The Evolutionary Theory of Sex

Total Page:16

File Type:pdf, Size:1020Kb

The Evolutionary Theory of Sex TWO SEXES. Why? ♂ ♀ ♀ ? ♀ The Evolutionary Theory of Sex \ Two Sexes – Why? The Evolutionary Theory of Sex Wilmington 2011 Two Sexes – Why? The Evolutionary Theory of Sex — Wilmington, 2011. What do we need two sexes for? What is the evolutionary significance of this phenomenon? What is the nature of the differences between the sexes? What is sexual dimorphism and what does it mean? How is it connected to other life phenomena? The new evolutionary theory of sex differentiation developed by Dr. Biol. Sci. Vigen Geodakian has the answers. It provides a scientific justification for the existence of men and women, and the evolutionary roles that they play in family and society. Pages: х + 230. Ill. 49, tabl. 33, bibliography 637. ISBN xxxxx (PDF) Compiled and edited by S. V. Geodakyan. The materials presented in the book, are submitted selectively and if they contain any omissions, the responsibility for them bears the originator of this work. In case of any questions and uncertainties readers should refer to the original papers listed in the bibliography. Copyright © 2011 by Sergey Geodakyan. All rights reserved. Printed in the United States of America. No part of this publication may be reproduced or transmittedin any form or by any means without the written permission of the publisher. E-mail: [email protected] PREFACE The first publication of Vigen Geodakian devoted to the problem of sex differentiation, appeared in 1965 in cybernetic magazine. Then came an article in popular science magazine “Science and Life”. Due to multimillion distribution this article received much attention. Many scientists still remember just this one small aspect of the theory—how to influence the sex of offspring. Hovewer, over 45 years of existence, the theory was greatly expanded. Since then, more than 150 articles has been published devoted to longevity, differentiation of a brain and hands, sex chromosomes, mechanisms of sex ratio regulation at plants and animals, congenital heart anomalies, and even culture. Newspapers and magazines repeatedly wrote about the theory. V. Geodakian made presentations at many Russian and international congresses, conferences and symposia, lectured hundreds of students. Two conferences have been devoted exclusively to the theory (Saint Petersburg, Russia, 1990, 1992). The theory has already entered the textbooks (V. Vasiltchenko, 1977, 1986, 2005; Tkatshenko et al., 2001; Nartova-Bashaver, 2003; Ilin, 2003; Zhukov, 2007) and was also included in study programs of some Russian (Physical-Technical College, Moscow College of Physical Engineers, Russian Humanitarian State University) and foreign (Tel-Aviv University) colleges and universities. The breadth of the theory, it’s explanatory and predictive capacity allow it to substitute many limited and imperfect theories of gender and related fields, in particular—C. Darwin’s theory of sexual selection. A look at dioecy as an effective way of evolution, and at males as an evolutionary “vanguard” of a population, allows to explain from the unified point of view many mysteries related to sex. The basic idea of asynchronous evolution led to the creation of two new theories—brain asymmetry and laterality (in 1993), and the evolutionary role of sex chromosomes (in 1996), as well as original interpretation of hormonal sex, homosexuality, cancer, and culture. Despite the large number of publications, the most complete presentation of the theory is still the V. Geodakian’s doctorate thesis (Institute of Developmental Biology, 1987). This book attempts to describe the current state of the theory including new developments and accumulated experimental data. General composition of the book is similar to that of the dissertation. First four chapters cover current state of the problem of sex, unclear phenomena and riddles, existing theories and their criticism. Chapters 5 and 6 are devoted to the classification and analysis of reproduction types, chapter 7—involvement of the sexes in the transmission of information to the progeny, chapter 8—sex ratio. Sexual dimorphism, mechanisms of it’s occurence, forms and rules are described in chapters 9–12. Sexual dimorphism in pathology is examined in chapter 13. Chapter 14 covers possible mechanisms of discovered relationships, chapter 15—predictions. Summary has all statements of the new theory and the general scheme of the trait’s evolution in ontogeny and phylogeny. The main goal was to and at the same time as much as possible to preserve the style of author’s publications. The book does not attempt to make a full review of the area of sex research, which can be found in several monographs. The areas where the thery was already applied were covered the most. The book is intended to scientists, teachers, students, and other professionals working in the fields of biology, medicine, psychology, sociology, anthropology, and many other related branches of science. Since sex-related questions are of great interest to almost everybody, more simplle description of the theory can be found on official theory’s web site (www.geodakian.com). Table of Contents INTRODUCTION ...................................................................................................................................... IX CHAPTER 1. ANALYSIS OF THE THREE MAIN WAYS OF REPRODUCTION ....................... 1 COMPARISON OF SEXUAL WAYS OF REPRODUCTION WITH ASEXUAL ................................................... 1 Hypotheses and Theories ................................................................................................................... 5 "Variation and Selection" hypotheses ........................................................................................... 