6 Principles of Ecology
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UNIT IX: Plant Ecology Chapter 6 Principles of Ecology Learning Objectives • How soil, climate and other physical features aff ect the fl ora and fauna or vice versa? Th e learner will be able to Th ese questions can be better answered with Understand the interaction between the study of ecology. organisms and their Ecology is essentially a practical science environment. involving experiments, continuous Describe biotic and observations to predict how organisms react abiotic factors that to particular environmental circumstances infl uence the dynamics of and understanding the principles involved in populations. ecology. Describe how organisms adapt themselves to environmental changes. 6.1 Ecology Learn the structure of various fruits and The term “ecology” seeds related to their dispersal mechanism. (oekologie) is derived from two Greek words – oikos (meaning house or dwelling Chapter outline place and logos meaning study) It was first proposed by 6.1 Ecology R. Misra Reiter (1868). However, the 6.2 Ecological factors most widely accepted definition of ecology was 6.3 Ecological adaptations given by Ernest Haeckel (1869). 6.4 Dispersal of seeds and fruits Alexander von Humbolt - Father of Ecology Ecology is a division of biology which deals Eugene P. Odum - Father of morden Ecology with the study of environment in relation to R. Misra - Father of Indian Ecology organisms. It can be studied by considering individual organisms, population, community, biome or biosphere and their environment. 6.1.1 Definitions of ecology While observing our different environments, “The study of living organisms, both plants and one can ask questions like animals, in their natural habitats or homes.” - Reiter (1885) • Why do plants or animals vary with places? “Ecology is the study of the reciprocal • What are the causes for variation in relationship between living organisms and their biological diversity of diff erent places? environment.” - Earnest Haeckel (1889) 136 Principles of Ecology TN_GOVT_BOTANY_XII_PAGES_136 -161 CH 06.indd 136 01-03-2019 17:00:30 6.1.2 Ecological hierarchy habitat and niche of any organism is called Ecotope The interaction of Biosphere organisms with their The differences between habitat and niche are environment results in Biome as follows. the establishment of grouping of organisms Landscape Habitat Niche which is called ecological 1. A specific A functional space hierarchy or ecological Ecosystem physical space occupied by an levels of organization. occupied by organism in the same The basic unit of an organism eco-system Community ecological hierarchy is (species) an individual organism. 2. Same habitat A single niche is Population The different hierarchy may be shared by occupied by a single of ecological systems is many organisms species Individual organism illustrated below: (species) 3. Habitat Organisms may 6.1.3 Branches of Ecology: specificity is change their niche Ecology is mainly divided into two branches, exhibited by with time and season. they are autecology and synecology. organism. 1. Autecology is the ecology of an individual Table 6.1: Difference between habitat and niche species and is also called species ecology. 2. Synecology is the ecology of a population or Applied ecology or community with one or more species and also environmental technology : called as community ecology. Application of the Many advances and developments in the field Science of ecology is ecology resulted in various new dimensions otherwise called as Applied ecology or and branches. Some of the advanced fields are Environmental technology. It helps us to Molecular ecology, Eco technology, Statistical manage and conserve natural resources, ecology and Environmental toxicology. particularly ecosystems, forest and wild 6.1.4 Habitat and Niche life conservative and management. Environmental management involves Habitat Bio-diversity conservation, Ecosystem Habitat is a specific physical place or locality restoration, Habitat management, occupied by an organism or any species which Invasive species management, Protected has a particular combination of abiotic or areas management and also help us plan environmental factors. But the environment of landscapes and environmental impact any community is called Biotope. designing for the futuristic ecology. Niche An ecological niche refers to an organism’s place 6.1.5 Ecological equivalents in the biotic environment and its functional Taxonomically different species occupying role in an ecosystem. The term was coined by similar habitats (Niches) in different geographical the naturalist Roswell Hill Johnson but Grinell regions are called Ecological equivalents. (1917) was probably first to use this term. The Principles of Ecology 137 