Occurrence of Saga Pedo (Orthoptera: Tettigoniidae) in the Czech Republic: Review of Faunistic Data

Home , Saga (bush cricket), Saga pedo

North-Western Journal of Zoology Vol. 6, No. 2, 2010, pp.218-224 P-ISSN: 1584-9074, E-ISSN: 1843-5629 Article No.: 061121

Occurrence of Saga pedo (Orthoptera: Tettigoniidae) in the Czech Republic: review of faunistic data

Jaroslav HOLUŠA1,*, Petr KOČÁREK2 and Robert VLK3

1. Department of Forest Protection and Game Management, Faculty of Forestry and Wood Sciences, Czech University of Life Sciences, Kamýcká 1176, CZ-165 21 Praha 6-Suchdol, Czech Republic 2. Department of Biology and Ecology, University of Ostrava, Chittussiho 10, CZ-710 00 Ostrava 2, Czech Republic 3. Department of Biology, Faculty of Education, Masaryk University, Poříčí 7, CZ-603 00 Brno, Czech Republic * Corresponding Author, J. Holuša, E-mail: [email protected]

Abstract. Predatory Katydid Saga pedo was recorded in the Czech Republic for the first time in 1900. Since then, it has been found in five areas: on Kamenný vrch (hill) at Kurdějov, in Pouzdřanská step- Kolby (national nature reserve), on Pavlovské vrchy (hills), near the Bulhary village, and in the environs of Znojmo (town). During the first half of the twentieth century, only eight specimens of S. pedo were observed or collected in the Czech Republic, and these were in three areas in the southeastern part of the country: Pavlovské vrchy hills, Pouzdřanská step-Kolby NNR, and environs of Znojmo. From 1950-1990, one locality (Kamenný vrch hill at Kurdějov) was added, but only eight individuals were observed during the same period for the entire country. From 1988-2009, 32 individuals were reported by incidental observers. In intensive surveys conducted during 2006-2009, 477 nymphs and 75 adults were found in the area of Pavlovské vrchy (hills). In the area of Pavlovské vrchy, S. pedo was detected in nine localities, i.e., the insect was widespread across the entire area of the hills, from Dívčí hrad ruin in the north to Turold hill in the south. During the surveys of 2006- 2009, however, S. pedo was not detected in any new locality, and in spite of intensive surveying, S. pedo was not found in two localities where it had previously been reported.

Key words: Saga pedo, faunistics, distribution, Czech Republic.

Introduction 1980, 1988, 1995, Labler 1933, Povolný 1994, Redtenbacher 1900). Currently, S. pedo is consi- Predatory Katydid Saga pedo (Pallas, 1771) is a dered a critically endangered species (Holuša parthenogenetic pentaploid (Dutrillaux et al. & Kočárek 2005). Between 2006 and 2009, the 2009) species distributed from the Pyrenees first systematic surveys for S. pedo in the Czech throughout the whole of southeastern Europe Republic were conducted and the methods and to western Siberia and the Caucasus (Kenyeres results are reported in this publication. et al. 2002). This is a threatened species and is This study summarizes all known findings listed in Appendix II of the Bern Convention of S. pedo in the Czech Republic, from the first and Annex IV of the Habitats Directive of published finding in 1900 up to the year 2009. Natura 2000. There are two subspecies, but only the nominotypical S. pedo pedo (Pallas, 1771) occurs in the Czech Republic (Kočárek et Material and methods al. 2005). Predatory Katydid Saga pedo inhabits All literature data concerning S. pedo were summarized only the warmest parts of southern Moravia, and all important national or provincial museum where it has been recorded at only few locali- collections were reviewed. The latter included collections ties (Ginter 1924a, 1924b, 1925, 1971, Chládek at Moravské zemské museum (Brno), Slezské zemské

