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Defining the Genus Homo
Defining the Genus Homo Mark Collard and Bernard Wood Contents Introduction ..................................................................................... 2108 Changing Interpretations of Genus Homo ..................................................... 2109 Is Genus Homo a “Good” Genus? ............................................................. 2114 Updating Wood and Collard’s (1999) Review of Genus Homo .............................. 2126 Conclusion ...................................................................................... 2137 Cross-References ............................................................................... 2138 References ...................................................................................... 2138 Abstract The definition of the genus Homo is an important but under-researched topic. In this chapter we show that interpretations of Homo have changed greatly over the last 150 years as a result of the incorporation of new fossil species, the discovery of fossil evidence that changed our perceptions of its component species, and reassessments of the functional capabilities of species previously allocated to Homo. We also show that these changes have been made in an ad hoc fashion. Criteria for recognizing fossil specimens of Homo have been outlined on a M. Collard (*) Human Evolutionary Studies Program and Department of Archaeology, Simon Fraser University, Burnaby, BC, Canada Department of Archaeology, University of Aberdeen, Aberdeen, UK e-mail: [email protected] B. Wood Center for the Advanced -
West Indian Manatee (Trichechus Manatus) Habitat Characterization Using Side-Scan Sonar
Andrews University Digital Commons @ Andrews University Master's Theses Graduate Research 2017 West Indian Manatee (Trichechus Manatus) Habitat Characterization Using Side-Scan Sonar Mindy J. McLarty Andrews University, [email protected] Follow this and additional works at: https://digitalcommons.andrews.edu/theses Part of the Biology Commons Recommended Citation McLarty, Mindy J., "West Indian Manatee (Trichechus Manatus) Habitat Characterization Using Side-Scan Sonar" (2017). Master's Theses. 98. https://digitalcommons.andrews.edu/theses/98 This Thesis is brought to you for free and open access by the Graduate Research at Digital Commons @ Andrews University. It has been accepted for inclusion in Master's Theses by an authorized administrator of Digital Commons @ Andrews University. For more information, please contact [email protected]. ABSTRACT WEST INDIAN MANATEE (TRICHECHUS MANATUS) HABITAT CHARACTERIZATION USING SIDE-SCAN SONAR by Mindy J. McLarty Chair: Daniel Gonzalez-Socoloske ABSTRACT OF GRADUATE STUDENT RESEARCH Thesis Andrews University School of Arts and Sciences Title: WEST INDIAN MANATEE (TRICHECHUS MANATUS) HABITAT CHARACTERIZATION USING SIDE-SCAN SONAR Name of researcher: Mindy J. McLarty Name and degree of faculty chair: Daniel Gonzalez-Socoloske, Ph.D. Date completed: April 2017 In this study, the reliability of low cost side-scan sonar to accurately identify soft substrates such as grass and mud was tested. Benthic substrates can be hard to classify from the surface, necessitating an alternative survey approach. A total area of 11.5 km2 was surveyed with the sonar in a large, brackish mangrove lagoon system. Individual points were ground-truthed for comparison with the sonar recordings to provide a measure of accuracy. -
Homo Habilis
COMMENT SUSTAINABILITY Citizens and POLICY End the bureaucracy THEATRE Shakespeare’s ENVIRONMENT James Lovelock businesses must track that is holding back science world was steeped in on surprisingly optimistic governments’ progress p.33 in India p.36 practical discovery p.39 form p.41 The foot of the apeman that palaeo ‘handy man’, anthropologists had been Homo habilis. recovering in southern Africa since the 1920s. This, the thinking went, was replaced by the taller, larger-brained Homo erectus from Asia, which spread to Europe and evolved into Nean derthals, which evolved into Homo sapiens. But what lay between the australopiths and H. erectus, the first known human? BETTING ON AFRICA Until the 1960s, H. erectus had been found only in Asia. But when primitive stone-chop LIBRARY PICTURE EVANS MUSEUM/MARY HISTORY NATURAL ping tools were uncovered at Olduvai Gorge in Tanzania, Leakey became convinced that this is