5 "Immediate Benefit" hypotheses ................................................................................................... 6 COMPARISON OF DIOECIOUS WAYS WITH HERMAPHRODITE ................................................................. 6 CHAPTER 2 MYSTERIES OF DIOECY: SEX RATIO ................................................................... 11 HIGH SECONDARY AND TERTIARY SEX RATIO ..................................................................................... 11 INFLUENCE OF HEREDITY ON PRIMARY AND SECONDARY SEX RATIO ................................................ 12 INFLUENCE OF ENVIRONMENT ON PRIMARY AND SECONDARY SEX RATIO ......................................... 13 “Phenomenon of war years” ........................................................................................................... 14 “Phenomenon of war years”—theories ....................................................................................... 14 MYSTERIES OF LONGEVITY ................................................................................................................... 16 Decreasing of Tertiary Sex Ratio in Ontogeny ................................................................................ 19 Men are “Champions” of Longevity ................................................................................................ 19 Early termination of fertility in women ............................................................................................ 19 Theories of High Male Mortality ..................................................................................................... 19 EVOLUTIONARY MECHANISMS OF SEX RATIO REGULATION ............................................................... 21 CHAPTER 3. MYSTERIES OF DIOECY: SEXUAL DIMORPHISM ........................................... 25 SEX DIFFERENCES IN HUMANS ............................................................................................................. 27 Anatomy ........................................................................................................................................... 27 Physiology ........................................................................................................................................ 28 Sense of Touch ............................................................................................................................ 28 Olfaction ..................................................................................................................................... 29 Vision .......................................................................................................................................... 29 Hearing and Voice ...................................................................................................................... 30 PSYCHOLOGICAL SEXUAL DIMORPHISM ............................................................................................... 31 ROLES IN SOCIETY ................................................................................................................................ 32 Work ................................................................................................................................................. 32 Science and Inventions ..................................................................................................................... 33 Culture ............................................................................................................................................. 34 Sport ................................................................................................................................................. 34 Religion ...........................................................................................................................................
Recommended publications
  • Stehlik 2002
    American Journal of Botany 89(12): 2007±2016. 2002. GLACIAL HISTORY OF THE ALPINE HERB RUMEX NIVALIS (POLYGONACEAE): A COMPARISON OF COMMON PHYLOGEOGRAPHIC METHODS WITH NESTED CLADE ANALYSIS1 IVANA STEHLIK2 Institute of Systematic Botany, University of Zurich, Zollikerstrasse 107, CH-8008 Zurich, Switzerland The glacial history of the alpine herb Rumex nivalis was investigated using ampli®ed fragment length polymorphisms (AFLPs) and restriction fragment length polymorphisms with polymerase chain reaction (PCR-RFLPs) of cpDNA. Both traditional statistical methods widely applied in phylogeographic research and nested clade analysis were used. The AFLPs indicated little geographic structure probably due to the wind-pollinated reproductive system of the dioecious R. nivalis. Because cpDNA haplotypes exhibited distinct distributional patterns, correlation between AFLPs and PCR-RFLPs was low. The results of common statistical methods and of nested clade analysis were largely congruent. Both supported in situ survival of one group of common haplotypes in the Central Alps. For another group of common haplotypes, classical phylogeographic analyses gave strong evidence for survival in peripheral refugia at the northern alpine border, whereas this conclusion was not as clearly supported in the nested clade analysis. Nested clade analysis provided several detailed insights on past and ongoing populational demographic processes. Thus, it is a valuable tool in the phylo- geographical analysis of haplotype data, but it should preferably be combined with other statistical analyses. In situations with low genetic variation in cpDNA, classical phylogeographic analytical tools on nuclear DNA will still be the methods of choice. Key words: AFLP; glacial survival; methodological comparison; migration; nested clade analysis; nunataks; PCR-RFLPs of cpDNA; Pleistocene glaciation.
