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Ceratodus Tunuensis, Sp. Nov., a New Lungfish (Sarcopterygii, Dipnoi) from the Upper Triassic of Central East Greenland

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Ceratodus Tunuensis, Sp. Nov., a New Lungfish (Sarcopterygii, Dipnoi) from the Upper Triassic of Central East Greenland UJVP #1439834, VOL 0, ISS 0 CERATODUS TUNUENSIS, SP. NOV., A NEW LUNGFISH (SARCOPTERYGII, DIPNOI) FROM THE UPPER TRIASSIC OF CENTRAL EAST GREENLAND FEDERICO L. AGNOLIN,OCTAVIO MATEUS,JESPER MILAN ,MARCO MARZOLA,OLIVER WINGS,JAN SCHULZ ADOLFSSEN, AND LARS B. CLEMMENSEN QUERY SHEET This page lists questions we have about your paper. The numbers displayed at left can be found in the text of the paper for reference. In addition, please review your paper as a whole for correctness. Q1. Au: Please check and confirm that all author names, affiliations, and corresponding author information are correct. Q2. Au: A ZooBank ID is required only if your manuscript introduces new zoological taxa at the family-group level or below. You do this at www.zoobank.org by: 1) creating an account (if you have not already done so); 2) registering the publication; 3) spec- ify an online archive. From the list you should select CLOCKSS; 4) separately registering each new taxon in the publication. Unless you register a taxon the publication will not be properly registered! If you have registered and received a ZooBank ID (LSID) for this manuscript, please provide the ID to the Production Editor: “pub” will appear in the ID string. 5) After the paper is published online you must use the LSID to return to ZooBank and edit the publication registration and change ‘not yet published’ to ‘published’. Q3. Au: GCMK. Location? Q4. Au: GCMK. This abbreviation not used in the paper? Delete? Q5. Au: Muller,€ 1845; Gill, 1873; Agassiz, 1838. Please provide the references. Q6. Au: Please cite Table 1. Q7. Au: Kirkland, 1987. Not listed? Or do you mean Kirkland, 1988? Q8. Au: Martin et al., 1999. Not cited? Q9. Au: Nelson et al., 2016. Place of publication? Q10. Au: Schultze, 1991. Please spell out YPFB if it is normally so. Publisher and place of publication? TABLE OF CONTENTS LISTING The table of contents for the journal will list your paper exactly as it appears below: Ceratodus Tunuensis, sp. nov., a new lungfish (Sarcopterygii, Dipnoi) from the Upper Triassic of central East Greenland FEDERICO L. AGNOLIN,OCTAVIO MATEUS,JESPER MILAN,MARCO MARZOLA,OLIVER WINGS,JAN SCHULZ ADOLFSSEN, AND LARS B. CLEMMENSEN Journal of Vertebrate Paleontology e1439834 (6 pages) Ó by the Society of Vertebrate Paleontology DOI: 10.1080/02724634.2018.1439834 ARTICLE Q1 CERATODUS TUNUENSIS, SP. NOV., A NEW LUNGFISH (SARCOPTERYGII, DIPNOI) FROM THE UPPER TRIASSIC OF CENTRAL EAST GREENLAND FEDERICO L. AGNOLIN,1,2 OCTAVIO MATEUS, *,3,4 JESPER MILAN, 5,6 MARCO MARZOLA, 3,4,7,8 5 OLIVER WINGS,9 JAN SCHULZ ADOLFSSEN,10 and LARS B. CLEMMENSEN 7 1Laboratorio de Anatomıa Comparada y Evolucion de los Vertebrados, Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia,’ Av. Angel Gallardo 470 (C1405DJR), Buenos Aires, Argentina, [email protected]; 2Fundacion de Historia Natural ‘Felix de Azara,’ Departamento de Ciencias Naturales y Antropologıa, CEBBAD–Universidad Maimonides, Hidalgo 775, Piso 7 (C1405BDB), Buenos Aires, Argentina; 10 3GeoBioTec, Departamento de Ciencias^ da Terra, Faculdade de Ciencias^ e Tecnologia, Universidade Nova de Lisboa, Quinta da Torre, 2829-516 