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Report of Field Trip (Guilan Province from 14 to 17 October 2019)
Third Plenary Conference and Field Trip of INQUA IFG 1709F Ponto-Caspian Stratigraphy and Geochronology Report of field trip (Guilan Province from 14 to 17 October 2019) First Day, 14 October 2019 Stations: (Manjil, Totkabon (Rashi cave), Rasht) On Monday, at 7:30 AM the group traveled to the Gilan province. The first station was Manjil City for visiting Manjil Dam and geology Formations (Shemshak formation) in Mnajil. Dr. Nazari from Geological Survey of Iran described that how the formations were formed and some short descriptions about its tectonic plate. The second station had been located in the Totkabon (near the Ruodbar city). After having lunch near the Rashi village, the group led by Dr. Biglari from the Ministry of Cultural Heritage, Tourism and Handicrafts moved to the Rashi cave. At 8:00 PM, the team accommodated in their Hotel and hostel in Rasht. Second day, 15 October 2019: Stations: Fuman city, Masuleh village On Tuesday, 15 October 2019, field work started at the 8:30 AM. The team traveled to near Rasht City to visit old beach deposition and to the Fuman city for visiting Qale-Rudkhan Castle that is located 25 km southwest of Fuman city. And it is built on two tips of a mount, with an area of 2.6 hectare at the elevations of 715 and 670 meters. To get to the Qale - Rudkhan Castle, the group had to climbed approximately 1000 steps. On Tuesday afternoon, team went to Masuleh for visiting old village. (Also, at night a city tour was organized by the Guilan University). -
Spatial Epidemiology of Rabies in Iran
Aus dem Friedrich-Loeffler-Institut eingereicht über den Fachbereich Veterinärmedizin der Freien Universität Berlin Spatial Epidemiology of Rabies in Iran Inaugural-Dissertation zur Erlangung des Grades eines Doktors der Veterinärmedizin an der Freien Universität Berlin vorgelegt von Rouzbeh Bashar Tierarzt aus Teheran, Iran Berlin 2019 Journal-Nr.: 4015 'ĞĚƌƵĐŬƚŵŝƚ'ĞŶĞŚŵŝŐƵŶŐĚĞƐ&ĂĐŚďĞƌĞŝĐŚƐsĞƚĞƌŝŶćƌŵĞĚŝnjŝŶ ĚĞƌ&ƌĞŝĞŶhŶŝǀĞƌƐŝƚćƚĞƌůŝŶ ĞŬĂŶ͗ hŶŝǀ͘ͲWƌŽĨ͘ƌ͘:ƺƌŐĞŶĞŶƚĞŬ ƌƐƚĞƌ'ƵƚĂĐŚƚĞƌ͗ WƌŽĨ͘ƌ͘&ƌĂŶnj:͘ŽŶƌĂƚŚƐ ǁĞŝƚĞƌ'ƵƚĂĐŚƚĞƌ͗ hŶŝǀ͘ͲWƌŽĨ͘ƌ͘DĂƌĐƵƐŽŚĞƌƌ ƌŝƚƚĞƌ'ƵƚĂĐŚƚĞƌ͗ Wƌ͘<ĞƌƐƚŝŶŽƌĐŚĞƌƐ ĞƐŬƌŝƉƚŽƌĞŶ;ŶĂĐŚͲdŚĞƐĂƵƌƵƐͿ͗ ZĂďŝĞƐ͕DĂŶ͕ŶŝŵĂůƐ͕ŽŐƐ͕ƉŝĚĞŵŝŽůŽŐLJ͕ƌĂŝŶ͕/ŵŵƵŶŽĨůƵŽƌĞƐĐĞŶĐĞ͕/ƌĂŶ dĂŐĚĞƌWƌŽŵŽƚŝŽŶ͗Ϯϴ͘Ϭϯ͘ϮϬϭϵ ŝďůŝŽŐƌĂĨŝƐĐŚĞ/ŶĨŽƌŵĂƚŝŽŶĚĞƌĞƵƚƐĐŚĞŶEĂƚŝŽŶĂůďŝďůŝŽƚŚĞŬ ŝĞĞƵƚƐĐŚĞEĂƚŝŽŶĂůďŝďůŝŽƚŚĞŬǀĞƌnjĞŝĐŚŶĞƚĚŝĞƐĞWƵďůŝŬĂƚŝŽŶŝŶĚĞƌĞƵƚƐĐŚĞŶEĂƚŝŽŶĂůďŝͲ ďůŝŽŐƌĂĨŝĞ͖ ĚĞƚĂŝůůŝĞƌƚĞ ďŝďůŝŽŐƌĂĨŝƐĐŚĞ ĂƚĞŶ ƐŝŶĚ ŝŵ /ŶƚĞƌŶĞƚ ƺďĞƌ фŚƚƚƉƐ͗ͬͬĚŶď͘ĚĞх ĂďƌƵĨďĂƌ͘ /^E͗ϵϳϴͲϯͲϴϲϯϴϳͲϵϳϮͲϯ ƵŐů͗͘ĞƌůŝŶ͕&ƌĞŝĞhŶŝǀ͕͘ŝƐƐ͕͘ϮϬϭϵ ŝƐƐĞƌƚĂƚŝŽŶ͕&ƌĞŝĞhŶŝǀĞƌƐŝƚćƚĞƌůŝŶ ϭϴϴ ŝĞƐĞƐtĞƌŬŝƐƚƵƌŚĞďĞƌƌĞĐŚƚůŝĐŚŐĞƐĐŚƺƚnjƚ͘ ůůĞ ZĞĐŚƚĞ͕ ĂƵĐŚ ĚŝĞ ĚĞƌ mďĞƌƐĞƚnjƵŶŐ͕ ĚĞƐ EĂĐŚĚƌƵĐŬĞƐ ƵŶĚ ĚĞƌ sĞƌǀŝĞůĨćůƚŝŐƵŶŐ ĚĞƐ ƵĐŚĞƐ͕ ŽĚĞƌ dĞŝůĞŶ ĚĂƌĂƵƐ͕ǀŽƌďĞŚĂůƚĞŶ͘<ĞŝŶdĞŝůĚĞƐtĞƌŬĞƐĚĂƌĨŽŚŶĞƐĐŚƌŝĨƚůŝĐŚĞ'ĞŶĞŚŵŝŐƵŶŐĚĞƐsĞƌůĂŐĞƐŝŶŝƌŐĞŶĚĞŝŶĞƌ&Žƌŵ ƌĞƉƌŽĚƵnjŝĞƌƚŽĚĞƌƵŶƚĞƌsĞƌǁĞŶĚƵŶŐĞůĞŬƚƌŽŶŝƐĐŚĞƌ^LJƐƚĞŵĞǀĞƌĂƌďĞŝƚĞƚ͕ǀĞƌǀŝĞůĨćůƚŝŐƚŽĚĞƌǀĞƌďƌĞŝƚĞƚǁĞƌĚĞŶ͘ ŝĞ tŝĞĚĞƌŐĂďĞ ǀŽŶ 'ĞďƌĂƵĐŚƐŶĂŵĞŶ͕ tĂƌĞŶďĞnjĞŝĐŚŶƵŶŐĞŶ͕ ƵƐǁ͘ ŝŶ ĚŝĞƐĞŵ tĞƌŬ ďĞƌĞĐŚƚŝŐƚ ĂƵĐŚ ŽŚŶĞ ďĞƐŽŶĚĞƌĞ <ĞŶŶnjĞŝĐŚŶƵŶŐ ŶŝĐŚƚ njƵ ĚĞƌ ŶŶĂŚŵĞ͕ ĚĂƐƐ ƐŽůĐŚĞ EĂŵĞŶ ŝŵ ^ŝŶŶĞ ĚĞƌ tĂƌĞŶnjĞŝĐŚĞŶͲ -
Assessing Chlorophyll-A in the Southwestern Coastal Waters of the Caspian Sea
