descriptions, management information and printing support toward the completion of this guide. Mountain thistle (Cirsium scopulorum) growing above timberline Casey Cisneros, Tim D’Amato and the Larimer County Department of Natural Resources Weed District collected, compiled and edited information, content and photos for this guide. Produced by the We welcome your comments, corrections, suggestions, and high Larimer County quality photos. If you would like to contribute to future editions, please contact the Larimer County Weed District at 970-498- Weed District 5769 or email [email protected] or [email protected]. Front cover photo of Cirsium eatonii var. hesperium by Janis Huggins Partners in Land Stewardship 2nd Edition 1 2 Table of Contents Introduction 4 Introduction Native Thistles (Pages 6-20) Barneyby’s Thistle (Cirsium barnebyi) 6 Cainville Thistle (Cirsium clacareum) 6 Native thistles are dispersed broadly Eaton’s Thistle (Cirsium eatonii) 8 across many Colorado ecosystems. Individual species occupy niches from Elk or Meadow Thistle (Cirsium scariosum) 8 3,500 feet to above timberline. These Flodman’s Thistle (Cirsium flodmanii) 10 plants are valuable to pollinators, seed Fringed or Fish Lake Thistle (Cirsium 10 feeders, browsing wildlife and to the centaureae or C. clavatum var. beauty and diversity of our native plant americanum) communities. Some non-native species Mountain Thistle (Cirsium scopulorum) 12 have become an invasive threat to New Mexico Thistle (Cirsium 12 agriculture and natural areas. For this reason, native and non-native thistles neomexicanum) alike are often pulled, mowed, clipped or Ousterhout’s or Aspen Thistle (Cirsium 14 sprayed indiscriminately.
    [Show full text]
  • Literature Cited
    Literature Cited Robert W. Kiger, Editor This is a consolidated list of all works cited in volumes 19, 20, and 21, whether as selected references, in text, or in nomenclatural contexts. In citations of articles, both here and in the taxonomic treatments, and also in nomenclatural citations, the titles of serials are rendered in the forms recommended in G. D. R. Bridson and E. R. Smith (1991). When those forms are abbre- viated, as most are, cross references to the corresponding full serial titles are interpolated here alphabetically by abbreviated form. In nomenclatural citations (only), book titles are rendered in the abbreviated forms recommended in F. A. Stafleu and R. S. Cowan (1976–1988) and F. A. Stafleu and E. A. Mennega (1992+). Here, those abbreviated forms are indicated parenthetically following the full citations of the corresponding works, and cross references to the full citations are interpolated in the list alphabetically by abbreviated form. Two or more works published in the same year by the same author or group of coauthors will be distinguished uniquely and consistently throughout all volumes of Flora of North America by lower-case letters (b, c, d, ...) suffixed to the date for the second and subsequent works in the set. The suffixes are assigned in order of editorial encounter and do not reflect chronological sequence of publication. The first work by any particular author or group from any given year carries the implicit date suffix “a”; thus, the sequence of explicit suffixes begins with “b”. Works missing from any suffixed sequence here are ones cited elsewhere in the Flora that are not pertinent in these volumes.
    [Show full text]
  • Milk Thistle
    Forest Health Technology Enterprise Team TECHNOLOGY TRANSFER Biological Control BIOLOGY AND BIOLOGICAL CONTROL OF EXOTIC T RU E T HISTL E S RACHEL WINSTON , RICH HANSEN , MA R K SCH W A R ZLÄNDE R , ER IC COO M BS , CA R OL BELL RANDALL , AND RODNEY LY M FHTET-2007-05 U.S. Department Forest September 2008 of Agriculture Service FHTET he Forest Health Technology Enterprise Team (FHTET) was created in 1995 Tby the Deputy Chief for State and Private Forestry, USDA, Forest Service, to develop and deliver technologies to protect and improve the health of American forests. This book was published by FHTET as part of the technology transfer series. http://www.fs.fed.us/foresthealth/technology/ On the cover: Italian thistle. Photo: ©Saint Mary’s College of California. The U.S. Department of Agriculture (USDA) prohibits discrimination in all its programs and activities on the basis of race, color, national origin, sex, religion, age, disability, political beliefs, sexual orientation, or marital or family status. (Not all prohibited bases apply to all programs.) Persons with disabilities who require alternative means for communication of program information (Braille, large print, audiotape, etc.) should contact USDA’s TARGET Center at 202-720-2600 (voice and TDD). To file a complaint of discrimination, write USDA, Director, Office of Civil Rights, Room 326-W, Whitten Building, 1400 Independence Avenue, SW, Washington, D.C. 20250-9410 or call 202-720-5964 (voice and TDD). USDA is an equal opportunity provider and employer. The use of trade, firm, or corporation names in this publication is for information only and does not constitute an endorsement by the U.S.