    [Show full text]
  • Města Brna – Historie a Současnost
    2020 Motýli (Lepidoptera) města Brna – historie a současnost Zdeněk Laštůvka, Aleš Laštůvka af.mendelu.cz 2020 Zdeněk Laštůvka, Aleš Laštůvka Motýli (Lepidoptera) města Brna – historie a současnost Vědečtí recenzenti: Ing. Jan Liška, Výzkumný ústav lesního hospodářství a myslivosti, v.v.i., Jíloviště-Strnady Ing. Jan Šumpich, Národní muzeum, Praha Možná citace: Laštůvka Z. & Laštůvka A., 2020: Motýli (Lepidoptera) města Brna – historie a současnost. Mendelova univerzita v Brně, Brno, 120 s. © Zdeněk Laštůvka & Aleš Laštůvka, 2020 © Mendelova Univerzita v Brně, Zemědělská 1, 613 00 Brno ISBN 978-80-7509-750-7 (Print) ISBN 978-80-7509-769-9 (On-line) Motýli (lepidoptera) Města Brna – historie a současnost OBSAH Abstrakt . 4 Abstract ...................................................................4 1 Úvod ....................................................................5 2 Historie studia motýlů na Brněnsku ........................................6 2.1 Nejstarší badatelé . 6 2.2 První polovina 20. století ................................................... 6 2.3 Poválečné období.......................................................... 7 2.4 Současnost ................................................................ 7 3 Přírodní poměry, proměny města a biotopy .................................8 4 Metodika ...............................................................13 5 Celkové výsledky . 17 6 Přehled zjištěných druhů.................................................20 7 Změny v průběhu času ...................................................53
    [Show full text]
  • A Guide to Frequent and Typical Plant Communities of the European Alps
    - Alpine Ecology and Environments A guide to frequent and typical plant communities of the European Alps Guide to the virtual excursion in lesson B1 (Alpine plant biodiversity) Peter M. Kammer and Adrian Möhl (illustrations) – Alpine Ecology and Environments B1 – Alpine plant biodiversity Preface This guide provides an overview over the most frequent, widely distributed, and characteristic plant communities of the European Alps; each of them occurring under different growth conditions. It serves as the basic document for the virtual excursion offered in lesson B1 (Alpine plant biodiversity) of the ALPECOLe course. Naturally, the guide can also be helpful for a real excursion in the field! By following the road map, that begins on page 3, you can determine the plant community you are looking at. Communities you have to know for the final test are indicated with bold frames in the road maps. On the portrait sheets you will find a short description of each plant community. Here, the names of communities you should know are underlined. The portrait sheets are structured as follows: • After the English name of the community the corresponding phytosociological units are in- dicated, i.e. the association (Ass.) and/or the alliance (All.). The names of the units follow El- lenberg (1996) and Grabherr & Mucina (1993). • The paragraph “site characteristics” provides information on the altitudinal occurrence of the community, its topographical situation, the types of substrata, specific climate conditions, the duration of snow-cover, as well as on the nature of the soil. Where appropriate, specifications on the agricultural management form are given. • In the section “stand characteristics” the horizontal and vertical structure of the community is described.
