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Willows of Interior Alaska
1 Willows of Interior Alaska Dominique M. Collet US Fish and Wildlife Service 2004 2 Willows of Interior Alaska Acknowledgements The development of this willow guide has been made possible thanks to funding from the U.S. Fish and Wildlife Service- Yukon Flats National Wildlife Refuge - order 70181-12-M692. Funding for printing was made available through a collaborative partnership of Natural Resources, U.S. Army Alaska, Department of Defense; Pacific North- west Research Station, U.S. Forest Service, Department of Agriculture; National Park Service, and Fairbanks Fish and Wildlife Field Office, U.S. Fish and Wildlife Service, Department of the Interior; and Bonanza Creek Long Term Ecological Research Program, University of Alaska Fairbanks. The data for the distribution maps were provided by George Argus, Al Batten, Garry Davies, Rob deVelice, and Carolyn Parker. Carol Griswold, George Argus, Les Viereck and Delia Person provided much improvement to the manuscript by their careful editing and suggestions. I want to thank Delia Person, of the Yukon Flats National Wildlife Refuge, for initiating and following through with the development and printing of this guide. Most of all, I am especially grateful to Pamela Houston whose support made the writing of this guide possible. Any errors or omissions are solely the responsibility of the author. Disclaimer This publication is designed to provide accurate information on willows from interior Alaska. If expert knowledge is required, services of an experienced botanist should be sought. Contents -
Vascular Plants of Kluane
26 Blueleaved strawberry Fragaria virginiana 63 Greyleaf willow Salix glauca Kluane National Park and Reserve 27 Bog blueberrry Vaccinium uliginosum 64 Ground cedar, Lycopodium complanatum 28 Bog labrador-tea Ledum groenlandica Creeping jenny 65 Hairy rockcress Arabis hirsuta 29 Boreal aster Aster alpinus 30 Boreal wormwood Artemisia arctica 66 Heart-leaf listera Listera borealis Vascular 31 Bristly stickseed Lappula myosotis 67 Heartleaf arnica Arnica cordifolia 32 Broadglumed wheatgrass Agropyron trachycaulum 68 High bush cranbery Viburnum edule Plants List 33 Broadleaf lupine Lupinus arcticus 69 Holboell's rockcress Arabis holboellii 34 Buffaloberry, Soapberry Sheperdia canadensis 70 Horned dandelion Taraxacum lacerum 35 Canada butterweed Senecio pauperculus 71 Kotzebue's grass-of- Parnassia kotzebuei 36 Chestnut rush Juncus castaneus parnassus 1 Alaska moss heath Cassiope stelleriana 37 Cleft-leaf groundsel Senecio streptanthifolius 72 Kuchei's lupine Lupinus kuschei 2 Alaska willow Salix alaxensis 38 Common horsetail Equisetum arvense 73 Labrador lousewort Pedicularis labradorica 3 Alkali bluegrass Poa juncifolia 39 Common mountain Juniperus communis 74 Lance-leaved draba Draba lanceolata 4 Alkali grass Puccinellia interior juniper 75 Lanceleaved stonecrop Sedum lanceolatum 5 Alpine bluegrass Poa alpina 40 Cow parsnip Heracleum lanatum 76 Lapland cassiope Cassiope tetragona 6 Alpine fescue Festuca ovina 41 Creeping juniper Juniperus horizontalis 77 Leafless pyrola Pyrola asarifolia 7 Alpine milk-vetch Astragalus alpinus 42 Creeping -
Chapter 10 • Principles of Conserving the Arctic's Biodiversity
Chapter 10 Principles of Conserving the Arctic’s Biodiversity Lead Author Michael B. Usher Contributing Authors Terry V.Callaghan, Grant Gilchrist, Bill Heal, Glenn P.Juday, Harald Loeng, Magdalena A. K. Muir, Pål Prestrud Contents Summary . .540 10.1. Introduction . .540 10.2. Conservation of arctic ecosystems and species . .543 10.2.1. Marine environments . .544 10.2.2. Freshwater environments . .546 10.2.3. Environments north of the treeline . .548 10.2.4. Boreal forest environments . .551 10.2.5. Human-modified habitats . .554 10.2.6. Conservation of arctic species . .556 10.2.7. Incorporating traditional knowledge . .558 10.2.8. Implications for biodiversity conservation . .559 10.3. Human impacts on the biodiversity of the Arctic . .560 10.3.1. Exploitation of populations . .560 10.3.2. Management of land and water . .562 10.3.3. Pollution . .564 10.3.4. Development pressures . .566 10.4. Effects of climate change on the biodiversity of the Arctic . .567 10.4.1. Changes in distribution ranges . .568 10.4.2. Changes in the extent of arctic habitats . .570 10.4.3. Changes in the abundance of arctic species . .571 10.4.4. Changes in genetic diversity . .572 10.4.5. Effects on migratory species and their management . .574 10.4.6. Effects caused by non-native species and their management .575 10.4.7. Effects on the management of protected areas . .577 10.4.8. Conserving the Arctic’s changing biodiversity . .579 10.5. Managing biodiversity conservation in a changing environment . .579 10.5.1. Documenting the current biodiversity . .580 10.5.2. -