    [Show full text]
  • Morphological and Biological Parameters of the Knapweed Nematode, Subanguina Picridis 1
    154 Journal of Nematology, Volume 18, No. 2, April 1986 weeds. 43. Acroptilon (Centaurea) repens (L.) DC. Ca- repens (L.) DC., Russian Knapweed (Compositae). Pp. nadian Journal of Plant Science 60:993-1004. 105-110 in J. S. Kelleher and M. A. Hulme, eds. 19. Watson, A. K. 1986. Host range of, and plant Biological control programmes against insects and reaction to, Subanguina picridis. Journal of Nematol- weeds in Canada 1969-1980. Slough: Common- ogy 18:112-120. wealth Agriculture Bureaux. 20. Watson, A. K. 1986. Morphological and bio- 22. Watson, A. K., and J. D. Shorthouse. 1979. logical parameters of the knapweed nematode, Suban- Gall formation on Cirsium arvense by Ditylenchus dip- guina _picridis.Journal of Nematology 18:154-158. saci. Journal of Nematology 11:16-22. 21. Watson, A. K., and P. Harris. 1984. Acroptilon Journal of Nematology 18(2):154-158. 1986. © The Society of Nematologists 1986. Morphological and Biological Parameters of the Knapweed Nematode, Subanguina picridis 1 A. K. WATSON 2 Abstract: Specimens of the knapweed nematode Subanguina picridis (Kirjanova) Brzeski obtained from different host plants were highly variable in measurement and structure. This variability refutes the validity of six Subanguina species attacking plants in the Asteraceae. Key words: Acroptilon repens (Russian knapweed), host specificity, Subanguina picridis (knapweed nematode), taxonomy, variability. The gall forming nematode Subanguina MATERIALS AND METHODS picridis (Kirjanova, 1944) Brzeski, 1981, Galls were obtained from A. repens, Cen- syn.: Paranguina picridis (Kirjanova, 1944) taurea diffusa Lam., Cynara sco$ymus L., and Kirjanova & Ivanova, 1968, has been eval- Onopordum acanthium L. plants inoculated uated in Canada as a potential biological with S.
    [Show full text]
  • Species List For: Valley View Glades NA 418 Species
    Species List for: Valley View Glades NA 418 Species Jefferson County Date Participants Location NA List NA Nomination and subsequent visits Jefferson County Glade Complex NA List from Gass, Wallace, Priddy, Chmielniak, T. Smith, Ladd & Glore, Bogler, MPF Hikes 9/24/80, 10/2/80, 7/10/85, 8/8/86, 6/2/87, 1986, and 5/92 WGNSS Lists Webster Groves Nature Study Society Fieldtrip Jefferson County Glade Complex Participants WGNSS Vascular Plant List maintained by Steve Turner Species Name (Synonym) Common Name Family COFC COFW Acalypha virginica Virginia copperleaf Euphorbiaceae 2 3 Acer rubrum var. undetermined red maple Sapindaceae 5 0 Acer saccharinum silver maple Sapindaceae 2 -3 Acer saccharum var. undetermined sugar maple Sapindaceae 5 3 Achillea millefolium yarrow Asteraceae/Anthemideae 1 3 Aesculus glabra var. undetermined Ohio buckeye Sapindaceae 5 -1 Agalinis skinneriana (Gerardia) midwestern gerardia Orobanchaceae 7 5 Agalinis tenuifolia (Gerardia, A. tenuifolia var. common gerardia Orobanchaceae 4 -3 macrophylla) Ageratina altissima var. altissima (Eupatorium rugosum) white snakeroot Asteraceae/Eupatorieae 2 3 Agrimonia pubescens downy agrimony Rosaceae 4 5 Agrimonia rostellata woodland agrimony Rosaceae 4 3 Allium canadense var. mobilense wild garlic Liliaceae 7 5 Allium canadense var. undetermined wild garlic Liliaceae 2 3 Allium cernuum wild onion Liliaceae 8 5 Allium stellatum wild onion Liliaceae 6 5 * Allium vineale field garlic Liliaceae 0 3 Ambrosia artemisiifolia common ragweed Asteraceae/Heliantheae 0 3 Ambrosia bidentata lanceleaf ragweed Asteraceae/Heliantheae 0 4 Ambrosia trifida giant ragweed Asteraceae/Heliantheae 0 -1 Amelanchier arborea var. arborea downy serviceberry Rosaceae 6 3 Amorpha canescens lead plant Fabaceae/Faboideae 8 5 Amphicarpaea bracteata hog peanut Fabaceae/Faboideae 4 0 Andropogon gerardii var.