    [Show full text]
  • Sooti Erinev Muna Suurus Liblikatel
    TARTU ÜLIKOOL ÖKOLOOGIA JA MAATEADUSTE INSTITUUT ZOOLOOGIA OSAKOND ENTOMOLOOGIA ÕPPETOOL Merili Martverk SOOTI ERINEV MUNA SUURUS LIBLIKATEL Bakalaureusetöö Juhendajad: prof. Toomas Tammaru PhD Tambet Tõnissoo MSc Riho Meier TARTU 2016 Sooti erinev muna suurus liblikatel Sooline dimorfism on bioloogide huviorbiidis olnud juba pikka aega, kuid suhteliselt vähe on olnud uurimistöid, mis käsitlevad dimorfismi kujunemist ontogeneesis. Viimasel ajal on dimorfismi ontogeneetilist kujunemist rohkem uuritud, kuid selliste uurimuste keskmes on olnud peamiselt kasvukiirus ja arengukestus. Ometi on dimorfismi kujunemisel ka võimalus, et erinevus on olemas juba algusest peale ehk siis sünnil või koorumisel. Käesolev töö keskendub dimorfismi uurimisel just nimelt sellele võimalusele. Töös antakse ülevaade munaraku arengust, soo määramise mehhanismidest ja empiirilistest uurimustest muna suuruse soolise dimorfismi kohta putukatel. Refereerides liblika munaraku arengu kohta teada olevat ei leitud ühtegi asjaolu, mis välistaks dimorfismi kujunemist juba munastaadiumis. Samas leiti, et sellist nähtust on dokumenteeritud putukate seas vaid haplodiploidse soo määramise mehhanismiga kiletiivalistel. Märksõnad: Sooline dimorfism, Lepidoptera, oogenees, soo määramine, muna suurus Sexual size dimorphism and egg size in Lepidoptera Even if studies on sexual size dimorphism (SSD) have a long history, ontogenetic mechanisms leading to unequal sizes of males and females have received little attention. Since recently, interest in ontogenetic patterns behind SSD has been growing but these studies have mostly been focusing on growth rate and development time. However, there is one more way to achieve SSD: initial size difference among new-born individuals or eggs. The present study focuses on that possibility. This study presents data on the empirical studies on egg size dimorphism in insects. It also gives an overview of oocyte development and sex determination mechanisms.
    [Show full text]
  • Identification of Micro-Moth Families
    Identification of micro-moth families Introduction This guide is intended for use by Garden Moth Scheme (GMS) recorders and other British and Irish moth enthusiasts. It is aimed at those who already have some understanding of moth taxonomy and morphology but struggle to easily home-in on which moth family a micro- moth belongs to. For further information about how to start out moth- trapping see www.gardenmoths.org.uk. There are currently over 1600 British and Irish micro-moth species, compared to around 800 species of macro-moth and 60 species of butterfly, so species identification is particularly challenging. Narrowing down identification to species requires additional information from training courses, websites and field guides. This guide shows photographs of 45 micro-moth families at rest, together with identification tips, similar species, and the number of species in that family. Taxonomists disagree on the order and arrangement of families, but this guide follows the numbering system of Bradley and Fletcher (B&F) wherever possible. The guide is intended to be used with the excellent Field Guide to the Micromoths of Great Britain and Ireland by Sterling, Parsons and Lewington, British Wildlife Publishing (2012, SPL), which covers 1000+ micro-moth species. However, for the uninitiated, facing a light trap full of micro-moths starting from scratch with SPL can be both daunting and time-consuming. Family names are different and are shown in a different order in SPL and B&F, so in order to reduce confusion, after the B&F numbers are quoted for each family the page numbers for the plates in SPL are also given.