A Vegetative Key to the Willows of Colorado© by Gwen Kittel
A Vegetative Key to the Willows of Colorado© By Gwen Kittel (March 6, 2016) This key uses only vegetative characteristics that are mature (or fully expanded). As The characteristics of willows are highly variable, be sure to look at several leaves and twigs on one shrub to get an overall feel (an average) for the size, shape and color. Leaf characteristics and measurements refer to the leaf blade only, unless otherwise noted. Ignore stipules (paired bracts at the base of leaf petioles) and leaves of sucker shoots (unusually large). Stipules occur on most Salix species, are very temporary and do not aid in distinguishing among them. Sucker shoots are vigorous, non-flowering twigs that grow from the base of the shrub, and can have really huge leaves, far beyond the parameters listed here. While this key is based only on vegetative characters, distinguishing catkin characteristics are occasionally noted. Two catkin characteristics are useful even when the catkin is over mature: the stipe length and the catkin length. I often find catkins late in the season still dangling on the shrub, or lying on the ground underneath the shrub. Key to Groups 1. Trees. Either large shade trees or at least tall, large diameter single trunk (occasionally split into two or more) up 30 feet (10 meters) in height, generally at lower elevations on the eastern plains and western valleys. ............................................................................................... Group A 1. Shrubs. Usually multi-stemmed, if single trunk than diameter not more than 10 inches (5 cm). Height from less than 10 cm creeping on the ground to large shrubs of stream banks and floodplains, up to 15 feet (5 meters) tall .........................................................................................2 2. -
Guide to the Willows of Shoshone National Forest
United States Department of Agriculture Guide to the Willows Forest Service Rocky Mountain Research Station of Shoshone National General Technical Report RMRS-GTR-83 Forest October 2001 Walter Fertig Stuart Markow Natural Resources Conservation Service Cody Conservation District Abstract Fertig, Walter; Markow, Stuart. 2001. Guide to the willows of Shoshone National Forest. Gen. Tech. Rep. RMRS-GTR-83. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station. 79 p. Correct identification of willow species is an important part of land management. This guide describes the 29 willows that are known to occur on the Shoshone National Forest, Wyoming. Keys to pistillate catkins and leaf morphology are included with illustrations and plant descriptions. Key words: Salix, willows, Shoshone National Forest, identification The Authors Walter Fertig has been Heritage Botanist with the University of Wyoming’s Natural Diversity Database (WYNDD) since 1992. He has conducted rare plant surveys and natural areas inventories throughout Wyoming, with an emphasis on the desert basins of southwest Wyoming and the montane and alpine regions of the Wind River and Absaroka ranges. Fertig is the author of the Wyoming Rare Plant Field Guide, and has written over 100 technical reports on rare plants of the State. Stuart Markow received his Masters Degree in botany from the University of Wyoming in 1993 for his floristic survey of the Targhee National Forest in Idaho and Wyoming. He is currently a Botanical Consultant with a research emphasis on the montane flora of the Greater Yellowstone area and the taxonomy of grasses. Acknowledgments Sincere thanks are extended to Kent Houston and Dave Henry of the Shoshone National Forest for providing Forest Service funding for this project. -