    [Show full text]
  • OLFS Plant List
    Checklist of Vascular Plants of Oak Lake Field Station Compiled by Gary E. Larson, Department of Natural Resource Management Trees/shrubs/woody vines Aceraceae Boxelder Acer negundo Anacardiaceae Smooth sumac Rhus glabra Rydberg poison ivy Toxicodendron rydbergii Caprifoliaceae Tatarian hone ysuckle Lonicera tatarica* Elderberry Sambucus canadensis Western snowberry Symphoricarpos occidentalis Celastraceae American bittersweet Celastrus scandens Cornaceae Redosier dogwood Cornus sericea Cupressaceae Eastern red cedar Juniperus virginiana Elaeagnaceae Russian olive Elaeagnus angustifolia* Buffaloberry Shepherdia argentea* Fabaceae Leadplant Amorpha canescens False indigo Amorpha fruticosa Siberian peashrub Caragana arborescens* Honey locust Gleditsia triacanthos* Fagaceae Bur oak Quercus macrocarpa Grossulariaceae Black currant Ribes americanum Missouri gooseberry Ribes missouriense Hippocastanaceae Ohio buckeye Aesculus glabra* Oleaceae Green ash Fraxinus pennsylvanica Pinaceae Norway spruce Picea abies* White spruce Picea glauca* Ponderosa pine Pinus ponderosa* Rhamnaceae Common buckthorn Rhamnus cathartica* Rosaceae Serviceberry Amelanchier alnifolia Wild plum Prunus americana Hawthorn Crataegus succulenta Chokecherry Prunus virginiana Siberian crab Pyrus baccata* Prairie rose Rosa arkansana Black raspberry Rubus occidentalis Salicaceae Cottonwood Populus deltoides Balm-of-Gilead Populus X jackii* White willow Salix alba* Peachleaf willow Salix amygdaloides Sandbar willow Salix exigua Solanaceae Matrimony vine Lycium barbarum* Ulmaceae
    [Show full text]
  • Canada Thistle Author: Various Cirsium Arvense (L.) Scop.; Parks Affected: All Breea Arvensis (L.) Lessing
    COLORADO STATE PARKS BEST MANAGEMENT PRACTICES WEED PROFILE Date Created: April 25, 2003 Revised: April 1, 2005 Canada thistle Author: Various Cirsium arvense (L.) Scop.; Parks Affected: All Breea arvensis (L.) Lessing Family: Asteraceae (Sunflower) Other Names: field thistle, Californian thistle USDA Code: CIAR4 Keys to Identification: Legal Status: Colorado Noxious List B (top ten worst). • Purple flowers form in clusters of 1-5 per branch. Identification • The floral bracts of Canada Growth form: Perennial forb. thistle are spineless. Flower: Flower heads are white to purple and borne in clusters • Small heads, vanilla scent of 1-5 per branch, with a strong vanilla scent. Heads are only about 1cm in diameter. Seeds/Fruit: One-seeded fruits (achenes) are straw or light brown in color, straight or slightly curved (Moore 1975). Leaves: Leaves are spiny, alternate, oblong or lance-shaped, with the base leaves stalkless and clasping, or extended down along the stem. Stems: Mature plants range from 2-4 feet in height. Roots: Canada thistle has two types of roots, horizontal and vertical. The horizontal roots produce numerous shoots, while vertical roots store water and nutrients in their many small branches. Seedling: Early spring growth appears as rosettes with spiny- tipped, wavy leaves. Other: The floral bracts of Canada thistle are spineless. Similar Species Exotics: Bull thistle (Cirsium vulgare); flower bracts are somewhat tapered and covered with spines. Scotch thistle (Onopordum acanthium); stems appear the have wings, floral bracts are covered with spines. Plumeless thistle (Carduus acanthoides); floral bracts are covered with sharp spines. Musk thistle (Carduus nutans); floral bracts are broad with spiny tips.