    [Show full text]
  • Survival in the Alpine Landscape Genetic, Demographic And
    Survival in the alpine landscape Genetic, demographic and reproductive strategies of the rare monocarpic perennial Campanula thyrsoides in the Swiss Alps INAUGURALDISSERTATION zur Erlangung der Würde eines Doktors der Philosophie vorgelegt der PHILOSOPHISCH-NATURWISSENSCHAFTLICHEN FAKULTÄT der Universität Basel von PATRICK KUSS aus Ludwigshafen am Rhein, Deutschland Basel, 2006 Genehmigt von der Philosphisch-Naturwissenschaftlichen Fakultät auf Antrag von Herrn Prof. Dr. Christian Körner Herrn PD Dr. Jürg Stöcklin Herrn Prof. Dr. Markus Fischer Basel, den 3. Juli 2006 Prof. Dr. Hans-Jakob Wirz Dekan Table of Contents Chapter 1 General Introduction 3 Chapter 2 Spatial differentiation and genetic differentiation in naturally 11 fragmented alpine plant populations P. Kuss, A.R. Pluess, H.H. Ægisdóttir, J. Stöcklin Chapter 3 Evolutionary demography of the long-lived monocarpic 37 perennial Campanula thyrsoides in the Swiss Alps P. Kuss, M. Rees, H.H. Ægisdóttir, J. Stöcklin Chapter 4 No inbreeding depression in an outcrossing Alpine species: 65 the breeding system of Campanula thyrsoides H.H. Ægisdóttir, D. Jespersen, P. Kuss, J. Stöcklin Chapter 5 The biological flora of Central Europe: Campanula thyrsoides L. 81 P. Kuss, H.H. Ægisdóttir, J. Stöcklin Chapter 6 General Discussion and Outlook 107 Acknowledgements 115 Curriculum Vitae 117 1 2 General Introduction Chapter 1 General Introduction 3 Chapter 1 The aim of this thesis This thesis comprehensively studies the genetic, demographic and reproductive strategies of the long-lived monocarpic perennial Campanula thyrsoides to assure survival in the alpine environment. The work is part of the larger project “How patchy habitats and isolation affect plant life: genetic diversity, gene flow and mating systems” which forms the framework for the PhD investigations of Hafdís Hanna Ægisdóttir and the author under the supervision of PD Dr.
    [Show full text]
  • Lepidoptera: Psychidae): Integrating Ecological and Molecular Analyses
    Eur. J. Entomol. 108: 635–650, 2011 http://www.eje.cz/scripts/viewabstract.php?abstract=1663 ISSN 1210-5759 (print), 1802-8829 (online) The parasitoid species complex associated with sexual and parthenogenetic Naryciinae (Lepidoptera: Psychidae): Integrating ecological and molecular analyses JELMER A. ELZINGA1, KEES ZWAKHALS2, JOHANNA MAPPES1 and ALESSANDRO GRAPPUTO3 1Centre of Excellence in Evolutionary Research, Department of Biological and Environmental Science, P.O. Box 35, 40014 University of Jyväskylä, Finland; e-mails: [email protected], [email protected] 2 Dr. Dreeslaan 204, 4241 CM Arkel, The Netherlands; e-mail: [email protected] 3Department of Biology, University of Padua, Via Ugo Bassi 58/B, I-35121 Padua, Italy; e-mail: [email protected] Key words. Barcoding, larval remains, parasitism, parthenogenesis, sex ratio, COI, COII, Wingless, Hymenoptera, Braconidae, Ichneumonidae, Lepidoptera, Psychidae, Naryciinae Abstract. This study describes the parasitoid species complex associated with seven closely related species of sexual (Siederia rupi- collella, S. listerella, Dahlica lazuri, D. charlottae and D. lichenella) and parthenogenetic (Dahlica fennicella and D. triquetrella) Naryciinae (Lepidoptera: Psychidae) in Central Finland. A thorough ecological analysis of all the species of parasitoids recorded was combined with analyses of molecular data. Mitochondrial and nuclear DNA data were obtained from all the species in order to (1) detect cryptic species associated with host specialization, (2) assign undescribed males to females, and (3) verify the morphological identification of closely related species. A DNA barcoding technique was employed to identify host species from parasitized larval remains. By sampling more than 10,000 host larvae, of which 25.7% were parasitized, nine parasitoid species were identified mor- phologically, including both koinobionts (Ichneumonidae: Diadegma incompletum, Macrus parvulus, Trachyarus borealis, T.
