Budapest 1982
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ANNALES HISTORICO-NATURALES MUSEI NATIONALIS HUNGARICI Tomus 74. Budapest, 1982 p. 85-91. Some notes on South Siberian Trechini (Coleoptera, Carabidae) with descriptions of three new species by V. G. SHILENKOV, Irkutsk Abstract — Three new species of Trechus from southern Siberia and northern Mongolia are described. Some new data on the taxonomy and distribution of other southern Siberian Trechini, as well as identification key to the species are given. With 17 figures. The ground-beetle fauna of Southern Siberia (from Altai to Transbaikal region) is not known adequately, whereas this interesting territory with expansive mountain systems appears to be a centre of endemism for some groups of insects. As for Trechini, only two species of Epaphius, six species of Trechus (three species described below) and one species of Altaiotrechus are now known from this re gion. Perhaps the Trechini fauna was severely affected during Pleistocene ice period. But undoubtedly, subsequent investigations of alpine and subalpine zones of Altai and Sajan Mountains will bring new species into science. In the present paper author will introduce into science three new species of Trechus from South Siberia and Northern Mongolia. Some new data on the taxonomy and distribution of other South Siberian Trechini will also be given. Before going further I wish to express my heartfull thanks to DR. Z. KASZAB for his indispensible support of this study, to DR. B. A. KOROTYAEV (Leningrad) and DR. V. G. MORDKOVITSCH (Novosi birsk) for kindly loaning specimens, and DR. S. I. KELEJNIKOVA (Moskow), who has provided facilites for studying type specimens of V. MOTSCHULSKY'S collection. The following abbreviations will be used below : ZIL — Zoological Institute, Academy of Sciences, Leningrad. ZMM — Zoological Museum of Moscow State University. HNHM — Hungarian Natural History Museum, Budapest. ISU — collection of V. G. SHILENKOV, Irkutsk State University. MN - collection of V. G. MORDKOVITSCH, Institut of Soil and Agrochemistry, Novosibirsk. IDENTIFICATION KEY TO THE SPECIES 1 (4) Stria 3 with three discal setiferous pores. 2 (3) Pronotum (fig. 1) strongly rounded at sides, with very obtuse hind angles, basal part prominent behind them level Epaphius (Epaphius) secalis (PAYK.) 3 (2) Pronotum (fig. 2) with rectangular or slightly acute hind angles, its basal part normal Epaphius (Epaphiolus) rivularis (GYLL.) 4 (1) Stria 3 with two dorsal setiferous pores. 5 (6) Larger (5.0-6.5 mm). Pronotum (fig. 3) with rectangular or slightly acute hind angles, basal margin stright, lateral sides rather widely reflexed, especially at basal half. Striae deep, strongly punctate Trechus rubens (F.) 6 (5) Smaller (2,9-3,6 mm). Hind angles of pronotum obtuse or rounded, if slightly acute, basal margin sinuated. Striae shallow, smooth or indistinct punctate. 7 (12) Eyes small, flat, scarcely longer or equal temporae. Aedeagus at apex with hook, curved dorsally. Fore tibiae without groove on external face. Mountaineous tundra of Altai, Sajan and Changai Mts. Annls hist.-nat. Mus. natn. hung., 74,1982 8 (9) Fore tibiae pubescent on external face. Recurrent striae free. Hind angles of pro notum obtuse, but extreme tip feebly prominent as small tooth. Altai Altaiotrechus alticola IABL. — KHNZ. 9 (8) Fore tibiae glabrous on external face. Recurrent stria joining stria 5. Hind angles of pronotum obtuse with rounded apex. 10 (11) Apical hook of aedeagus long and strongly curved dorsally (fig. 11). East Sajan Mts. Trechus almonius REITT. 11 (10) Apical hook of aedeagus short and feebly curved dorsally (fig. 12). Mongolia, Changai Mts (fig. 4) Trechus hangaicus sp. n. 12 (7) Eyes larger, more protrudent, longer than temporae. Aedeagus without apical hook. Fore tibiae with groove, or at least with its trace near apex on external face. 13 (14) Aedeagus (fig. 13) massive, thickened at apex, inner sac with two spines and groups of small teeth. Pronotum narrow, with obtuse hind angles. Uniformly pale-brown. Tannu-Ola Mts. (fig. 6) Trechus korotyaevi sp. n. 14 (13) Aedeagus (figs 8-10) smaller, with well developed copulatory piece and tufts of long teeth. Body colour darker. 