©NwjZ, Oradea, Romania, 2010 North-West J Zool, 6, 2010 www.herp-or.uv.ro/nwjz Oradea, Romania Occurrence of Saga pedo in the Czech Republic: review of faunistic data 219 muzeum (Opava), Národní muzeum (Praha), Muzeum Koutný, 20.v.1923, 1n, leg. Stejskal, coll. jihovýchodní Moravy (Zlín), Ostravské Muzeum Museum of south-eastern Moravia in Zlín, (Ostrava), and Východočeské muzeum (Hradec Králové). revid. J. Holuša (Ginter 1924a, 1924b, Ginter Data were also obtained by appeals to the scientific community (Holuša et al. 2007, Holuša, Kočárek 2007, 1925); Pausram (=Pouzdřany), 1A, 1936, leg. Kočárek, Holuša 2008). Finally, the Internet was searched Plitschk, coll. Moravian provincial museum thoroughly for references to S. pedo (photographs, re- (Brno), det. I. Malenovský; 15.vii.1946, 2n, leg. marks about accidental findings, etc.). B. Dobšík (Chládek 1988); 5.vi.2008, 1n (Dvořák During the period of 2006-2009, intensive surveys were carried out at all localities where S. pedo had been et al. 2008); 2.vi.2009, 1n, observ. J. Holuša.

reported and also at some other localities potentially (iii) Area of the Pavlovské vrchy hills (7165) suitable for S. pedo occurrence. Surveys were done by Pavlovské vrchy hills/Pavlovské kopce groups of 2-4 people and between 10 and 16 o’clock on clear, sunny days. Localities were systematically hills/Pálava hills (not specified), 28.vii.1932, surveyed, and the vegetation was pushed aside with help 1n, 1A, leg. Hoffer (Labler 1933), 1n, coll. of 1-m-long sticks. Many localities were surveyed more National museum Prague, but leg. Kempný than once, and the dates of the surveys and locations indicated on the locality label, revid. M. surveyed are listed in Table 1. A survey of a particular Fikáček; 19.vii.1957, 1A, leg. Hoberlandt, coll. location on a particular date required 1 to 2 hours. One plot (10000-50000 m2) was surveyed for most localities National museum Prague, revid. M. Fikáček; but two to four plots were surveyed for larger localities. 30.ix.1977, 1A, leg. B. Dobšík, det. et coll. P. Surveys were not done in the environs of the town of Kočárek. Znojmo because the locality in Redtenbacher (1900) is Děvín hill, 2.vii.1952, 1n, 1A, unknown unclear and because the area had already been surveyed for orthopteran insects in another study (Holuša 2003). collector, coll. National museum Prague, revid. Names of localities were taken from recent maps M. Fikáček (Kempný 1953: 3n found Mařan (Anonymous 2005, www.mapy.cz), and names of hills, and Hoffer in 1952); without indicating a rocks, and ruins were used: Děvín hill (549m asl), Obora concrete finding (Chládek 1995); under posts of hill (483m asl), Pálava hill (461m asl), Stolová hora hill the power line, 27.v.1998, 3 n, obsev. J. Hájek; (458m asl), Turold hill (385m asl), Martinka rock, Trůn rock, Dívčí hrad ruin, and Kaple ruin of chapel. Several S. 30m SW from broadcaster on Děvín hill, pedo were not precisely localized because different 14.ix.2002, 1ex; c. 150m from broadcaster on collectors used different names for the same general area; the edge of the slope, 11.x.2003, 1ex, 6.x. 2004, these names were Pavlovské kopce (hills), Pavlovské 1ex, M. Škorpík observ. (M. Škorpík pers. vrchy (hills), and Pálava hills. comm.); 28.v.2002, 1n, 4.-10.vii.2003, 5ex Abbreviations: n = nymph; A = adult (female); ex = (Chytil 2003); hill-top part above steep slope in specimen without specification of stage; NNR = National its SE part, August 1999-2008, altogether 8 ex, Nature Reserve; NR = Nature Reserve; (7066) = code of observ. Kavka (Kavka in litt.); 6.viii.2008, 1A, standard grid square (11 x 12 km) of faunistic grid mapping in the Czech Republic. photo Z. Souček, det. J. Holuša, P. Kočárek (http://zdeneksoucek.blog.cz/0808/za-kudlan kou-a-sagou-na-palavu); under posts of the Results and Discussion power line, 1998-2002, 2ex, obsev. J. Matuška (Matuška pers. comm); 26.viii.2005, Holuša List of records of Saga pedo from the observ.; by the view-point at Soutěska, 3n, Czech Republic 29.iv.2007, foto J. Korba, det. J. Holuša, P.