    [Show full text]
  • Proceedings and Transactions of the British Entomological and Natural History Society
    ^ D.C C2n.dc; :!z c/J — c/> iiiSNi NviNOSHiii^s S3iyvaan libraries Smithsonian inj z ^ 2: " _ ^W^:^^ r- \i^A liars:'/ -^ "^M^^///'y^rj^j'' t*— \RIES*^SMITHSONiAN INSTITUTION NOIinillSNI NVIN0SHillMs'^S3 to 5 to — C/5 a: DO \^ iIiSNI~NV!NOSHimS S3IMVHan LIBRARIES SMITHS0NIAN"'|N! 03 73 ^ C/^ ± C/5 \RIES SMITHSONIAN INSTITUTION NOIiniliSNI NVINOSHlllNS S3 to (J) "Z. t t^ .*r^-. < Wp/^^ iiiSNi_NViN0SHims S3iavyan libraries Smithsonian in: X-'i\ _i ^RIES^SMITHSONIAN INSTITUTION NOIiniUSNI NVINOSHill^S S* z r- 2: r- z: to _ to uiiSNi NViNOSHims S3iyvdan libraries Smithsonian in z: CO z >•• to 2 X-H COo Z > '-i^ :s: *\. > _ * c/5 Z c/7 Z c/5 ARIES SMITHSONIAN INSTITUTION NOIinillSNI NVINOSHlllNS S c^ CO 5: ^ £/i ^ ^ .- < m . '^ m m ^OIinillSNI~NVINOSHiIlMS S3 I d VM 9 n~L I B R A R I Es'^SMITHSONIA 2 C/5 Z ... C/) O X o l?.l 'V IBRARiES SMITHSONIAN ~ INSTITUTION NOIlDillSNIlillSNI NVINOSHimNVINOSHl CO _ m '^\ or >v = 1 < )0iiniiisNi"'NviN0SHims s3iMVMan libraries^smithsonia r— TT »— -» C/7 _ If) _ IBRARIES SMITHSONIAN INSTITUTION NOIiniliSNI NVINOSHll^^ to Z ' CO z -^,; CO -'^ CO ^^^ O x/ 'x >?',.,> X v-> /-'' z /^-^ t v?,«^.. ^ to ' Z £/) IOIiniliSNI_NVINOSHlll^S SHiyVHSIl LIBRARIES SMITHSONIA 2 "•• </' . r; tn ^ ' -I z _j IBRARIES SMITHSONIAN INSTITUTION NOIinillSNI NVINOSHIIIM — - c/' - CO goiiniiiSNi NviNOSHims saiHVdan libraries smithsonia Z CO Z -,-. to .':/.^/ I /f1 'y^y braries Smithsonian ~ institution NoiiniiiSNi NviNOSHim <.n *fi ~ < Xi>Nr APRIL 1978 Vol. 11, Parts 1/2 Proceedings and Transactions of The British Entomological and Natural History Society iUN90 1978 Price: £3.00 / : Officers and Council for 1978 President: G.