    [Show full text]
  • Field Release of the Hoverfly Cheilosia Urbana (Diptera: Syrphidae)
    USDA iiillllllllll United States Department of Field release of the hoverfly Agriculture Cheilosia urbana (Diptera: Marketing and Regulatory Syrphidae) for biological Programs control of invasive Pilosella species hawkweeds (Asteraceae) in the contiguous United States. Environmental Assessment, July 2019 Field release of the hoverfly Cheilosia urbana (Diptera: Syrphidae) for biological control of invasive Pilosella species hawkweeds (Asteraceae) in the contiguous United States. Environmental Assessment, July 2019 Agency Contact: Colin D. Stewart, Assistant Director Pests, Pathogens, and Biocontrol Permits Plant Protection and Quarantine Animal and Plant Health Inspection Service U.S. Department of Agriculture 4700 River Rd., Unit 133 Riverdale, MD 20737 Non-Discrimination Policy The U.S. Department of Agriculture (USDA) prohibits discrimination against its customers, employees, and applicants for employment on the bases of race, color, national origin, age, disability, sex, gender identity, religion, reprisal, and where applicable, political beliefs, marital status, familial or parental status, sexual orientation, or all or part of an individual's income is derived from any public assistance program, or protected genetic information in employment or in any program or activity conducted or funded by the Department. (Not all prohibited bases will apply to all programs and/or employment activities.) To File an Employment Complaint If you wish to file an employment complaint, you must contact your agency's EEO Counselor (PDF) within 45 days of the date of the alleged discriminatory act, event, or in the case of a personnel action. Additional information can be found online at http://www.ascr.usda.gov/complaint_filing_file.html. To File a Program Complaint If you wish to file a Civil Rights program complaint of discrimination, complete the USDA Program Discrimination Complaint Form (PDF), found online at http://www.ascr.usda.gov/complaint_filing_cust.html, or at any USDA office, or call (866) 632-9992 to request the form.
    [Show full text]
  • List of Plants for Great Sand Dunes National Park and Preserve
    Great Sand Dunes National Park and Preserve Plant Checklist DRAFT as of 29 November 2005 FERNS AND FERN ALLIES Equisetaceae (Horsetail Family) Vascular Plant Equisetales Equisetaceae Equisetum arvense Present in Park Rare Native Field horsetail Vascular Plant Equisetales Equisetaceae Equisetum laevigatum Present in Park Unknown Native Scouring-rush Polypodiaceae (Fern Family) Vascular Plant Polypodiales Dryopteridaceae Cystopteris fragilis Present in Park Uncommon Native Brittle bladderfern Vascular Plant Polypodiales Dryopteridaceae Woodsia oregana Present in Park Uncommon Native Oregon woodsia Pteridaceae (Maidenhair Fern Family) Vascular Plant Polypodiales Pteridaceae Argyrochosma fendleri Present in Park Unknown Native Zigzag fern Vascular Plant Polypodiales Pteridaceae Cheilanthes feei Present in Park Uncommon Native Slender lip fern Vascular Plant Polypodiales Pteridaceae Cryptogramma acrostichoides Present in Park Unknown Native American rockbrake Selaginellaceae (Spikemoss Family) Vascular Plant Selaginellales Selaginellaceae Selaginella densa Present in Park Rare Native Lesser spikemoss Vascular Plant Selaginellales Selaginellaceae Selaginella weatherbiana Present in Park Unknown Native Weatherby's clubmoss CONIFERS Cupressaceae (Cypress family) Vascular Plant Pinales Cupressaceae Juniperus scopulorum Present in Park Unknown Native Rocky Mountain juniper Pinaceae (Pine Family) Vascular Plant Pinales Pinaceae Abies concolor var. concolor Present in Park Rare Native White fir Vascular Plant Pinales Pinaceae Abies lasiocarpa Present
    [Show full text]
  • Plant Community Composition and Structure Monitoring for Agate Fossil Beds National Monument 2011-2015 Summary Report
    National Park Service U.S. Department of the Interior Natural Resource Stewardship and Science Plant Community Composition and Structure Monitoring for Agate Fossil Beds National Monument 2011-2015 Summary Report Natural Resource Report NPS/NGPN/NRR—2016/1198 ON THIS PAGE Photograph of riparian long-term monitoring plot 261 at Agate Fossil Beds National Monument, 2015. Photograph courtesy of the National Park Service. ON THE COVER Photograph of plant community monitoring at Agate Fossil Beds National Monument, 2015. Photograph courtesy of the National Park Service. Plant Community Composition and Structure Monitoring for Agate Fossil Beds National Monument 2011-2015 Summary Report Natural Resource Report NPS/NGPN/NRR—2016/1198 Isabel W. Ashton Christopher J. Davis National Park Service Northern Great Plains Inventory & Monitoring Network 231 East St. Joseph Street Rapid City, SD 57701 April 2016 U.S. Department of the Interior National Park Service Natural Resource Stewardship and Science Fort Collins, Colorado The National Park Service, Natural Resource Stewardship and Science office in Fort Collins, Colorado, publishes a range of reports that address natural resource topics. These reports are of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, conservation and environmental constituencies, and the public. The Natural Resource Report Series is used to disseminate comprehensive information and analysis about natural resources and related topics concerning lands managed by the National Park Service. The series supports the advancement of science, informed decision-making, and the achievement of the National Park Service mission. The series also provides a forum for presenting more lengthy results that may not be accepted by publications with page limitations.