(i) NR Kamenný vrch at Kurdějov (7066) Kočárek (http://www.equi-shop.cz/rubriky/ Kamenný vrch (hill) at Kurdějov, 9.vi.1979, mdiskuse/show.php?tema=180397&r=2&m=9 1n (Chládek 1980). 97&page=14); SE slopes of the Děvín hill, 8.vi.2008, 2n, photo A. Poledníček, det. J. (ii) NNR Pouzdřanská step-Kolby (7065) Holuša (Poledníček in litt., http://www.biolib. Pouzdřanská step-Kolby, 8.ix.1922, 1A, leg. cz/cz/taxon/id150/).

North-West J Zool, 6, 2010 220 Holuša, J. et al.

Table 1. Names and locations of plots, and dates on which the plots were surveyed.

Latitude Longitude Locality (plot number) 2006 2007 2008 2009 ( N ) ( E ) Anenský vrch NM 48°49'33.7" 16°37'5.6" - 7.VIII. - - Bavory 48°49‘32.5" 16°37‘45.5" - 7.VIII. - - Čejkovické Špidláky NR 48°54'58.3" 16°57'49.6" 6.IX. - - - Dívčí hrad ruin - plot 1 48°52'19.4" 16°39'21.5" - 10.VI.; 7.VIII. 2.V.; 6.VIII. 10.V. Dívčí hrad ruin - plot 2 48°52'28.4" 16°39'39.9" - - 2.V.; 6.VIII. 10.V. Dívčí hrad ruin - plot 3 48°52'14.9" 16°39'30.1" - - 1.V.;7.VIII. 10.V. Dívčí hrad ruin - plot 4 48°52'9.4" 16°39'23.2" - - 1.V.;6.VIII. - Děvín hill - plot 1 48°52'8.3" 16°38'56.1" 17.,24.IX. 10.VI.; 7.VIII. 2.V.;6.VIII. 13.VI. Děvín hill - plot 2 48°52'8.3" 16°38'59.7" 17.,24.IX. 10.VI.; 7.VIII. 2.V.;6.VIII. 13.VI. Děvín hill - plot 3 48°52'13.3" 16°39'10.5" 17.,24.IX. 7.VIII. 2.V.;6.VIII. 13.VI. Děvín hill - plot 4 48°51'54.0" 16°38'36.1" 2.,5.VII.; 4.V.;10.VI.; 1.V.;6.VIII. 13.VI. 8.,9.VIII. 6.VIII. Dunajovické kopce NR 48°50'48.6" 16°33'19.3" - - - 8.V. Kamenný vrch hill (Kurdějov) 48°57'57.4" 16°45'8.1" 16.IX. - 10.VI. - Kaple - ruin of chappel 48°51'32.2" 16°38'10.2" 10.,15.VIII. 8.VIII. 1.V.;7.VIII. - Kočičí kámen NM 48°49'49.7" 16°38'12.6" - 6.VIII. - - Martinka hill - plot 1 48°51'56.9" 16°38'9.4" - - 1.,17,29.V.; 9.,30.V.; 14.VI.; 5.VII.; 19.VI.;12.VII. 5.VIII. Martinka hill - plot 2 48°51'58.2" 16°38'12.9" 10.,11.,18.VIII. 12.V.;6.VIII.; 1.V., 17.V., 9.V. ;17.,18.IX. 8.VIII. 29.V., 14.VI., 5.VII., 5.VIII. Milovická stráň NM 48°50'54.4" 16°41'33.4" - - 6.VIII. - Obora hill 48°51'52.3" 16°38'18.2" 24.VII;10.,14., 12.V.;6.VIII. 1.V.;5.VIII. 9.V. 18.,VIII. Pálava (Kotel) hill 48°51'23.4" 16°38'32.9" 2.VII. - 5.VIII. 12.VII. Pouzdřanská step-Kolby NNR - pl. 1 48°56'30.9" 16°38'30.3" 16.IX. - 6.VIII. - Pouzdřanská step-Kolby NNR - pl. 2 48°56'51.5" 16°38'43.5" - - - 31.V. Růžový kopec NM 48°49'18.4" 16°37'27.0" - 7.VIII. - - Skalka hill (Sedlec) 48°46'19.8" 16°40'34.6" 18.VIII. - - - Stolová hora hill - plot 1 48°50'16.2" 16°38'16.5" 2.,5.VII.; 9.- 9.VIII. 30.IV.;4.VIII. 14.VI. 11.VIII. Stolová hora hill - plot 2 48°50'7.2" 16°38'20.1" 9.-11.,15., 10.VI.;9.VIII. 30.IV.;4.VIII. 14.VI.;5.VII. 19.VIII. Svatý kopeček hill - plot 2 48°48'21.7" 16°39'1.2" 9.VIII. - - 5.VII. Trůn hill (=Obří palec) 48°51'50.8" 16°38'24.4" - 8.VIII. 1.V.;5.VIII. 9.V. Turold hill - plot 1 48°49'0.5" 16°38‘24.0" 9.,19.VIII. 7.VIII. 1.V.;7.VIII. 8.V. Turold hill - plot 2 48°49‘0.5" 16°38'18.6" 9.,19.VIII. 7.VIII. 1.V.;7.VIII. 8.V. Turold hill - plot 3 48°48'54.1" 16°38'21.0" 9.,19.VIII. 7.VIII. 1.V.;7.VIII. 8.V. Turold hill - plot 4 48°48'54.1" 16°38'23.9" 9.,19.VIII. 7.VIII. 1.V.;7.VIII. 8.V. Výchoz NM (Čejč) 48°56'14.1" 16°59'4.8" 6.IX. - - - Zimarky hill (Velké Pavlovice) 48°52'31.7" 16°53'9.3" 6.IX. - - -

North-West J Zool, 6, 2010 Occurrence of Saga pedo in the Czech Republic: review of faunistic data 221