15 (16) Pitchy-brown. Pronotum narrower, 1.32-1.37 times as wide as long, hind angles prominent as short acute teeth. Aedeagus (fig. 10) with slightly thickened tip, copu latory piece as long bar. West Sajan Mts. (fig. 7) Trechus mordkovitschi sp. n. 16 (15) Dark-brown. Pronotum wider, 1.40-1.47 times as wide as long, hind angles obtuse, no prominent. Aedeagus (figs. 8-9) with simple tip, inner sac armed two discoid plates and long bar, bifurcate at apex (fig. 5) Trechus montanus MÖTSCH. Epaphius (Epaphius) secalis (PAYKULL, 1790) Carabus Secalis PAYKULL, 1790: Monogr. Carab. Suec. : 94 (type locality: Sweden). Trechus laticollis MOTSCHULSKY, 1844: 235 (type locality: village Turka on the shore of Baikal). For other synonyms and bibliography, see JEANNEL, 1927: 132, 1941: 317; CSIKI, 1928: 245; LINDROTH, 1945: 666; BURAKOWSKI & al., 1973: 179; PAWLOWSKI, 1975: 30. Distribution. A widely distributed Euro-Siberian species restricted in the East by Lena river and Transbaikal region. Ecology. Inhabits several types of birch forests, sometimes in lowland forests and gardens, rare in open country with grass cover, near water. Material. Novosibirsk distr. : environs of Barabinsk, village Karachi (MN, ISU). Irkutsk distr.: Irkutsk (ISU, ZIL); Bolshaja Glubokaja (Shelekhov env.) (ISU); Troitzk (Zalari env.) (ISU); Vydrino (ISU), Kultuk (ZIL) (Sljudjanka env.); Elantzy (ISU), Kosaja Stepp (ZIL) (Olkhon env.); Belaja river (ZIL). Burjatskaja ASSR: Iljinka (ISU). Taxomical remark. Lectotype, rf of Trechus laticollis MÖTSCH. (ZMM), designated here, bears two labels: "L. Baikal" and "Trechus laticollis mihi, Turkinsk". This specimen is, however, in fact, an Epaphius secalis (PAYK.) with slightly monstrous pronotum. Synonymy was ascertained for the first time by JEANNEL (1927: 132). Epaphius (Epaphiolus) rivularis (GYLLENHAL, 1810) Bembidium rivulare GYLLENHAL, 1810, Ins. Suec, II: 33 (type locality : Sweden: Kinnekulle). Trechus nigricomis MOTSCHULSKY, 1844: 235, tab. X, fig. 5-5', syn. n. (type locality: Kiachta). For other synonyms, distribution and ecology, see JEANNEL, 1927: 138; CSIKI, 1928: 244; LINDROTH, 1945: 661; BURAKOWSKI & all, 1973: 197; PAWLOWSKI, 1975: 27. Distribution. Until recently it had been known only from Europe, where it is widely distributed in the North and very sporadically in the mountains of Central Europe. In fact it possesses a Euro-Siberian area of distribution. It was found in South Baikal region (SHILENKOV, 1979: 41), and now I have specimens from other parts of Siberia. Ecology. This species prefers marshes with Sphagnum and lowland forests with soft soil rich in organic matters (LINDROTH, 1945, PAWLOWSKI, 1975). Immature beetles 16. VI. (Iljinka). Material. Novosibirsk distr. : environs ot Barabinsk, village Karachi, 1-13. VIII. 1973, leg. V. G. MORDKOVITSCH, 7 ex (MN, ISU). Irkutsk distr. : Bolshaja Murozhnaja river, tributary of Anga ra river, birch forest, 23-26. VII. 1978, leg. V. K. DMITRIJENKO, 2 ex. (ISU); the same locality, birch and silver fir forest, 18. VIII. 1978, leg. V. K. DMITRIJENKO, 1 ex. (ISU), Burjatskaja ASSR: Iljinka settlement, 16. VI-28. VII. 1975, leg. V. G. SHILENKOV and A. VOINKOV, 3 ex. (ISU). Taxonomical remarks. Lectotype, Ç of Trechus nigricornis MÖTSCH. (ZMM), designated here, bears two labels: "Kiachta" and "Trechus nigricornis mihi, 10 v. v. Kiach.". Length 3.5 mm. This is an immature specimen of Epaphius rivularis (GYLL.). JEANNEL (1928: 792) presumed, that T. nigricornis MÖTSCH, is related to or synonymous with T. montanus MÖTSCH., but short description by MOTSCHULSKY was not sufficient for exact conclusion. Trechus (Trechus) rubens (FABRICIUS, 1792) Carabus rubens FABRICIUS, 1792, Entomol. syst, emend, et aucta, I, 1 (type locality: Germany: Kiel). For synonymy, distribution and ecology, see JEANNEL, 1927: 180, 1941: 319; CSIKI, 1928: 247; LINDROTH, 1945: 664, 1961: 194; PAWLOWSKI, 1975: 50; SHILENKOV, 1979: 41. Distribution. Borei-montaneous in Europe, in Siberia to the East as far as Lena river and Baikal. Introduced in Canada. Material. Irkutsk distr.: "Irkutsk province", 31. V. 1907, leg. DR. SMIRNOV, 2 ex. (ZIL); Baikalsk, in open forest under stones near small stream, 16. VII. 1980, leg. A. D. SHILENKOVA, 2 ex. <1SU). Figs 1-3. Pronotum. 1 : Epaphius secalis (PAYK.) ; 2 : Epaphius rivularis (GYLL.) ; 3 : Trechus rubens (F.). — Figs 4-7. Habitus 4: Trechus hangaicus sp. n.; 5: Trechus montanus MÖTSCH.; 6: Trechus koro- tyaevisp. n.; 7: Trechus mordkovitschisp. n. / Trechus (Trechus) montanus MOTSCHULSKY, 1844. Trechus montanus MOTSCHULSKY, 1844: 236, tab. X, fig. 6-6' (type locality : mountains to the South of Baikal). Trechus montanus, J. SAHLBERG, 1880: 21. Trechus curvatus J. SAHLBERG, 1880: 20 (type locality: vicinities of Krasnojarsk and Fatjanovsk). Trechus perminutus REITTER, 1898: 112 (type locality: source of Irkut). Trechus perminutus, REITTER, 1903: 5. Trechus (s. str.) montanus, JEANNEL, 1927: 301, fig. 806-809. Trechus (s. str.) montanus, CSIKI, 1928: 258. Trechus montanus, JEANNEL, 1962: 87, 89, fig.2-4 . Trechus montanus, SHILENKOV, 1979: 40. Distribution. After Jeannel (1927: 302, fig. 806, map), widely distributed in Siberia from Ob river to Transbaikal region and Northern Mongolia (Changai Mts.). Perhaps it is farther distributed to the East, in any case I have seen specimens from Jakutija. Material. Krasnojarsk distr. (ISU). Irkutsk distr.: