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A Revision of the Species of Geostiba THOMSON and Tropimenelytron

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A Revision of the Species of Geostiba THOMSON and Tropimenelytron © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at Linzer biol. Beitr. 37/2 903-1006 16.12.2005 A revision of the species of Geostiba THOMSON and Tropimenelytron PACE of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae, Aleocharinae) V. ASSING Abstract: Based on an examination of types and non-type material, the previously unrevised species of the Eastern Mediterranean, including all of the Balkans and the southeast of Central Europe, and the Caucasus region are (re-)described and illustrated, among them five species new to science: Geostiba (Tropogastrosipalia) mosorica sp.n. (Croatia), G. (Sipalotricha) incognita sp.n. (Bulgaria), G. (S.) extorta sp.n. (Turkey), G. (S.) gontarenkoi sp.n. (Ukraine), and G. (Sibiota) dinarica sp.n. (Croatia). 21 synonymies are proposed: Geostiba mihoki (BERNHAUER 1932) = C. biharica PACE 1990, syn.n.; G. spinicollis (KRAATZ 1862) = G. croatica EPPELSHEIM 1880, syn.n., = G. cahnthiaca SCHEERPELTZ 1957, syn.n., = G. krapinensis PACE 1990, syn.n.; G. armicollis (BREIT 1917) = G. tergestina PACE 1988, syn.n.; G. apfelbecki EPPELSHEIM 1892 = G. wunderlei PACE 1996, syn.n.; G. spizzana (BERNHAUER 1932) = G. maderi PACE 1996, syn.n.; G. biokovensis PACE 1990 = G. cribripennis PACE 1990, syn.n.; G. tiflisensis PACE 1996 = G. arnica PACE 1996, syn.n.; G. infirma (WEISE 1878) = G. pacei ZERCHE 1988, syn.n.; G. cuneiformis (KRAATZ 1856) = G. gyorffyi (BERNHAUER 1929), syn.n., = G. hcejkai (ROUBAL 1932), syn.n.; G. oertzeni (EPPELSHEIM 1888) = G. tenenbaumi (BERNHAUER 1940), syn.n.; G. zoufali (RAMBOUSEK 1915) = G. optima PACE 1983, syn.n.; G. samai PACE 1977 = G. coiffaili PACE 1983, syn.n.; G. meixneri (BERNHAUER 1910) = G. mostarensis PACE 2002, syn.n.; G. weiratheri PACE 1984 = G. behnei ZERCHE 2002, syn.n.; G. carinicollis (EPPELSHEIM 1878) = G. medea PACE 1996; syn.n.; G. kobrisensis PACE 1996 = G. crucis PACE 1996, syn.n.; G. rhilensis (RAMBOUSEK 1924) = G. bulgarica PACE 1983, syn.n.; G. cingulata (EPPELSHEIM 1878) = G. tbilisensis PACE 1996, syn.n. Three species remain of doubtful identity, because they are represented only by females: G. (Tropogastrosipalia) bernhaueri (BREIT), G. (T.) huberi PACE, and G. (Sibiota) zerchei PACE 1996. Three species are referred to different subgenera: G. zerchei PACE (ex Lioglutosipalia SCHEERPELTZ) to Sibiota CASEY, G. arida (EPPELSHEIM) (ex Trachyglutosipalia SCHEERPELTZ) and G. cingulata (EPPELSHEIM) (ex Chondridiosipalia) both to Sipalotricha SCHEERPELTZ. Lectotypes are designated for Homalota spinicollis KRAATZ, H. chyzeri EPPELSHEIM, H. croatica EPPELSHEIM, Leptusa arida EPPELSHEIM, L bituberculata EPPELSHEIM, L. carinicollis EPPELSHEIM, L cingulata EPPELSHEIM, Sipalia mihoki BERNHAUER, S. hummleri BERNHAUER, S. paganettiana BERNHAUER, S. spizzana BERNHAUER, S. deubeli BERNHAUER, S. kocsii BERNHAUER, S. stussineri BERNHAUER, S. bernhaueri BREIT, S. armicollis BREIT, S. hcejkai ROUBAL, S. carinicollis krzysztofi ROUBAL, S. carinthiaca SCHEERPELTZ, and Geosliba apfelbecki EPPELSHEIM. Numerous