Blueberry Willow): a Technical Conservation Assessment
Salix myrtillifolia Anderss. (blueberry willow): A Technical Conservation Assessment Prepared for the USDA Forest Service, Rocky Mountain Region, Species Conservation Project June 29, 2006 Stephanie L. Neid, Ph.D., Karin Decker, and David G. Anderson Colorado Natural Heritage Program Colorado State University Fort Collins, CO Peer Review Administered by Society for Conservation Biology Neid, S.L., K. Decker, and D.G. Anderson. (2006, June 29). Salix myrtillifolia Anderss. (blueberry willow): a technical conservation assessment. [Online]. USDA Forest Service, Rocky Mountain Region. Available: http://www.fs.fed.us/r2/projects/scp/assessments/salixmyrtillifolia.pdf [date of access]. ACKNOWLEDGMENTS This research was greatly facilitated by the helpfulness and generosity of many experts, particularly George Argus, Robert Dorn, Bud Kovalchik, Steve Brunsfield, John Sanderson, and Barry Johnston. Their interest in the project, valuable insight, depth of experience, and time spent answering questions were extremely valuable and crucial to the project. Herbarium specimen label data were provided by Tim Hogan (COLO), Ron Hartman and Ernie Nelson (RM), and Janet Wingate (KDH). Thanks also to Cathy Cripps, Bonnie Heidel, George Jones, Chris Chappel, Rex Crawford, Florence Caplow, Scott Mincemoyer, Andy Kratz, Kathy Carsey, and Beth Burkhart for assisting with questions and project management. Jane Nusbaum, Dawn Taylor, and Barbara Brayfield provided crucial financial oversight. AUTHORS’ BIOGRAPHIES Stephanie L. Neid is an ecologist with the Colorado Natural Heritage Program (CNHP). Her work at CNHP includes ecological inventory and assessment throughout Colorado, beginning in 2004. Prior to this, she was an ecologist with the New Hampshire Natural Heritage Bureau and the Massachusetts Natural Heritage and Endangered Species Program and was a Regional Vegetation Ecologist for NatureServe. -
George Argus
Skvortsovia: 1(2): 99–111 (2014) Skvortsovia ISSN 2309-6497 (Print) Copyright: © 2014 Russian Academy of Sciences http://skvortsovia.uran.ru/ ISSN 2309-6500 (Online) Editorial George W. Argus, salicologist for life Irina Belyaeva1,2* and Keith Chamberlain3 1 Royal Botanic Gardens, Kew, Richmond, TW9 3AE, UK 2 Russian Academy of Sciences, Ural Branch: Institute Botanic Garden, 8 Marta, 202A, 620144, Yekaterinburg, Russia 3 Rothamsted Research, Harpenden, AL5 2JQ, UK *Corresponding author. Email: [email protected] Published on line: 30 April 2014 This April saw the 85th birthday of one of the world’s greatest salicologists, George W. Argus. Despite having retired, officially, almost 20 years ago he continues his scientific research, editorial work and the identification of specimens at the Canadian Museum of Nature in Ottawa where he is Professor and Curator Emeritus. He also runs workshops on the identification of willows anywhere that he is invited to do so. However, he did not start his career as a botanist, having studied engineering at Valparaiso University, Indiana, and subsequently worked as a labourer on the Alaskan Railroad and a lineman’s assistant in the Alaskan gold mines. It was in this special environment of mountains and glaciers that he Ottawa, local field trip, 2006 developed a lifelong interest in natural history, especially in geology and palaeontology and completed his undergraduate degree at the University of Alaska in biology and geology in 1952. After a brief spell as a surveyor on the Juneau Icefield Research Project George entered the US army as an instructor in arctic techniques. He was a founder of the Alaska Alpine Club and while he was in the army, in April, 1954, George joined with three others in an expedition to climb Mount McKinley (Denali) in what was to become an impressive story of survival in extreme conditions. -
Height and Growth Rings of Salix Lanata Ssp. Richardsonii Along the Coastal Temperature Gradient of Northern Alaska