where he would find the earliest stone- tool makers, who he assumed would belong to our genus. Maybe, like the australopiths, our human ancestors also originated in Africa. In 1931, Leakey began intensive prospect ing and excavation at Olduvai Gorge, 33 years before he announced the new human species. Now tourists travel to Olduvai on paved roads in air-conditioned buses; in the 1930s in the rainy season, the journey from Nairobi could take weeks. The ravines at Olduvai offered unparalleled access to ancient strata, but field work was no picnic in the park. Water was often scarce. Leakey and his team had to learn to share Olduvai with all of the wild animals that lived there, lions included. -
Hands-On Human Evolution: a Laboratory Based Approach
Hands-on Human Evolution: A Laboratory Based Approach Developed by Margarita Hernandez Center for Precollegiate Education and Training Author: Margarita Hernandez Curriculum Team: Julie Bokor, Sven Engling A huge thank you to….. Contents: 4. Author’s note 5. Introduction 6. Tips about the curriculum 8. Lesson Summaries 9. Lesson Sequencing Guide 10. Vocabulary 11. Next Generation Sunshine State Standards- Science 12. Background information 13. Lessons 122. Resources 123. Content Assessment 129. Content Area Expert Evaluation 131. Teacher Feedback Form 134. Student Feedback Form Lesson 1: Hominid Evolution Lab 19. Lesson 1 . Student Lab Pages . Student Lab Key . Human Evolution Phylogeny . Lab Station Numbers . Skeletal Pictures Lesson 2: Chromosomal Comparison Lab 48. Lesson 2 . Student Activity Pages . Student Lab Key Lesson 3: Naledi Jigsaw 77. Lesson 3 Author’s note Introduction Page The validity and importance of the theory of biological evolution runs strong throughout the topic of biology. Evolution serves as a foundation to many biological concepts by tying together the different tenants of biology, like ecology, anatomy, genetics, zoology, and taxonomy. It is for this reason that evolution plays a prominent role in the state and national standards and deserves thorough coverage in a classroom. A prime example of evolution can be seen in our own ancestral history, and this unit provides students with an excellent opportunity to consider the multiple lines of evidence that support hominid evolution. By allowing students the chance to uncover the supporting evidence for evolution themselves, they discover the ways the theory of evolution is supported by multiple sources. It is our hope that the opportunity to handle our ancestors’ bone casts and examine real molecular data, in an inquiry based environment, will pique the interest of students, ultimately leading them to conclude that the evidence they have gathered thoroughly supports the theory of evolution. -
Neither Chimpanzee Nor Human, Ardipithecus Reveals the Surprising Ancestry of Both Tim D
SPECIAL FEATURE: PERSPECTIVE PERSPECTIVE SPECIAL FEATURE: Neither chimpanzee nor human, Ardipithecus reveals the surprising ancestry of both Tim D. Whitea,1, C. Owen Lovejoyb, Berhane Asfawc, Joshua P. Carlsona, and Gen Suwad,1 aDepartment of Integrative Biology, Human Evolution Research Center, University of California, Berkeley, CA 94720; bDepartment of Anthropology, School of Biomedical Sciences, Kent State University, Kent, OH 44242–0001; cRift Valley Research Service, Addis Ababa, Ethiopia; and dThe University Museum, The University of Tokyo, Hongo, Bunkyo-ku Tokyo 113-0033, Japan Edited by Neil H. Shubin, University of Chicago, Chicago, IL, and approved September 10, 2014 (received for review April 25, 2014) Australopithecus fossils were regularly interpreted during the late 20th century in a framework that used living African apes, especially chimpanzees, as proxies for the immediate ancestors of the human clade. Such projection is now largely nullified by the discovery of Ardipithecus. In the context of accumulating evidence from genetics, developmental biology, anatomy, ecology, biogeography, and geology, Ardipithecus alters perspectives on how our earliest hominid ancestors—and our closest living relatives—evolved. human evolution | Australopithecus | hominid | Ethiopia “...the stock whence two or more species have chimpanzees, can serve as adequate repre- (5). Indeed, a widely used textbook still pro- sprung, need in no respect be intermediate sentations of the ancestral past. claims that, “Overall, Au. afarensis seems very between those species.” much like a missing link between the living Background T. H. Huxley, 1860 (1) Africanapesandlaterhomininsinitsdental, ’ Darwin s human evolution scenario attemp- cranial, and skeletal morphology” (6). Charles Darwin famously suggested that ted to explain hominid tool use, bipedality, Australopithecus can no longer be legiti- Africa was humanity’s most probable birth enlarged brains, and reduced canine teeth (2). -