Journal of the Persian Gulf (Marine Science)/Vol. 5/No. 16/June 2014/14/1-14 Assessing Chlorophyll-a in the Southwestern Coastal Waters of the Caspian Sea Jamshidi, Siamak Iranian National Institute for Oceanography and Atmospheric Science, Tehran, IR Iran Received: October 2013 Accepted: April 2014 © 2014 Journal of the Persian Gulf. All rights reserved. Abstract Caspian Sea with an average depth of 27m is the largest enclosed water body in the world. Despite its enormity and valuable biotic and economic resources, investigations on the biota and seawater properties are mosaic at best. In previous studies, the monitoring of the chlorophyll-a concentrations in the Southern Caspian Sea was organized based on satellite data sets; however, vertical distribution of chlorophyll-a concentrations and its variations in deeper layers of the southern Caspian seawater are not well known. The aim of this research was to study the variations of seasonal distributions of chlorophyll-a near Anzali Port in northern Iran. Data collection was performed at 23 stations, 2 km apart, along four survey lines perpendicular to the coastline and two transects parallel to the coast every season using a research vessel. Field measurements in the study area were carried out. A portable CTD probe was applied for profiling from sea surface to bottom. Results of seasonal field measurements showed variations of the seawater properties (e.g. temperature, salinity, density and chlorophyll-a). Maximum depth of the profiling stations was more than 470 m and vertical structure of temperature in the southern Caspian Sea waters with a significant seasonal thermocline between 20-50 m depths in summer with a vertical difference of 16oC were characterized. -
High Prevalence of Hepatitis B Virus Infection in the Village of Esfandiar
ArchiveHepat Mon of. 2018 SID August; 18(8):e65473. doi: 10.5812/hepatmon.65473. Published online 2018 August 4. Research Article High Prevalence of Hepatitis B Virus Infection in the Village of Esfandiar in South Khorasan Province, Iran Davod Javanmard 1, 2, Seyed Moayed Alavian 3, Farshid Abedi 1, Mohammad Hasan Namaei 1, Arghavan Asghari 1 and Masood Ziaee 1, * 1Infectious Diseases Research Center, Birjand University of Medical Sciences, Birjand, Iran 2Department of Virology, School of Medicine, Iran University of Medical Sciences, Tehran, Iran 3Baqiyatallah Research Center for Gastroenterology and Liver Diseases, Baqiyatallah University of Medical Sciences, Tehran, Iran *Corresponding author: Infectious Diseases Research Center, Birjand University of Medical Sciences, Birjand, Iran. Tel: +98-9151613942, Fax: +98-5632433004, Email: [email protected] Received 2017 December 23; Revised 2018 July 15; Accepted 2018 July 24. Abstract Background: Infection with Hepatitis B virus (HBV) is still a leading cause of cirrhosis and hepatocellular carcinoma around the world. Objectives: Having recently received reports of high HBV prevalence in the South Khorasan Province in the Esfandiar village, we aimed to evaluate the prevalence of HBV in this region in comparison to neighboring villages, with associations to risk factors. Methods: This was a cross sectional study that included the Esfandiar village and neighboring villages, Marghoub, and Zenowghan. All of the Esfandiar and certain neighboring villages were summoned and blood sampling was performed along with the comple- tion of a questionnaire. Positivity for HBsAg, anti-HBc along with the seromarkers of HCV and HDV were assayed using ELISA method. Statistical analysis was performed using the SPSS software with chi-square tests, logistic regression, and risk-estimation analysis. -
Reproduction Characteristics of the Vimba Vimba Persa (Pallas, 1811), in Coastal Waters of the Caspian Sea