Height and growth rings of Salix lanata ssp. richardsonii along the coastal temperature gradient of northern Alaska D. A. WALKER Institute of Arctic and Alpine Research, UniversiQ of Colorado, Boulder, CO, U.S.A. 80309 Received March 14, 1986 WALKER,D. A. 1987. Height and growth rings of Salix lanata ssp. richardsonii along the coastal temperature gradient of northern Alaska. Can. J. Bot. 65: 988-993. Salix lanata ssp. richardsonii from open-tundra and streamside populations were studied at seven sites along a 100-km north-south transect following the Sagavanirktok River from the Alaskan arctic coast to the foothills of the Brooks Range. Mean July temperatures along this transect vary from 2.6 at the coast to 10°C at the base of the foothills. Mean maximum heights of the sampled open-tundra willows increased from 10 + 2 at the coast to 37 + 8 cm at the southern end of the transect. Mean maximum heights of sampled streamside willows increased from 0 at the coast to 147 i 25 cm. The mean maximum height of willows in both habitats showed very strong correlations with thawing degree-days. Mean growth-ring widths increased from 92 + 20 at the coast to 188 i 57 pm at the southern end of the transect and were also highly correlated with the temperature gradient. The results are discussed in light of other arctic studies of willow growth rings and Cantlon's system of vegetation subdivisions within the Alaskan arctic. WALKER,D. A. 1987. Height and growth rings of Salix lanata ssp. richardsonii along the coastal temperature gradient of northern Alaska. -
Routt National Forest Riparian Vegetation Classification
• Routt National Forest Riparian Vegetation Classification January 12, 1996 Prepared for: • Routt National Forest 29587 West U.S. 40, Suite 20 Steamboat Springs, CO 80487 Prepared by: Steve Kettler - Ecologist Amy McMullen - Research Associate Colorado Natural Heritage Program 254 General Services Building Colorado State University Ft. Collins, CO 80523 • Table of Contents • EXECUTIVE SUMMARY ...........•........••.................... , IV ACKNOWLEDGEMENTS V INTRODUCTION ............................................... 1 Colorado's Natural Heritage Program . .. 2 STUDY AREA DESCRIPTION . .. 4 METHODS. .. 6 Representative site selection .................................. 6 Vegetation and environmental data collection . .. 7 Classification . .. 8 RESULTS .................................................... 10 Key to Routt National Forest Riparian Plant Associations . .. 10 Evergreen Forests ............................................. 20 Abies lasiocarpa Series ..................................... 20 Abies lasiocarpa/Alnus incana-Cornus sericea p.a. ............ 20 Abies lasiocarpa/Alnus incana-Salix drummondiana p.a.......... 21 Abies lasiocarpa/Calamagrostis canadensis p.a. .............. 22 • Abies lasiocarpa/Mertensia ciliata p.a. .................... 24 Abies lasiocarpa-Picea engelmanniilActaea rubra p.a............ 25 Abies lasiocarpa-Picea engelmanniilRibes spp. p.a.............. 26 PIcea· engeI" mannn Senes . .................................... 27 Picea engelmanniilCarex aquatilis p.a. .................... 27 Picea engelmanniilEquisetum -
Willows of Southcentral Alaska
1 Willows of Southcentral Alaska Dominique M. Collet Kenai Watershed Forum 2002 2 Willows of Southcentral Alaska Acknowledgements The development of this willow guide has been made possible thanks to funding from the Cook Inlet Coastal Program, U.S. Fish and Wildlife Service– Grant 70181-1-G092. Additional funding for printing was made available through a collabo- rative partnership of the US Forest Service, the Alaska Department of Transpor- tation, the Plant Materials Center of the Alaska Department of natural Re- sources, the Kenai Peninsula Borough through the Coastal Impact Assistance Program of the National Oceanic and Atmospheric Administration, and the Alaska Native Plant Society. Many individuals have contributed to this guide. Special thanks go to John DeLapp (USFWS) for his support from the beginning of the project, Robert Lipkin (Alaska Natural Heritage Program) who helped initiate and define the project, Robert Ruffner (Kenai Watershed Forum) for his administrative support, George Argus, who has been the leading teacher and mentor to the author, and numerous willow students in Alaska. Suggestions and editing notes from Ed Berg, Kelley Shea, and Pauline Simmons provided much improvement to the original manuscript. Field-testing was expedited through the assistance of John Mohorchich of the Kenai River Center, where numerous landowners and contractors seek assistance with revegetation projects. The data for the distributions maps were provided by George Argus, Al Batten, Garry Davies, Rob deVelice, Mike Gracz, Carolyn Parker, and Mike Tetreau. Special thanks go to Roy Baldwin for his information on diamond willows. Most of all, I am especially grateful to Pamela Houston, whose support made the writing of this guide possible. -