'Lucy' Fossil Found
Published online 20 September 2006 | Nature | doi:10.1038/news060918-5 News Little 'Lucy' fossil found Toddler hominin has arms for swinging and legs for walking. Rex Dalton The 3.3-million-year-old bones of a female toddler from Ethiopia are telling scientists a story about the route human ancestors took from the trees to the ground. In today's issue of Nature, an Ethiopian-led international team reports the discovery of a juvenile skeleton of the species commonly known as 'Lucy', or Australopithecus afarensis.1,2 The researchers have named her Selam, after an Ethiopian word for 'peace'. The specimen, which is the oldest and most complete juvenile of a human relative ever found, has features that stand as striking examples of part-way evolution between primitive apes and modern humans. Although many other samples of A. afarensis have been found before, this is the first one reported to come complete with a whole shoulder-blade bone (scapula). In modern humans the scapula has a ridge running horizontally across the top of the bone; in apes the scapula's ridge reaches further down the Little Salem is the most back, where it can help to throw more muscle into arm action, as would be needed to swing from trees. ancient toddler ever found. In the young A. afarensis, the scapula looks to be part-way between. Zeresenay Alemseged and Copyright Authority for Research and Conservation "The animal was losing its capacity to be arboreal — heading right toward being human," says of Cultrual Heritages anthropologist Owen Lovejoy of Kent State University in Ohio. -
Mechanics of Bipedalism: an Exploration of Skeletal Morphology and Force Plate Anaylsis Erin Forse May 04, 2007 a Senior Thesis
MECHANICS OF BIPEDALISM: AN EXPLORATION OF SKELETAL MORPHOLOGY AND FORCE PLATE ANAYLSIS ERIN FORSE MAY 04, 2007 A SENIOR THESIS SUBMITTED IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF BACHELOR OF ARTS IN ARCHAEOLOGICAL STUDIES UNIVERSITY OF WISCONSIN- LA CROSSE Abstract There are several theories on how humans learned to walk, and while these all address the adaptations needed for walking, none adequately describes how our early ancestors developed the mechanism to walk. Our earliest recognizable relatives, the australopithecines, have several variations on a theme: walking upright. There are varied changes as australopithecines approach the genus Homo. These changes occurred in the spine, legs, pelvis, and feet, and changes are also in the cranium, arms and hands, but these are features that may have occurred simultaneously with bipedalism. Several analyses of Australopithecus afarensis, specifically specimen A.L. 288-1 ("Lucy"), have shown that the skeletal changes are intermediate between apes and humans. Force plate analyses are used to determine if the gait pattern of humans resembles that of apes, and if it is a likely development pattern. The results of both these analyses will give insight into how modern humans developed bipedalism. Introduction Bipedalism is classified as movement of the post-cranial body in a vertical position, with the lower limbs shifting as an inverted pendulum, progressing forward. Simply, it is upright walking. Several theories have addressed why bipedalism evolved in hominids, with some unlikely ideas taking hold throughout the history of the issue. Other theories are more likely, but all lack the same characteristic: answering how bipedalism developed. -
Human Evolution: a Paleoanthropological Perspective - F.H