Dívčí hrad ruin, 30.viii.1974, 1A, leg. Z. placed onto the locality of NR Havranické Navrátil let., coll. Moravian provincial mu- vřesoviště during autumn 2005 (Kavka in litt.). seum (Brno), det. I. Malenovský; 27.ix.1988, 1A, During the year 2007 as well as 2008, several leg. D. Povolný (Povolný 1992, 1994); without individuals were recorded there (Kavka in indicating a concrete finding (Chládek 1995); litt.). This amateur illegal introduction is not 1995, 1A, obsev. J. Troneček (Troneček pers. well documented. comm.); 1km N of the ruin, 29.v.2007, 1n, excursion of the Univerzity of Ostrava, det. P. So far, the current report summarizes all histo- Kočárek; under the ruin, 13.ix.2007, 1A, foto P. rical S. pedo findings but does not include Molitor, det. J. Holuša, P. Kočárek. several unpublished findings found in Kotel (former NNR comprising studied different museum collections. Almost all of localities Martinka rock, Trůn rock, zříc. Kaple those findings were known to Ginter (1977). ruin of chapel, Obora hill and Pálava hill), During the first half of twentieth century, only among Pinus nigra trees, 1993, 1A, obsev. J. eight individuals of S. pedo were observed or Troneček (Troneček pers. comm.); without collected in the Czech Republic, and these were indicating a concrete finding (Chládek 1995). collected in three areas in the southeastern part Obora hill, 3.x.2007, photo P. Muckstein, of the country (Pavlovské vrchy hills, Pouz- det. J. Holuša (Muckstein in litt., http://www. dřanská step-Kolby NNR, and the environs of biolib. cz/cz/taxon/id150). Znojmo). From 1950 to 1990, one locality (Ka- Stolová hora hill (=Tabulová hora hill), menný vrch at Kurdějov) was added, and S edge of the plateau, 2001, 1n, observ. T. observations of only eight individuals were Kuras (Kuras pers. comm.); above the quarry recorded for all locations. After 1987, the with the ranch; 28.viii.2005, 1ex, observ. R. Vlk number of observations of S. pedo increased. (Vlk et al. 2006; Vlk pers. comm.); in N From 1988 to 2009, 32 S. pedo individuals were projection above Perná village (at the locality reported by incidental observers. These repre- called “Kučerova mulda”), 11.viii.2006, 1A, sented casual findings in almost all cases; only photo volunteers of international camp INEX a few of these observations were reported by SDA, organized by the Centre of ecological entomologists, and most were reported by education Pálava ZO ČSOP 56/15, det. J. vertebrate zoologists, botanists, or nature pho- Holuša; 28.v.2008, 1n, R. Gabriš; 50m down- tographers. Accidental observations of S. pedo wards the slope from the barrier railing, 29.v. by naturalists during their field researches 2008, 1 n, photo M. Sovíček, det. J. Holuša seem to be quite general (see e.g. Zamfirescu et (http://www.biolib.cz/cz/taxon/id150). al. (2009)). Trůn rock, 1994, 1A, obsev. J. Troneček In the surveys conducted from 2006 to (pers. comm.). 2009, the occurrence of S. pedo was confirmed for only two areas (Pavlovské vrchy hills and (iv) Surroundings of Bulhary village (7166) at Pouzdřanská step-Kolby NNR) in altitudes Surroundings of Bulhary village, during 220-540m asl (Fig. 1), what corresponds with sweeping along the road from Bulhary village French data (Darcemont et al. 2009). The to Lednice town, 18.v.2002, 1n, photo J. largest numbers of S. pedo specimens (477 Dvořák, det. J. Holuša. nymphs and 75 adults) were found in nine (v) Environs of Znojmo town (7162) localities representing the area of the Pavlov- Znojmo, 1ex (Redtenbacher 1900). ské vrchy hills (Table 2). Saga pedo is wide- Note: Eggs of one keeping female of S. spread in this area, and analysis of S. pedo pedo, native at the locality of Děvín hill, were population trends in this area has been pub-

North-West J Zool, 6, 2010 222 Holuša, J. et al.

Figure 1. Occurrence of Saga pedo in the Czech Republic.

Table 2. Detection of Saga pedo specimens in nine localities representing the area of the Pavlovské vrchy hills (n.s.- not surveyed).

2006 2007 2008 2009 Locality (no. of plots) nymphs adults nymphs adults nymphs adults nymphs adults Děvín hill (4) 3 0 3 4 1 1 15 n.s. Dívčí hrad ruin (4) n.s. n.s. 3 2 7 7 50 n.s. Kaple - ruin of chappel n.s. 2 0 0 n.s. n.s. n.s. n.s. Pálava (Kotel) hill 0 0 n.s. n.s. 0 3 0 n.s. Martinka hill (2) n.s. 12 42 5 169 9 125 1 Obora hill n.s. 11 1 0 1 0 9 n.s. Stolová hora hill (2) n.s. 6 0 0 1 1 17 n.s. Trůn hill (=Obří palec) n.s. n.s. n.s. 3 2 5 19 n.s. Turold hill (4) n.s. 2 0 0 7 1 2 n.s. Totals 3 33 49 14 188 27 237 1 36 63 215 238