Reproduction characteristics of the Vimba vimba persa (Pallas, 1811), in coastal waters of the Caspian Sea Item Type article Authors Chaichi, A.R.; Vosoughi, G.H.; Kaymaram, F.; Jamili, S.; Fazli, H. Download date 28/09/2021 21:28:45 Link to Item http://hdl.handle.net/1834/37234 Iranian Journal of Fisheries Sciences 10(4) 585-595 2011 Reproduction characteristics of the Vimba vimba persa (Pallas, 1811), in coastal waters of the Caspian Sea Chaichi A. R. 1*; Vosoughi G. h.1; Kaymaram F.2; Jamili S.1,2; Fazli H.3 Received: October 2010 Accepted: March 2011 Abstract Some reproduction characteristics of the Caspian Vimba, Vimba vimba persa, were studied from Oct. 2008 to Sep. 2009 in coastal waters of the Caspian Sea (Mazandaran province). 994 specimens were monthly collected from 6 fish landing sites of Ramsar, Tonekabon, Chalus, Mahmood Abad, Sari and Behshahr. The fecundity was measured using 92 specimens. This species was found to have more abundance in spring (especially April and May). The samples composed of 397(42.6%) male, 537(57.4%) female; the sex ratio was M: F = 1:1.35. The advanced stages of maturity (4th & 5th) were found in April and May. The highest Gonadosomatic Index in females was in May and the lowest one was in July. The average of absolute and relative fecundities was 17198±7710 and 171.85±48.8, respectively. Keywords: Caspian Vimba, Vimba vimba persa, Fecundity, GSI, Sex ratio, Caspian Sea Downloaded from jifro.ir at 16:34 +0330 on Monday February 12th 2018 ____________________ 1- Graduate School of Marine Science & Technology, Department of Marine Biology, Science and Research, Islamic Azad University, Tehran- Iran. -
We Intend to Set a New RECORD IMIDRO Exclusive
Rls. 300,000 / €30 June 2015 / IMIDRO Exclusive www.iraninternationalmagazine.com www.iraninternational.ir Iran International, the Forum for Partners in Iran’s Marketplace Double Issue We Intend to Set a New RECORD IMIDRO Exclusive Iran’s Mining Projects Open to Foreign Investors 2 IRAN INTERNATIONAL, June 2015 IRAN INTERNATIONAL, June 2015 3 Competition between Mine and Oil IMIDRO Exclusive ith regard to the started the above-mentioned tendency of the studies so that after identifi- international com- cation and discovery of de- munityW towards advanced posits and their processing Important industries (Hi Tec) in such and production we would fields as electronic and tele- be able to meet domestic communication industries, demands with regard to the automotive, catalysts, perma- development of modern in- Development nent magnets, metallurgical dustries. In this regard, the alloys, phosphorescence and plan for “processing and fea- pigments, glass and ceram- sibility study of rare earth el- ics, aerospace, etc., produc- ements” has been defined and Factor tion and consumption of rare started implementation. earth elements and strategic Iron and phosphate ore metals such as lanthanide, and tailings are among re- scandium and yttrium, as the sources possessing rare earth raw materials for the manu- elements and with their si- facture of high technology multaneous extraction more products have taken a mo- value added can be created. mentum. In all iron ore mining com- IMIDRO in an effort panies affiliated to IMIDRO not to lag behind the trend of sampling of rare elements international developments was conducted on tailings has formulated a plan for and studies on the amount exploration of rare earth ele- of deposits of the elements ments (REE) and has started are being conducted. -
Award Rationale Esfahk Historic Village in Iran