    [Show full text]
  • Asteraceae – Aster Family
    ASTERACEAE – ASTER FAMILY Plant: herbs (annual or perennial), some shrubs, rarely vines or trees. Stem: Root: Often with tubers, rhizomes, stolons, or fleshy roots Leaves: mostly simple, some compound, alternate or opposite, rarely whorled. Flowers: flower head supported by an involucre (whorl of green bracts or phyllaries); each head composed of small flowers (composite) of flat ray-like (ligulate) flowers on the outside (ray flowers) and central tube-like flowers (disk flowers) – some species may have only one or the other. Calyx absent or modified into hairs, bristles, scales or a crown (pappus); 5 stamens (syngenesious -united by anthers); 5 united petals (sympetalous), receptacle may also have hairs or bristles. Both pappus and receptacle hairs/bristles may be used in ID. Fruit: achene (small, one-seeded, inferior ovule, 2 carpels, hard shell fruit) often with persisting crowned pappus which helps with seed dispersal. Other: Very large family, divided into sub-families and tribes, once named Compositae; 1-2,000 genera, 20,000+ species. Dicotyledons Group WARNING – family descriptions are only a layman’s guide and should not be used as definitive ASTERACEAE – ASTER FAMILY Straggler Daisy; Calyptocarpus vialis Less. (Introduced) Nodding [Plumeless] Thistle; Carduus nutans L. (Introduced) Garden Cornflower [Bachelor’s Button; Blue Bottle]; Centaurea cyanus L. (Introduced) Spotted Knapweed; Centaurea stoebe L. ssp. micranthos (Gugler) Hayek (Introduced) Woody [Bush] Goldenrod; Chrysoma pauciflosculosa (Michx.) Greene (Solidago pauciflosculosa) Green and Gold; Chrysogonum virginianum L. Soft Goldenaster; Chrysopsis pilosa Nutt. Chicory; Cichorium intybus L. (Introduced) Tall Thistle; Cirsium altissimum (L.) Hill Canada Thistle; Cirsium arvense (L.) Scop. Soft [Carolina] Thistle; Cirsium carolinianum (Walter) Fernald & B.G.
    [Show full text]
  • The Herbaceous Vascular Plants of Blackacre Preserve a Preliminary List
    The Herbaceous Vascular Plants of Blackacre Preserve A Preliminary List December 3, 2010 Submitted to: Kentucky State Nature Preserves Commission Submitted by: William E. Thomas Herbarium Indiana University Southeast Photo: Spiked Crested Coralroot by Richard Lyons 1 Scope The aim of this survey was to compile a rough list of herbaceous vascular plant species on the below described tract and was conducted from July 11, 2010 through the end of the growing season. In addition any extensive populations of invasive alien species were noted. Locale Description The Blackacre Preserve website states that the property consists of 170 acres in eastern Jefferson County Kentucky. It is the authors understanding that some additional acreage (size?) was appended to the southern border of the original 170 acre tract. The property is located at 3200 Tucker Station Rd. The tract is bordered on all sides by housing and urban areas; a railroad track runs along the north border. The terrain is of mostly gentle slopes with some wooded areas and open fields formerly used for pasture or crops. There are several ponds on the property; a limestone glade area constitutes the northeast corner of the tract. A small creek flows east to west across the tract north of the center. There are numerous foot trails, some designated and some rogue. An old section of Mann’s Lick road runs northward about midway in the tract. Map #1 from the Blackacre Preserve website provides a general layout of this tract. Map #2 is a topographic map with a NAD83 UTM 16 grid superimposed and the foot trails plotted in various colors.
    [Show full text]