additional records of previously revised Geosliba and Tropimenelytron species from the study area are reported. The known Geostiba fauna of the study region now includes 136 species. The highest diversities were observed for Turkey (44 species) and Greece (42 © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at 904 species). The distributions of 27 species are mapped. A catalogue and an updated key to species are provided. Key words: Coleoptera, Staphylinidae, Aleocharinae, Geostiba, Tropimenelytron, Palaearctic region, Central Europe, Balkans, Turkey, Middle East, Caucasus region, taxonomy, revision, new species, new synonyms, catalogue, key to species. Contents 1. Introduction 906 2. Material, methods, and abbreviations 906 3. Revision of the Geostiba species of the Eastern Mediterranean, the Caucasus, and adjacent regions: general results 908 4. New species, «descriptions, and new records 909 4.1. Geostiba {Geosliba) circellaris (GRAVENHORST) 909 4.2. Geostiba (Tropogastrosipalia) chyzeri (EPPELSHEIM) 911 4.3. Geostiba {Tropogastrosipalia) rodopensis PACE 914 4.4. Geostiba {Tropogastrosipalia) bernhaueri (BREIT) 914 4.5. Geostiba {Tropogastrosipalia) mihoki (BERNHAUER) 915 4.6. Geostiba {Tropogastrosipalia) torisuturalis ASSING 917 4.7. Geostiba {Tropogastrosipalia) taygetana (BERNHAUER) 917 4.8. Geostiba {Tropogastrosipalia) meschniggi PACE 917 4.9. Geostiba {Tropogastrosipalia) spinicollis (KRAATZ) 917 4.10. Geostiba {Tropogastrosipalia) hummleri (BERNHAUER) 919 4.11. Geostiba {Tropogastrosipalia) armicollis (BREIT) 920 4.12. Geostiba {Tropogastrosipalia) apfelbecki EPPELSHEIM 921 4.13. Geostiba {Tropogastrosipalia)paganettiana (BERNHAUER) 923 4.14. Geostiba {Tropogastrosipalia) spizzana (BERNHAUER) 923 4.15. Geostiba {Tropogastrosipalia) curzolae (BERNHAUER) 925 4.16. Geostiba {Tropogastrosipalia) biokovensis PACE 926 4.17. Geostiba {Tropogastrosipalia) winkleriana PACE 927 4.18. Geostiba {Tropogastrosipalia) mosorica sp.n 928 4.19. Geostiba {Tropogastrosipalia) slaviankaensis ZERCHE 928 4.20. Geostiba {Tropogastrosipalia) ossogovskaensis ZERCHE 929 4.21. Geostiba {Tropogastrosipalia) ilievi ZERCHE 930 4.22. Geostiba {Tropogastrosipalia) belasizaensis ZERCHE 930 4.23. Geostiba {Tropogastrosipalia) khnzoriani PACE 931 4.24. Geostiba {Tropogastrosipalia) winkleri (BERNHAUER) 931 4.25. Geostiba {Tropogastrosipalia) tißisensis PACE 932 4.26. Geostiba {Tropogastrosipalia) sengleti PACE 933 4.27. Geostiba {Tropogastrosipalia) huberi PACE 934 4.28. Geostiba {Tropogastrosipalia) turcica (BERNHAUER) 934 4.29. Geostiba {Tropogastrosipalia) sinuosa ASSING 934 © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at 905 4.30. Geostiba (Sipalotricha) deubeli (BERNHAUER) 934 4.31. Geostiba (ßipalotricha) infirma (WEISE) 936 4.32. Geostiba (Sipalotricha) cuneiformis (KRAATZ) 938 4.33. Geostiba (Sipalotricha) matajurensis (SCHEERPELTZ) 940 4.34. Geostiba (Sipalotricha) bulbifera ZERCHE 941 4.35. Geostiba (Sipalotricha) incognita sp.n 942 4.36. Geostiba (Sipalotricha) euboica PACE 942 4.37. Geostiba (Sipalotricha) ulcerifera ASSING 943 4.38. Geostiba (Sipalotricha) arida (EPPELSHEIM) 943 4.39. Geostiba (Sipalotricha) extorta sp.n 945 4.40. Geostiba (Sipalotricha) lucens (BENICK) 946 4.41. Geostiba (Sipalotricha) gontarenkoi sp.n 946 4.42. Geostiba (Sipalotricha) cingulala (EPPELSHEIM) 948 4.43. Geostiba (Sibiota) oertzeni (EPPELSHEIM) 949 4.44. Geostiba (Sibiota) zoufali (RAMBOUSEK) 950 4.45. Geostiba (Sibiota) samai PACE 951 4.46. Geostiba (Sibiota) sculpticollis (APFELBECK) 952 4.47. Geostiba (Sibiota) galicicana ASSING 952 4.48. Geostiba (Sibiota) kasyi (SCHEERPELTZ) 952 4.49. Geostiba (Sibiota) meixneri (BERNHAUER) 952 4.50. Geostiba (Sibiota) stussineri (BERNHAUER) 953 4.51. Geostiba (Sibiota) weiratheri PACE 953 4.52. Geostiba (Sibiota) bituberculata (EPPELSHEIM) 954 4.53. Geostiba (Sibiota) carinicollis (EPPELSHEIM) 955 4.54. Geostiba (Sibiota) krzysztofl (ROUBAL) 956 4.55. Geostiba (Sibiota) kobrisensis PACE 957 4.56. Geostiba (Sibiota) zerchei PACE 959 4.57. Geostiba (Sibiota) batumiensis PACE 959 4.58. Geostiba (Sibiota) uhligi PACE 960 4.59. Geostiba (Sibiota) dinarica sp.n 960 4.60. Geostiba (Typhlusida) rhilensis (RAMBOUSEK) 961 4.61. Geostiba (Typhlusida) flava (KRAATZ) 963 4.62. Geostiba scheerpeltziana (FAGEL) 963 4.63. Tropimenelytron tuberiventris (EPPELSHEIM) 963 5. Key to the species of Geostiba and Paraleptusa of the Eastern Mediterranean and the Caucasus region 980 6. Catalogue of the Geostiba species of the Eastern Mediterranean and the Caucasus region 998 7. Acknowledgements 1002 8. Zusammenfassung 1002 9. References 1003 © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at 906 1. Introduction The Geostiba species of the southeast of Central Europe, the southern Balkans, and Turkey have been revised in several steps (ASSING 1999, 2000a, 2000b, 2000c, 2001a, 2001b, 2003, 2004a); The objective of the present paper is a revision of the previously unexamined species from the Caucasus region and from the northern parts of southeast- em Europe. In the 19th and 20th centuries, collecting activity was much higher in the southeast of Central Europe and the northern Balkans than elsewhere in the Eastern Mediterranean, so that numerous Geostiba species were described from that region. Except for most of the Bulgarian species, which were revised by ZERCHE (2002), practically all of these descriptions were made on a more or less additive basis, without revisory work. The known distributions were often either restricted to the respective type localities, very vaguely indicated, or zoogeographically not plausible. The latter is true especially of some species described from the southeast of Central Europe (e. g. G. gyorffyi and G. hcejkai). Moreover, the previous revisions of the Geostiba fauna of the southern Balkans and Turkey had revealed that intraspecific variation had often been underestimated and, in addition, Zoogeographie considerations had often been neglected. In particular, many descriptions were based on the assumption that Geostiba species always have more or less restricted distributions. It turned out, however, that some species (e. g. G. oertzeni, G. lucens, G. euboica) are widespread and at the same time subject to pronounced intraspecific variation, which explains why they now have so many synonyms; in the case of G. oertzeni, the number of junior synonyms amounts to 16 (including a new synonym proposed in this paper). At least three species
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