    [Show full text]
  • Program and Abstracts
    PROGRAM AND ABSTRACTS FRONTIERS OF SEXUAL PLANT REPRODUCTION III OCTOBER 17 – 19, 2008 MARRIOTT UNIVERSITY PARK TUCSON, ARIZONA, USA MEETING ORGANISERS: Dr. Ravi Palanivelu, University of Arizona, Tucson, USA Dr. Dmitry Belostotsky, University of Missouri, Kansas City, USA 1 2 MEETING OVERVIEW Friday, October 17, 2008 7:30 am – 8:30 am Breakfast 8:30 am - 9 am Opening Remarks by Dr. Vicki Chandler, Director, BIO5 Institute, University of Arizona. 9 am - 12:15pm Session I: Pollen Development and Tube Growth (Marriott Sabino/Pima rooms) 12 noon - 1 pm Lunch break 1 pm - 2 pm Poster session (please stand by your poster if it is even numbered) 2 pm – 5 pm Session II: Female Gametophyte Development and Function (Marriott Sabino/Pima rooms) 5 pm - 6 pm Refreshments and poster session (please stand by your poster if it is odd numbered) 6 pm – 7 pm Dinner break 7 pm – 10 pm Session III: Endosperm and Imprinting (Marriott Sabino/Pima rooms) Saturday, October 18 2008 7:30 am – 9 am Breakfast 9 pm – 12 noon Session IV: Pollen Pistil Interactions (Marriott Sabino/Pima rooms) 12 noon – 1 pm Lunch break 1 – 2 pm Transport to Sonoran Desert Museum 2 pm - 4 pm Free time - Enjoy the desert museum! 4 pm – 7 pm Session V: Evolution (Desert Museum Warden Oasis Theater) 7 - 10 pm Banquet dinner in desert museum Sunday, October 19, 2008 7:30 am – 8:30 am Breakfast 8:30 am - 11:30 am Session VI: Novel Systems and Emerging Technologies (Marriott Sabino/Pima rooms) 11:30am - 1 pm Lunch break END OF MEETING 3 MEETING SPONSORS AND EXHIBITORS University of Arizona Federal agencies 4 Other sponsors and exhibitors Society for Developmental Biology DEVELOPMENTAL BIOLOGY FSPRIII is partly supported by a grant from the Society for Developmental Biology and by Developmental Biology, the SDB official journal.
    [Show full text]
  • Absence of W Chromosome in Psychidae Moths and Implications for the Theory of Sex Chromosome Evolution in Lepidoptera
    G C A T T A C G G C A T genes Article Absence of W Chromosome in Psychidae Moths and Implications for the Theory of Sex Chromosome Evolution in Lepidoptera 1,2 2 1 1,2, Martina Hejníˇcková , Petr Koutecký , Pavel Potocký , Irena Provazníková y , Anna Voleníková 1,2, Martina Dalíková 1,2 , Sander Visser 1,2, František Marec 1 and Magda Zrzavá 1,2,* 1 Biology Centre of the Czech Academy of Sciences, Institute of Entomology, 37005 Ceskˇ é Budˇejovice, Czech Republic 2 Faculty of Science, University of South Bohemia, 37005 Ceskˇ é Budˇejovice,Czech Republic * Correspondence: [email protected]; Tel.: +420-387775282 Current address: European Molecular Biology Laboratory, Meyerhofstrasse 1, 69117 Heidelberg, Germany. y Received: 8 November 2019; Accepted: 2 December 2019; Published: 5 December 2019 Abstract: Moths and butterflies (Lepidoptera) are the largest group with heterogametic females. Although the ancestral sex chromosome system is probably Z0/ZZ, most lepidopteran species have the W chromosome. When and how the W chromosome arose remains elusive. Existing hypotheses place the W origin either at the common ancestor of Ditrysia and Tischeriidae, or prefer independent origins of W chromosomes in these two groups. Due to their phylogenetic position at the base of Ditrysia, bagworms (Psychidae) play an important role in investigating the W chromosome origin. Therefore, we examined the W chromosome status in three Psychidae species, namely Proutia betulina, Taleporia tubulosa, and Diplodoma laichartingella, using both classical and molecular cytogenetic methods such as sex chromatin assay, comparative genomic hybridization (CGH), and male vs. female genome size comparison by flow cytometry.