Award Rationale Esfahk Historic Village in Iran Klaus Betz On behalf of Studienkreis für Tourismus und Entwicklung e.V. November 2019 Contents Maps [1] 1. Introduction [2] 2. Backround [2] 3. The Project Esfahk Historic Village [3] 4. Evaluation Against the TO DO Award Criteria [5] 5. Conclusion and Comments [9] Project Location of Esfahk Historic Village Source: Google Maps Source: Google Maps 1 “Then let us turn what was bad into something good!“ 1 INTRODUCTION The application for the TO DO Award 2020 was submitted by the village council of Esfahk and refers to the original Esfahk Historic Village. On the basis of the documents submitted, the application was nominated by the TO DO Award jury. From 6th to 15th November 2019, expert Klaus Betz travelled in Iran on behalf of the Institute for Tourism and Development (Studienkreis für Tourismus und Entwicklung e. V.) in order to evaluate Esfahk Historic Village on location. The village of Esfahk (also spelt Esfahak) with a population of 800 inhabitants is situated at an altitude of 1,100 m, along the national road 68, not far from the district capital of Tabas City in South Khorasan Province in the East of Iran. The small oasis village with its date palm grove and irrigated gardens and fields all around is situated in or rather on the edge of the Parvedeh salt desert/salt steppe, with the rocky “camel mountains“ at its back. Their highest elevation is the Jamal peak (”the beautiful one”, “the handsome one“), rising to an altitude of about 3,000 metres. -
Mid-Term Plan for Conservation of the Anzali Wetland for 2020 – 2030
Japan International Department of Environment Cooperation Agency Gilan Provincial Government Islamic Republic of Iran Mid‐term Plan for Conservation of the Anzali Wetland for 2020 ‐ 2030 May 2019 Anzali Wetland Ecological Management Project ‐ Phase II Department of Environment Japan International Gilan Provincial Government Cooperation Agency Islamic Republic of Iran MID-TERM PLAN FOR CONSERVATION OF THE ANZALI WETLAND FOR 2020 – 2030 (Prepared under The Anzali Wetland Ecological Management Project - Phase II) May 2019 NIPPON KOEI CO., LTD. Exchange Rate JPY 100 = IRR 38,068 USD 1 = IRR 42,000 (as of 23 May, 2019) Source: Central Bank of the Islamic Republic of Iran Preface The Mid-term Plan for Conservation of the Anzali Wetland for 2020 – 2030 (Mid-term Plan) was prepared as a final product of the Anzali Wetland Ecological Management Project - Phase II (Phase II Project). The Phase II Project was a 5-year technical cooperation project of the Japan International Cooperation Agency (JICA) between May 2014 and May 2019. JICA has supported Iranian government on conservation of the Anzali Wetland since 2003 through “The Study on Integrated Management for Ecosystem Conservation of the Anzali Wetland (2003-2005)” (Master Plan Study) and “Anzali Wetland Ecological Management Project (2007-2008, 2011-2012)” (Phase I Project). The Mid-term Plan will succeed the previous Master Plan for 2005 - 2019, which was prepared under the Master Plan Study. In the 1st year of the Phase II Project, actual implementation status of the Master Plan was reviewed and an Action Plan for 5 years, which is the last 5-year of the Master Plan and period of the Phase II Project, was prepared to facilitate the conservation activity of the Anzali Wetland. -
Ecological Study of Phytoplankton of the Anzali Lagoon (N Iran) and Its Outflow Into the Caspian Sea