lished (Holuša et al. 2009). During the current 2008 but had not been recorded in the previous study, however, no other locality was discove- 62 years. Given that locality Pouzdřanská step- red, and despite intensive and repeated sur- Kolby NNR is frequently explored and that S. veys, S. pedo was not observed in two localities pedo is an attractive insect, it is possible that S. (name the localities) where it had been pre- pedo was absent from this locality after 1946 viously reported (see Table 1 and 2). and was only recently reintroduced (Dvořák et At the locality Pouzdřanská step-Kolby al. 2008). The presence of S. pedo at this locality NNR, the occurrence of S. pedo was recorded in was confirmed in 2009, indicating the possible

North-West J Zool, 6, 2010 Occurrence of Saga pedo in the Czech Republic: review of faunistic data 223 presence of a reproducing population of S. pedo whole area of these hills, from the Dívčí hrad in the Pouzdřanská step-Kolby NNR. ruin in the north to the Turold hill in the south. The detection of S. pedo at NR Havranické vřesoviště (near Znojmo town) in 2005, 2007, and 2008 is probably the consequence of introduction in 2005. Authors of this contribution Acknowledgements. The research was supported by the Agency for Nature Conservation and Landscape Protection of repeatedly carried out orthopterological re- the Czech Republic and partially by a project of the Ministry search at this locality between 1991-2002, and of Agriculture of the Czech Republic QH 71094. The authors S. pedo was not recorded there before its intro- would like to thank to Dr. Bruce Jaffee (USA) for linguistic and editorial improvements and many colleagues that help us duction in 2005. Like Pouzdřanská step-Kolby with field surveys (Jiří Dvořák, Tomáš Kavka, Martin NNR, NR Havranické vřesoviště is intensively Škorpík, Igor Malenovský, Petr Muckstein, Pavel Marhoul, visited by entomologists. The repeated detec- Jiří Matuška, David Hauck, Lenka Rožánková, Jolana tion of S. pedo at this locality indicates there are Szusková). still many habitats suitable for the occurrence of S. pedo in the eastern part of the Podyjí National Park. References The finding of a single S. pedo nymph along the road not far from the Bulhary village Anonymous (2005): Pavlovské vrchy a dolní Podyjí. Soubor in 2002 (see List of records above) is interest- turistických map 1:50 000. Trasa, spol. s.r.o., Praha, 1 map. [in Czech] ing. The finding is properly documented and Chládek , F. (1980): Poecilimon intermedius (Fieb.) neu für die localized, but its circumstances – sweeping Tschechoslowakei (Orthoptera, Ensifera). Articulata ditch along the road – do not indicate an au- 1(15): 152. tochtonous occurrence. In the area between Chládek, F. (1988): Rovnokřídlý hmyz (Orthoptera – Saltatoria), švábi (Blattoptera) a škvoři (Dermaptera) Bulhary village and Lednice town, all poten- státní přírodní rezervace "Pouzdřanská step" a jejího tially suitable localities were searched but nejbližšího okolí. Práce z oboru botaniky a zoologie, could not confirm the occurrence of S. pedo. Klub přírodovědecký v Brně 1988: 22-25. [in Czech] Chládek, F. (1995): Ensifera and Caelifera. pp. 121-128. In: The specimen was released near the monument Rozkošný, R., Vaňhara, J. (eds.), Terrestrial Invertebrates “Tři Grácie” in the area of Lednice Park. of the Pálava Biosphere Reserve of UNESCO I. Folia In summary, since 1900, in the Czech Re- Facultatis Scientiarum Naturalium Universitatis Masarykianae Brunensis, Biologia 92 (1-206 pp). public, S. pedo has been reported in five areas: Chytil, J. (2003): Návrat kobylky sága. Veronica 18(6): 25. [in Kamenný vrch hill at Kurdějov, Pouzdřanská Czech] step-Kolby NNR, Pavlovské vrchy hills, Dutrillaux, A.M., Lemonnier-Darcemont, M. Darcemont, C., Krpac, V., Fouchet, P., Dutrillaux, B. (2009): Origin of the Bulhary village, and the environs of Znojmo complex karyotype of the polyploid parthenogenetic (town). These localities represent the northern grasshopper Saga pedo (Orthoptera: Tettigoniidae). edge of the species range in central Europe European Journal of Entomology 106: 477-483. (Krištín, Kaňuch 2007). During the recent Dvořák, D., Roleček, J., Fajmon, K. (2008): A recent record of the predatory bush cricket Saga pedo Pallas, 1771, in the intensive research, which was carried out from Pouzdřanská step Steppe (southern Moravia). Acta 2006 to 2009, occurrences were confirmed only Musei Moraviae, Scientiae Biologicae 93: 7-9. in two areas: in the area of Pavlovské vrchy Ginter, O. (1924a): Saga serrata F. na Moravě. Sborník Přírodovědecké Společnosti v Moravské Ostravě 2(1922- hills and at Pouzdřanská step-Kolby NNR. The 1923): 1-4. [in Czech] most abundant recent population occurs in the Ginter , O. (1924b): Seznam moravských orthopter I. Časopis area of the Pavlovské vrchy hills, where during Československé Společnosti Entomologické 21: 7-11. [in Czech] the period of the research, 552 S. pedo indivi- Ginter, O. (1925): Acridiodea Burm. a Locustoidea Handl. na duals were recorded. Saga pedo occurs there at Moravě. Acridiodea Burm. et Locustoidea Handl. nine localities and is widespread over the Moraviae. Práce Moravské Přírodovědecké Společnosti 2: 21-84. [in Czech]