    [Show full text]
  • Polygonum Cognatum Meissn. (Madimak) Tohumlarinin
    POLYGONUM COGNATUM MEISSN. (MADIMAK) TOHUMLARININ ÇİMLENME B İYOLOJİSİ ve KLONLARI ARASINDAK İ GENETİK ÇEŞİTLİLİĞİN BELİRLENMESİ Mesut ÖZKURT Y.Lisans Tezi Bitki Koruma Anabilim Dalı Doç. Dr. Hüse yin ÖNEN T.C. GAZİOSMANPAŞA ÜNİVERSİTESİ FEN BİLİMLERİ ENSTİTÜSÜ BİTKİ KORUMA ANABİLİM DALI Y.LİSANS TEZİ POLYGONUM COGNATUM MEISSN. (MADIMAK) TOHUMLARININ ÇİMLENME BİYOLOJİSİ ve KLONLARI ARASINDAKİ GENETİK ÇEŞİTLİLİĞİN BELİRLENMESİ Mesut ÖZKURT TOKAT 2008 Her hakkı saklıdır TEZ BEYANI Tez yazım kurallarına uygun olarak hazırlanan bu tezin yazılmasında bilimsel ahlak kurallarına uyulduğunu, başkalarının eserlerinden yararlanılması durumunda bilimsel normlara uygun olarak atıfta bulunulduğunu, tezin içerdiği yenilik ve sonuçların başka bir yerden alınmadığını, kullanılan verilerde herhangi bir tahrifat yapılmadığını, tezin herhangi bir kısmının bu üniversite veya başka bir üniversitedeki başka bir tez çalışması olarak sunulmadığını beyan ederim. Mesut ÖZKURT Doç. Dr. Hüseyin ÖNEN danışmanlığında, Mesut ÖZKURT tarafından hazırlanan bu çalışma 07/04/2008 tarihinde aşağıdaki jüri tarafından oy birliği/oy çokluğu ile Bitki Koruma Anabilim Dalı’nda Yüksek Lisans tezi olarak kabul edilmiştir. Başkan : Prof. Dr. İzzet KADIOĞLU İmza : Üye : Doç. Dr. Hüseyin ÖNEN İmza : Üye : Yrd. Doç. Dr. İskender PARMAKSIZ İmza : Yukarıdaki sonucu onaylarım Prof. Dr. Metin YILDIRIM Enstitü Müdürü ÖZET Y. Lisans Tezi POLYGONUM COGNATUM MEISSN. (MADIMAK) TOHUMLARININ ÇİMLENME BİYOLOJİSİ ve KLONLARI ARASINDAKİ GENETİK ÇEŞİTLİLİĞİN BELİRLENMESİ Mesut ÖZKURT Gaziosmanpaşa
    [Show full text]
  • Patterns of Endemism and Comparative Phylogeography Confirm Palaeo- Environmental Evidence Forpleistocene Refugia in the Eastern Alps
    52 August 2003: 477–497 Tribsch & Schönswetter Pleistocene refugia in E Alps Patterns of endemism and comparative phylogeography confirm palaeo- environmental evidence forPleistocene refugia in the Eastern Alps Andreas Tribsch 1 & PeterSchö nswetter 2 1Department of Higher Plant Systematics and Evolution, Institute of Botany, University of Vienna, Rennweg 14, A-1030 Vienna, Austria. [email protected] (author for correspondence) 2Department of Chorology and Vegetation Science, Institute of Botany, University of Vienna, Rennweg 14, A- 1030 Vienna, Austria. [email protected] Climatic fluctuations during Quaternary glaciations had a significant influence on the distribution of taxa and on their intraspecific genetic structure. In this paper, we test hypotheses on Pleistocene refugia for mountain plants in the eastern part of the European Alps derived from palaeoenvironmental and geological results, with new data on distributional patterns of 288 vascular plant endemics and molecular phylogeographies of select- ed species. High numbers of endemics are found in calcareous regions at the southern and the eastern border of the Eastern Alps, which remained unglaciated during the Pleistocene. The distribution of local endemic taxa in general, and of silicicolous taxa in particular, shows a clear relationship with hypothetical glacial refugia in the southern, southeastern, easternmost, and northeastern Alps. Molecular phylogeographic data from several silicicolous alpine species ( Androsace alpina , Androsace wulfeniana , Eritrichium nanum , Phyteuma globu- lariifolium , Ranunculus glacialis , Saponaria pumila ) are not completely congruent. However, all genetically defined population groups are in congruence with hypothetical refugia. In general, results from distributions of endemic taxa and data from intraspecific phylogeography are compatible with previously hypothesized refu- gia suggesting that refugial situations have shaped the current patterns.
    [Show full text]