PHYSICAL (BIOLOGICAL) ANTHROPOLOGY - Human Evolution: A Paleoanthropological Perspective - F.H. Smith HUMAN EVOLUTION: A PALEOANTHROPOLOGICAL PERSPECTIVE F.H. Smith Department of Anthropology, Loyola University Chicago, USA Keywords: Human evolution, Miocene apes, Sahelanthropus, australopithecines, Australopithecus afarensis, cladogenesis, robust australopithecines, early Homo, Homo erectus, Homo heidelbergensis, Australopithecus africanus/Australopithecus garhi, mitochondrial DNA, homology, Neandertals, modern human origins, African Transitional Group. Contents 1. Introduction 2. Reconstructing Biological History: The Relationship of Humans and Apes 3. The Human Fossil Record: Basal Hominins 4. The Earliest Definite Hominins: The Australopithecines 5. Early Australopithecines as Primitive Humans 6. The Australopithecine Radiation 7. Origin and Evolution of the Genus Homo 8. Explaining Early Hominin Evolution: Controversy and the Documentation- Explanation Controversy 9. Early Homo erectus in East Africa and the Initial Radiation of Homo 10. After Homo erectus: The Middle Range of the Evolution of the Genus Homo 11. Neandertals and Late Archaics from Africa and Asia: The Hominin World before Modernity 12. The Origin of Modern Humans 13. Closing Perspective Glossary Bibliography Biographical Sketch Summary UNESCO – EOLSS The basic course of human biological history is well represented by the existing fossil record, although there is considerable debate on the details of that history. This review details both what is firmly understood (first echelon issues) and what is contentious concerning humanSAMPLE evolution. Most of the coCHAPTERSntention actually concerns the details (second echelon issues) of human evolution rather than the fundamental issues. For example, both anatomical and molecular evidence on living (extant) hominoids (apes and humans) suggests the close relationship of African great apes and humans (hominins). That relationship is demonstrated by the existing hominoid fossil record, including that of early hominins. -
Energetic Tradeoffs Control the Size Distribution of Aquatic Mammals William Gearty
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Faculty Publications in the Biological Sciences Papers in the Biological Sciences 4-17-2018 Energetic tradeoffs control the size distribution of aquatic mammals William Gearty Craig R. McClain Jonathan Payne Follow this and additional works at: https://digitalcommons.unl.edu/bioscifacpub Part of the Biology Commons, Evolution Commons, and the Terrestrial and Aquatic Ecology Commons This Article is brought to you for free and open access by the Papers in the Biological Sciences at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Faculty Publications in the Biological Sciences by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Energetic tradeoffs control the size distribution of aquatic mammals William Geartya,1, Craig R. McClainb, and Jonathan L. Paynea aDepartment of Geological Sciences, Stanford University, Stanford, CA 94305; and bLouisiana Universities Marine Consortium, Chauvin, LA 70344 Edited by Nicholas D. Pyenson, Smithsonian Institution, Washington, DC, and accepted by Editorial Board Member David Jablonski February 23, 2018 (received for review August 8, 2017) Four extant lineages of mammals have invaded and diversified in the entering the water will increase in average size, these theories differ in water: Sirenia, Cetacea, Pinnipedia, and Lutrinae. Most of these aquatic their predictions for how such a size change is achieved. More spe- clades are larger bodied, on average, than their closest land-dwelling cifically, they differ in their predictions both about the rate of evo- relatives, but the extent to which potential ecological, biomechanical, lution toward the new, larger average size as well as the variance of and physiological controls contributed to this pattern remains untested the aquatic size distribution relative to its terrestrial sister group (22). -
Paranthropus Boisei: Fifty Years of Evidence and Analysis Bernard A
Marshall University Marshall Digital Scholar Biological Sciences Faculty Research Biological Sciences Fall 11-28-2007 Paranthropus boisei: Fifty Years of Evidence and Analysis Bernard A. Wood George Washington University Paul J. Constantino Biological Sciences, [email protected] Follow this and additional works at: http://mds.marshall.edu/bio_sciences_faculty Part of the Biological and Physical Anthropology Commons Recommended Citation Wood B and Constantino P. Paranthropus boisei: Fifty years of evidence and analysis. Yearbook of Physical Anthropology 50:106-132. This Article is brought to you for free and open access by the Biological Sciences at Marshall Digital Scholar. It has been accepted for inclusion in Biological Sciences Faculty Research by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected], [email protected]. YEARBOOK OF PHYSICAL ANTHROPOLOGY 50:106–132 (2007) Paranthropus