Czech Phycology, Olomouc, 4: 145-154, 2004 145 Ecological study of phytoplankton of the Anzali lagoon (N Iran) and its outflow into the Caspian Sea Ekologická studie fytoplanktonu laguny Anzali (severní Irán) a jejího vtoku do Kaspického moře Zohreh R a m e z a n p o o r International Sturgeon Research Institute. P. O. Box: 41635 – 3464 Rasht – Iran, e-mail: [email protected] Abstract A study of phytoplankton communities of western regions of the Anzali lagoon, rivers flowing from the lagoon, and Anzali Harbor (navigation channel), located in the southern coast of the Caspian Sea was carried out. Samples were collected monthly at nine sites from May 1992 to April 1993 from the surface, 1 m, 5 m and from the final depths of each sampling site using Ruttner sampler with 1 lit. volume. Water and air temperature, - 3- - 2- turbidity, pH, Cl , PO4 , NO3 , SO4 and salinity were measured. Phytoplankton samples were fixed in 4% formaldehyde solution and then identified and counted using inverted microscope. A total of 97 taxa of phytoplankton were identified during the studied period. Phytoplankton species and environmental variables confirmed that the western part of the Anzali Lagoon has fresh water and eutrophic character. However, alkalinity in this region was high and high amount of sulfate at this region indicates the influence of the Caspian Sea in the area. As the Harbor is under combined influence of the Caspian Sea and freshwater rivers, it showed mostly estuarine phytoplankton communities with some species of marine origin. Few sources of pollution were identified that reduced the species diversity and alternatively some other species became dominant. -
Astacus Leptodactylus) Based on Microsatellite DNA Loci
Archive of SID Iranian Journal of Animal Biosystematics (IJAB) Vol.14, No.1, 15-27, 2018 ISSN: 1735-434X (print); 2423-4222 (online) DOI: 10.22067/ijab.v14i1.54723 Population molecular analysis among the narrow-clawed crayfish (Astacus leptodactylus) based on microsatellite DNA loci Khoshkholgh, M.1* and Nazari, S. 2 1Department of Fisheries, School of Natural Resources, Guilan university, P.O. Box 1144 Sowmehsara, Iran 2Shahid Motahary Cold-water Fishes Genetic and Breeding Research Center, Iranian Fisheries Sciences Research Institute, Agricultural Research, Education and Extension Organization (AREEO), Yasuj, Iran (Received: 15 October 2017; Accepted: 10 February 2018) The genetic variation of narrow-clawed crayfish (Astacus leptodactylus, Eschscholtz, 1823) from Caspian Sea including two sites (Astara and Kiashahr), Aras region, Anzali lagoon and three rivers (Chafrood, Masuleh, and Siah Darvishan) was evaluated and their genetic variation on the basis of DNA microsatellite loci was estimated. DNA from pleopods of 194 specimens extracted and was examined with six microsatellite markers. Genetic differences between the populations were discerned by pairwise comparison based on allelic distribution. The average numbers of alleles per locus ranged from 2 to 10, while the average observed heterozygosity (Ho) at various loci varied between 0.222 to 0.732, implying that a midway level of genetic variation. Among seven populations Siah Darvishan population displayed the highest level of variability in terms of heterozygosity. Tests of Hardy-Weinberg showed that the microsatellite loci deviated significantly in most populations indicating deficit of heterozygote. The results indicate that some of the above populations are significantly differentiated from one another based on pairwise FST estimates. -
Highlights: Trip Overview
Highlights: Trip Overview: St. Thaddeus Monastery Iran is truly one of the worlds beautiful countries, not just in nature but in its culture and variety of ethnic groups inhabiting Masuleh Village it. Compared to trekking in other countries in the region, Iran Alamout Castel offers better serviced trekking routes making trekking easy and Persepolis (UNESCO World Heritage Sites) comfortable. Trekking in Iran has captured the imaginations of Services: mountaineers and explorers for more than 1000 years. The B / B, H / B, F / B (Based on Your Choice) lifestyle and habits of the inhabitants of the mountain regions Hotels: has not changed for generations, the village people offer a 3 Stars, 4 Stars, 5 Stars or Camp huge sense of hospitability and welcome trekkers and (Based on The Program) mountaineers alike. 2nd floor, NO 40, Shahid Beheshti Ave, Tehran, Iran Tel: +98 21 88 46 07 55 / +98 21 88 46 09 78 Fax: +98 21 88 46 10 32 Email: [email protected] / [email protected] www.pitotour.net Day 1: Pre reserve symbol of Iran high up in the the Arasbaran forest near Kaleybar City. It was also one of the last regional Day 2 Tabriz: Morning arrival Tabriz, meet the Guide and strongholds to fall to Arab invaders in the 9th Century. transfer to the Hotel. After that drive to Jolfa Border, to O/N in Kaleybar visit two of the best churches in iran. St Stepanos Monastery and St. Thaddeus Monastery. The Saint Day 5 Kaleybar - Sareyn: Drive to Sareyn through Ahar. Thaddeus Monastery is an ancient Armenian monastery Sareyn, is a city and the capital of Sareyn County, in located in the mountainous area of Iran's West Ardabil Province, Iran. -
Heavy Metals Phytoremediation by Aquatic Plants (Hyrocotyle Ranocloides, Ceratophyllum Demersum) of Anzali Lagoon
Int. J.Mar.Sci.Eng., 2(4), 249-254, Autumn 2012 ISSN 2251-6743 © IAU Heavy metals phytoremediation by aquatic plants (Hyrocotyle ranocloides, Ceratophyllum demersum) of Anzali lagoon 1*L. VahdatiRaad; 2H. Khara 1 Department of Marine Biology, Graduate School of Marine Science and Technology, Science and Research Branch, Islamic Azad University, Tehran, Iran 2 Department of Fisheries, Faculty of Natural Resources, Islamic Azad University of Lahijan Branch, P.O. Box 1616, Lahijan, Iran Received 10 August 2012; Revised 29 August 2012; Accepted 28 September 2012 ABSTRACT: Wetlands are often considered sinks for contaminants, and Aquatic plants have been shown to play important roles in wetland biogeochemistry through their active and passive circulation of elements. In this research, in order to evaluate the concentration level of heavy metals released by the urban, industrial and agricultural activities at the margin of the rivers flowing into Anzali lagoon, as well as finding lead and cadmium accumulated in two plant species quoted below, a sampling has been made on two kinds of aquatic plants in the lagoon at three randomized stations based on the density of the plant coverage. In this project two regional aquatic plant species asHyrocotyleranocloides, Ceratophyllumdemersumhave been investigated for absorption of lead and cadmium. Results show an average concentration of lead in Ceratophyllum being 53.11 ppm, for Hydrocotyle 77.8 ppm, and of cadmium in Ceratophyllum being 4.46 ppm and for Hydrocotyle 6.28 ppm. According to results, lead has been the most abundant between these two metals inside plant organs regardless the specie. The statistic test gives the results under the confidence level 95% proving the amount of the absorption being significant in the stem of Hydrocotyle plant rather than in the root or the leaf compared with other species.