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Holuša, J., Kočárek, P., Drozd, P., Vlk, R. (2009): Analysis of k potravě kobylky ságy. In: Bryja, J., Zukal, J. (eds) population trend in Saga pedo (Orthoptera: Tettigoniidae) Zoologické dny Brno 2006. Sborník abstraktu on the edge of its range: more abundant or more z konference 9.-10. února 2006 [abstract of contribution intensively studied? Pp. 122-123. In: Metaleptea – was not printed]. [in Czech] special conference issue, 10th International Congress of Zamfirescu, S.R., Zamfirescu, O., Popescu, I.E., Ion, C. (2009): Orthopterology, 21-25 June 2009, Antalya, Turkey. Preliminary data on the population characteristics of Kempný, L. (1953): Nález Saga pedo (Pall.) (Sagidae, Vipera ursinii moldavica from “Dealul lui Dumnezeu” Orthoptera) na jižním Slovensku (Zobor u Nitry). (Iaşi County, Romania) with notes on conservation. Přírodovědný Sborník Ostravského Kraje 14: 529-534. North-Western Journal of Zoology 5: 85-96. [in Czech] Kenyeres, Z., Bauer, N., Rácz, I. (2002): Saga pedo Pallas dans le bassin des Carpates, synthese et nouvelles données (Orthoptera, Tettigoniidae). Bulletin de la Societe Submitted: 01 November 2009 Entomologique de France 107: 149-156. / Accepted: 20 August 2010 Kočárek, P., Holuša, J., Vidlička, Ľ. (2005): Blattaria, Mantodea, Orthoptera & Dermaptera of the Czech and Published Online: 23 August 2010 Slovak Republics. Blattaria, Mantodea, Orthoptera & Dermaptera České a Slovenské republiky. Kabourek, Zlín. [in Czech] Kočárek, P., Holuša, J. (2008): Kobylka sága a problematika její ochrany v České republice. Pp. 73. In: Tuf, I.H.,

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