Twenty of the Most Thermophilous Vascular Plant Species in Svalbard and Their Conservation State
Twenty of the most thermophilous vascular plant species in Svalbard and their conservation state Torstein Engelskjøn, Leidulf Lund & Inger Greve Alsos An aim for conservation in Norway is preserving the Svalbard archi- pelago as one of the least disturbed areas in the Arctic. Information on local distribution, population sizes and ecology is summarized for 20 thermophilous vascular plant species. The need for conservation of north- ern, marginal populations in Svalbard is reviewed, using World Conser- vation Union categories and criteria at a regional scale. Thirteen species reach their northernmost distribution in Svalbard, the remaining seven in the western Arctic. Nine species have 1 - 8 populations in Svalbard and are assigned to Red List categories endangered or critically endangered: Campanula rotundifolia, Euphrasia frigida, Juncus castaneus, Kobresia simpliciuscula, Rubus chamaemorus, Alchemilla glomerulans, Ranuncu- lus wilanderi, Salix lanata and Vaccinium uliginosum, the last four spe- cies needing immediate protective measures. Five species are classifi ed as vulnerable: Betula nana, Carex marina ssp. pseudolagopina, Luzula wahlenbergii, Ranunculus arcticus and Ranunculus pallasii. Six species are considered at lower risk: Calamagrostis stricta, Empetrum nigrum ssp. hermaphroditum, Hippuris vulgaris (only occurring on Bjørnøya), Juncus triglumis, Ranunculus lapponicus and Rhodiola rosea. The warmer Inner Arctic Fjord Zone of Spitsbergen supports most of the 20 target species and is of particular importance for conservation. Endan- gered or vulnerable species were found in a variety of edaphic conditions; thus, several kinds of habitats need protection. T. Engelskjøn, I. G. Alsos, Tromsø Museum, University of Tromsø, NO-9037 Tromsø, Norway, torstein@ tmu.uit.no; L. Lund, Phytotron, University of Tromsø, NO-9037 Tromsø, Norway. -
Rare Vascular Plants of the North Slope a Review of the Taxonomy, Distribution, and Ecology of 31 Rare Plant Taxa That Occur in Alaska’S North Slope Region
BLM U. S. Department of the Interior Bureau of Land Management BLM Alaska Technical Report 58 BLM/AK/GI-10/002+6518+F030 December 2009 Rare Vascular Plants of the North Slope A Review of the Taxonomy, Distribution, and Ecology of 31 Rare Plant Taxa That Occur in Alaska’s North Slope Region Helen Cortés-Burns, Matthew L. Carlson, Robert Lipkin, Lindsey Flagstad, and David Yokel Alaska The BLM Mission The Bureau of Land Management sustains the health, diversity and productivity of the Nation’s public lands for the use and enjoyment of present and future generations. Cover Photo Drummond’s bluebells (Mertensii drummondii). © Jo Overholt. This and all other copyrighted material in this report used with permission. Author Helen Cortés-Burns is a botanist at the Alaska Natural Heritage Program (AKNHP) in Anchorage, Alaska. Matthew Carlson is the program botanist at AKNHP and an assistant professor in the Biological Sciences Department, University of Alaska Anchorage. Robert Lipkin worked as a botanist at AKNHP until 2009 and oversaw the botanical information in Alaska’s rare plant database (Biotics). Lindsey Flagstad is a research biologist at AKNHP. David Yokel is a wildlife biologist at the Bureau of Land Management’s Arctic Field Office in Fairbanks. Disclaimer The mention of trade names or commercial products in this report does not constitute endorsement or rec- ommendation for use by the federal government. Technical Reports Technical Reports issued by BLM-Alaska present results of research, studies, investigations, literature searches, testing, or similar endeavors on a variety of scientific and technical subjects. The results pre- sented are final, or a summation and analysis of data at an intermediate point in a long-term research project, and have received objective review by peers in the author’s field.