TN_GOVT_BOTANY_XII_PAGES_136 -161 CH 06.indd 137 01-03-2019 17:00:30 Examples: • Certain species of epiphytic orchids of Sun light Western Ghats of India differ from the Precipitation epiphytic orchids of South America. But they are epiphytes. Carbon • Species of the grass lands of Western Ghats dioxide and Water vapour Climatic factors of India differ from the grass species of Wind temperate grass lands of Steppe in North America. But they are all ecologically primary producers and fulfilling similar roles in their respective communities. Biotic factors Grazing animals Birds Rodents Insects Plant Pathogens 6.2 Ecological factors Man Epiphytes Many organisms, co-exist in an environment. Soil slope Edaphic factors The environment (surrounding) includes Soil water Minerals physical, chemical and biological components. Physical nature When a component surrounding an organism of the soil affects the life of an organism, it becomes a Soil air factor. All such factors together are called Figure 6.1: Environmental factors affecting a plant environmental factors or ecological factors. a. Light These factors can be classified into living Light is a well known factor needed for the (biotic) and non-living (abiotic) which make basic physiological processes of plants, such as the environment of an organism. However the photosynthesis, transpiration, seed germination ecological factors are meaningfully grouped and flowering. The portion of the sunlight which into four classes, which are as follows: can be resolved by the human eye is called i. Climatic factors visible light. The visible part of light is made- ii. Edaphic factors up of wavelength from about 400 nm (violet) to 700 nm (red). The rate of photosynthesis is iii. Topographic factors maximum at blue (400 – 500 nm) and red (600 – iv. Biotic factors 700 nm). The green (500 – 600 nm) wave length We will discuss the above factors in a concise of spectrum is less strongly absorbed by plants. manner. Effects of light on plants Flowers of poppy, chicory, dog rose and many other Light regulates Stem and plants, blossom before the Leaf formation Photosynthesisd break of dawn (4 – 5 am), a s ProductionRunner evening primrose open up with the onset Opening an Closing of stomat of dusk (5 – 6 pm) due to diurnal rhythm. Movements Tuber formation Tuber Germination of seedFlowering 6.2.1 Climatic Factors Climate is one of the important natural factors controlling the plant life. The climatic factors includes light, temperature, water, wind and Figure 6.2: Various effects of light upon a fire. green plant 138 Principles of Ecology TN_GOVT_BOTANY_XII_PAGES_136 -161 CH 06.indd 138 01-03-2019 17:00:32 Based on the tolerance to intensities of light, 1. Eurythermal: Organisms which can tolerate the plants are divided into two types. They are a wide range of temperature fluctuations. 1. Heliophytes - Light loving plants. Example: Zostera (A marine Angiosperm) and Example: Angiosperms. Artemisia tridentata. 2. Sciophytes - Shade loving plants. 2. Stenothermal: Organisms which can Example: Bryophytes and Pteridophytes. tolerate only small range of temperature variations. Example: Mango and Palm In deep sea (>500m), the environment is (Terrestrial Angiosperms). dark and its inhabitants are not aware of the Mango plant donot and cannot grow in existence of celestial source of energy called Sun. What, then is their source of energy? temperate countries like Canada and Germany. Thermal Stratification Palaeoclimatology–Helps to It is usually found in aquatic habitat. The reconstruct past climates of change in the temperature profile with our planet and flora, fauna increasing depth in a water body is called and ecosystem in which they thermal stratification. There are three kinds lived. Example: Air bubbles trapped in ice of thermal stratifications. for tens of thousands of years with fossilized pollen, coral, plant and animal debris. Epilimnion b. Temperature Metalimnion Temperature is one of the important factors which affect almost all the metabolic activities Hypolimnion of an organism. Every physiological process in an organism requires an optimum temperature at which it shows the maximum metabolic rate. Figure 6.3: Thermal stratification of pond Three limits of temperature can be recognized 1. Epilimniotn – The upper layer of warmer for any organism. They are water. 1. Minimum temperature - Physiological 2. Metalimnion – The middle layer with a activities are lowest. zone of gradual decrease in temperature. 2. Optimum temperature - Physiological 3. Hypolimnion - The bottom layer of colder activities are maximum. water. 3. Maximum temperature - Physiological Temperature based zonation activities will stop. Variations in latitude and altitude do affect the Based on the temperature prevailing in temperature and the vegetation on