boisei: Fifty Years of Evidence and Analysis Bernard Wood* and Paul Constantino Center for the Advanced Study of Hominid Paleobiology, George Washington University, Washington, DC 20052 KEY WORDS Paranthropus; boisei; aethiopicus; human evolution; Africa ABSTRACT Paranthropus boisei is a hominin taxon ers can trace the evolution of metric and nonmetric var- with a distinctive cranial and dental morphology. Its iables across hundreds of thousands of years. This pa- hypodigm has been recovered from sites with good per is a detailed1 review of half a century’s worth of fos- stratigraphic and chronological control, and for some sil evidence and analysis of P. boi se i and traces how morphological regions, such as the mandible and the both its evolutionary history and our understanding of mandibular dentition, the samples are not only rela- its evolutionary history have evolved during the past tively well dated, but they are, by paleontological 50 years. -
Anthro Notes : National Museum of Natural History Bulletin for Teachers
LUCY,UPATREE? Also, they argue, at least some of the A. afarensis hominids, especially the smaller ones like Lucy, no doubt slept, Paleoanthropologists no longer ques- hid, and fed in trees enough of the tion that Lucy, a 3 1/2 foot hominid time so that we can recognize some female with a chimp- sized brain, walked arboreal features in their anatomy. on two legs in Ethiopia about 3.5 million years ago. Neither do they argue that Susman and Stern presented their the anatomy of Lucy's species, Australo - evidence and analysis in an extensive pithecus afarensis , is fully modern; all article in the Journal of Physical agree it is a "mosaic of human-like and Anthropology (March 1983) , and at an ape-like features." No one seriously exciting and often boisterous confer- disputes that bipedal ism was more impor- ence in April. The conference, held tant to their lifestyle than for any non- at the Institute of Human Origins in human primate, living or dead. However, Berkeley, was directed by Donald C. Lucy's discoverers, Donald C. Johanson Johanson, founder of the Institute. and Tim White, claim that the bipedalism There the different factions met to seen in A. afarensis differs insignifi- examine the bones and thrash out their cantly from that of modern humans. Other many different views about two con- scientists disagree. troversies: 1) When did bipedalism begin and to what extent was Lucy Recently two noted anatomists from bipedal? 2) Did A. afarensis make the State University of New York at Stony the footprints at Laetoli or did Brook, Jack Stern and Randall L. -
What Makes a Modern Human We Probably All Carry Genes from Archaic Species Such As Neanderthals
COMMENT NATURAL HISTORY Edward EARTH SCIENCE How rocks and MUSIC Philip Glass on Einstein EMPLOYMENT The skills gained Lear’s forgotten work life evolved together on our and the unpredictability of in PhD training make it on ornithology p.36 planet p.39 opera composition p.40 worth the money p.41 ILLUSTRATION BY CHRISTIAN DARKIN CHRISTIAN BY ILLUSTRATION What makes a modern human We probably all carry genes from archaic species such as Neanderthals. Chris Stringer explains why the DNA we have in common is more important than any differences. n many ways, what makes a modern we were trying to set up strict criteria, based non-modern (or, in palaeontological human is obvious. Compared with our on cranial measurements, to test whether terms, archaic). What I did not foresee evolutionary forebears, Homo sapiens is controversial fossils from Omo Kibish in was that some researchers who were not Icharacterized by a lightly built skeleton and Ethiopia were within the range of human impressed with our test would reverse it, several novel skull features. But attempts to skeletal variation today — anatomically applying it back onto the skeletal range of distinguish the traits of modern humans modern humans. all modern humans to claim that our diag- from those of our ancestors can be fraught Our results suggested that one skull nosis wrongly excluded some skulls of with problems. was modern, whereas the other was recent populations from being modern2. Decades ago, a colleague and I got into This, they suggested, implied that some difficulties over an attempt to define (or, as PEOPLING THE PLANET people today were more ‘modern’ than oth- I prefer, diagnose) modern humans using Interactive map of migrations: ers.