Caparica, Portugal, [email protected]; [email protected]; 4Museu da Lourinha,~ Rua Joao~ Luıs de Moura, 95, 2530-158 Lourinha,~ Portugal; 5Geomuseum Faxe/Østsjællands Museum, Højerup Bygade 38, DK-4660 Store Heddinge, Denmark, [email protected]; 6Natural History Museum of Denmark, Øster Voldgade 5–7, DK-1350 Copenhagen K, Denmark; 15 7Department of Geosciences and Natural Resource Management, University of Copenhagen, Øster Voldgade 10, DK-1350 Copenhagen K, Denmark, [email protected]; 8Geocenter Møns Klint, Stengardsvej 8, DK-4751 Borre, Denmark; 9Zentralmagazin Naturwissenschaftlicher Sammlungen (ZNS), Martin-Luther-Universitat€ Halle-Wittenberg, Domplatz 4, 06108 Halle (Saale), Germany; 20 10Ministry of Mineral Resources, Department of Geology, Imaneq 1, DK-3900 Nuuk, Greenland, [email protected] Q2 ABSTRACT—The fossil record of post-Paleozoic lungfishes in Greenland is currently restricted to a few brief reports of isolated and undetermined tooth plates coming from the uppermost Fleming Fjord Formation (late Norian) in Jameson Land, central East Greenland. Here, we describe Ceratodus tunuensis, sp. nov., a new dipnoan from a thin bed of calcareous lake mudstone from the Ørsted Dal Member of the Fleming Fjord Formation. The Ceratodus fossil record indicates that 25 during the Late Triassic, this genus was restricted to the middle latitudes of the Northern Hemisphere. This record matches previous paleobiogeographical analyses and indicates that terrestrial biota during the Late Triassic was strongly influenced by paleolatitude. Citation for this article: Agnolin, F. L., O. Mateus, J. Milan, M. Marzola, O. Wings, J. Schulz Adolfssen, and L. B. Clemmensen. 2018. Ceratodus tunuensis, sp. nov., a new lungfish (Sarcopterygii, Dipnoi) from the Upper Triassic of central 30 East Greenland. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2018.1439834. INTRODUCTION Here, we describe tooth plates and jaw bones belonging to Ceratodus tunuensis, sp. nov., a new lungfish taxon from the Lungfishes have a very long evolutionary history. The oldest uppermost Fleming Fjord Formation. As the first post-Paleozoic 50 fossil record of Dipnoi is Devonian (see Long, 2011), but finds record of fossil lungfishes from Greenland, these specimens add become notably abundant in post-Paleozoic deposits (Martin, valuable information on the genus Ceratodus. 35 1982; Schultze, 2004). Living forms are represented by three gen- Institutional Abbreviations—GCMK—GeoCenter Møns era in South America, Africa, and Australia (Nelson et al., Klint, xx, Denmark; NHMD, Natural History Museum of Den- 2016). mark, Copenhagen, Denmark. 55 Fossil lungfish tooth plates are only relatively common in Q3 Upper Triassic rocks from Europe (Schultze, 2004). Apart from 40 that, European dipnoans still have a patchy fossil record. Several MATERIALS AND METHODS Q4 genera of dipnoans have been described from the Upper Devo- Geological Setting nian of East Greenland (Lehman, 1959; Bendix-Almgreen, 1976), but so far the only post-Paleozoic reports of dipnoans During the Triassic, what is now East Greenland was located have been restricted to brief mentions of isolated tooth plates at the northern rim of the Pangaean supercontinent at about 45 coming from the uppermost Fleming Fjord Formation (late Nor- 40N and bounded to the north by the Boreal Sea (Nøttvedt 60 ian) in Jameson Land (Jenkins et al., 1994; Clemmensen et al., et al., 2008; Clemmensen et al., 2016). Triassic sediments are 1998, 2016). well exposed in the Jameson Land Basin, located in central East Greenland at about 71N in the present-day land areas of Jameson Land and Scoresby Land (Fig. 1). The thickness of the Triassic succession in East Greenland 65 *Corresponding author varies between 1.0 and 1.7 km (Nøttvedt et al., 2008). At the Color versions of one or more of the figures in the article can be found beginning of the Triassic, the Jameson Land Basin is inter- online at www.tandfonline.com/ujvp. preted as a marine bay (Boreal Sea), which gradually Agnolin et al.—New Late Triassic Ceratodontidae from Greenland (e1439834-2) underwent regression and continental emergence up through 70 the Early Triassic (Clemmensen, 1980a, 1980b; Nøttvedt et al., 2008). These overlying sediments are continental and com- prise alluvial fan, river, aeolian dune, saline lake, and freshwa- ter lake deposits (Clemmensen, 1980a; Clemmensen et al., 1998). The prominent freshwater lake deposits from the 75 uppermost part of the Triassic succession (the Fleming Fjord Formation) are particularly well exposed in cliff sides facing the Carlsberg Fjord (Fig. 2). The Fleming Fjord Formation is of Norian–early Rhaetian age, and the fossil-bearing layer in the Carlsberg Ford beds has a likely age of about 208 Ma (see 80 Clemmensen et al., 1998). The new finds of fossil lungfish remains are from a thin (0.1–0.5 m thick) calcareous mudstone bed in the Carlsberg Fjord beds of the Ørsted Dal Member from the uppermost part of the Fleming Fjord Formation (Jenkins et al., 1994; Clemmensen et al., 1998, 2016). 85 The calcareous mudstone bed formed in an ephemeral to semiper- ennial lake. The lungfish remains are associated with numerous freshwater bivalves, undetermined amphibian remains, and frag- ments of testudinatans. Eight specimens were collected during the GeoCenter Møns 90 Klint Dinosaur Expeditions in 2012 and 2016 and are deposited in the NHMD under institutional numbers 115910 to 115917. Terminology In the present article, we follow the terminology of Churcher et al. (2006; see also Churcher and De Iuliis, 2001) for the tooth 95 plates, and that of Martin (1981) for jaw-bone structures. Dis- tinction between upper and lower tooth plates follows the FIGURE 2. A, stratigraphic scheme of the Triassic succession exposed at Carlsberg Fjord. B, field photo showing the reddish-brown bedded lacustrine sediments of the Ørsted Dal Member. criteria detailed by Martin (1980, 1981). The angle between the medial and lingual margins of the tooth plate is termed ‘inner angle’ following Vorobyeva and Minikh (1968). Histological terminology follows Kemp (2001). Measurements were taken 100 as in Apesteguıa et al. (2007). SYSTEMATIC PALEONTOLOGY DIPNOI Muller,€ 1845 CERATODONTIDAE Gill, 1873 CERATODUS Agassiz, 1838 105 CERATODUS TUNUENSIS, sp. nov. (Figs. 3–5) Q5 Holotype—NHMD 11590, a right upper tooth plate lacking the labial half of the first ridge (Fig. 3A). Referred Material—NHMD 11911, an upper left tooth plate 110 with incomplete mesial margin and first ridge (Fig. 3B); FIGURE 1. Location map of the East Greenland showing the Mesozoic HNMD 115912, an upper left tooth plate (Fig. 3C); NHMD deposits in the Jameson Land Basin. A, small insert showing Greenland, with East Greenland indicated by the box. B, geological map of East 115913, a fragmentary left lower tooth plate (Fig. 3D); Greenland, with the Carlsberg Fjord locality indicated by the box.
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