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III Joséphine+Th. Koster INULEAE The compositae of New Guinea III Joséphine+Th. Koster Rijksherbarium, Leiden 4. INULEAE*) 88 & Nat. Pfl. Fam. Cass., J. Phys. Chim. Hist. Nat. (1819) 193; Hoffmann, E. P., 4, 5 (1894) 172. Herbs. Leaves nearly always alternate, sometimes rosulate, mostly entire, sometimes dentate, rarely pinnatifid. Heads solitary or in inflorescences, homogamous or heteroga- herbaceous corolla of mous; phyllaries one- to many-seriate, or membranous; marginal anthers flowers filiform, dentate, or ligulate, of disc flowers tubular, (4- or) 5-dentate; achene sagittate and mostly caudate at the base; style two-armed; small, pappus setaceous, sometimes consisting of scales, or wanting; receptacle naked. KEY TO THE GENERA IN NEW GUINEA Leaves decurrent the heads clustered into 2 I a. along stem; small, densely glomerules b. Leaves not decurrent along the stem; heads solitary, few together, corymbose, paniculate, or clustered, but not in glomerules 3 Pterocaulon 2 a. Pappus setaceous I. 2 Sphaeranthus b. Pappus wanting . 3. Epaltes 3 a. Heads small, numerous, disc-shaped; pappus wanting Heads setaceous b. variously shaped; pappus 4 18— orange-yellow, afterwards glossy. 4 a. Headsjlarge; involucre 25 mm long; phyllaries yellow-brown, 4. Helichrysum than 18 coloured but not 5 b. Involucre much shorter mm; phyllaries variously yellow .... Phacellothrix 5 a. Heads solitary, homogamous; corolla tubular 5- several when then corolla of b. Heads solitary or together,mostly heterogamous, homogamous marginal flowers filiform 6 less whitish brown. 6 a. Whitish lanate herbs; phyllaries rigid, membranous,more or glossy, or light 7 membranous, not whitish or light brown 8 b. Herbs not whitish lanate; phyllaries herbaceous or Heads feminine flowers often numerous, sometimes 7 a. densely corymbose, shortly pedunculate; marginal wanting; disc flowers often masculine, sometimes bisexual, 1 — 50; exclusively masculine heads frequent, number of flowers and disc flowers in head variable in and the same species. marginal a very one 6. Anaphalis flowers disc flowers b. Heads clustered or solitary, sessile or subsessile; feminine marginal numerous; flowers masculine heads not occurring, number of and disc bisexual, 1—7; exclusively marginal in in and the flowers a head not much variable one same species 7. Gnaphalium 8 undivided 8. Pluchea a. Disc flowers few, masculine; style or shortly two-armed, ovary rudimentary . Blumea b. Disc flowers few ornumerous, bisexual; style arms fairly long, ovary normally developed 9. Bot. of tribes in accordance with this II in *) I in Nova Guinea, 24 (1966) 497—614 (numbering paper); Blumea 18 (1970) 137—145. 193 BLUMEA VOL. No. 194 XX, I, 1972 I. PTEROCAULON Sketch Bot. Austr. Elliot, S.-Carol. and Geo. 2 (1823) 323; Benth., Fl. 3 (1866) 522; Benth. & Hook, /, Gen. Pi. 2 (1873) 294; Clarke, Comp. Ind. (1876) 98; Hook. /, Fl. & Pfl. Br. Ind. 3 (1881) 275; Hoffmann, E. P., Nat. Fam. 4, 5 (1894) 178; Koster in Fl. — Backer & Bakh., Java 2 (1965) 391. Monenteles Labill., Sert. Austro-caled. 2 (1825) 42. Grey-tomentose herbs. Leaves alternate, decurrent along the stem. Heads small, densely clustered into globose, ellipsoid, or cylindrical glomerules, heterogamous; involucre ovoid or campanulate, phyllaries in few rows, narrow, inner gradually shorter, linear, rigidly scarious, deciduous together with flowers, outer much shorter. Marginal flowers corolla several, female; filiform, minutely 2- or 3-dentate or truncate; style-arms filiform, rather achene hairs long, exserted; small, oblong, 4- or 5-costate, pappus setaceous, white, ciliate. Disc often corolla fine, flowers few or one, bisexual, sterile; tubular, thin, 5-lobate; anthers and caudate the obtuse the auricles sagittate at base, at apex, very fine, connate; style-arms lanceolate, short; achene and pappus like those of marginal flowers; receptacle minute. Distribution: about and China 30 species in N., C., S. America, Burma, S. Malesia, Australia, New Caledonia. KEY TO THE SPECIES IN NEW GUINEA I a. Leaves obovate-oblong, minutely dentate; glomerules of heads in terminal spikes I. P. redolens Leaves of b. oblong, repand or subentire; glomerules heads solitary, terminal, sessile or on winged peduncles 2. P. sphacelatum I. Pterocaulon redolens (Forst. f. ex Willd.) F.-Vill., Noviss. App. Blanco, Fl. Filip., 3rd ed., 4, 3 (1880) 116; Mattfeld, Bot. Jahrb. 62 (1929) 426; Merr. & Chun, Sunyatsenia 2 Craib Fl. — redolens (1935) 329; in Kerr, Siam. 2 (1936) 268. Conyza Forst. / ex Willd., Sp. Pi. — Monenteles Austro-caled. t. Benth. 3 (1803) 1951. spicatus Labill., Sert. 2 (1825) 43, 43; — Ind. Fl. Austr. 3 (1866) 523. Pterocaulon cylindrostachyum Clarke, Comp. (1876) 98; Fl. Ind. Weeds Pi. f. Hook./, Br. 3 (1881) 275; Bailey, Pois. Qld (1906) 135; Gagnepain, Fl. Gen. I.-C. 3 (1924) 550, f. 57:1—7. — Pterocaulon spicatum (Labill.) Domin, Bibl. Bot., Heft 89 (1929) 1218. often much branched the Erect herb, 30—100 cm tall, in upper part, stem sparsely in greyish lanate-tomentose, the lower part glabrescent, 2—6 mm thick, wing entire, greyish lanate-tomentose, internodes 1—2 cm long. Leaves obovate-oblong, narrowed the rounded the to base, or nearly so at apex, minutely dentate, chartaceous, densely broad. greyish lanate-tomentose below, thinly so above, 1—5.5 cm long, 0.5—2 cm of heads Glomerules small, globose or ellipsoid-oblong, sessile, in terminal 2—7 cm long heads involucre spikes; ellipsoid-oblong; ovoid, 5 mm long, phyllaries 4-seriate, inner linear-lanceolate, acuminate and acute, rigid, stramineous, pale brownish, shining, glabrous, outer narrowly spathulate, obtuse or rounded, densely dingy crisp-lanate. Marginal flowers 20—25; corolla 2- or 3-dentate, 3 mm long; achene dark brown, 0.75 mm corolla long, sparsely shortly pilose, pappus 2.5 mm long. Disc flower one; 3 mm long, lobes deltoid-ovate, at the achene glandular upper part; 0.5 mm long, empty, pappus 2.5 mm long. Distribution: Burma, Thailand, Laos, Vietnam, Malay Peninsula, Philippines, New Guinea, Australia, New Caledonia, New Hebrides. Josephine Th. Koster: The Compositae of New Guittea 111 195 — Dist.: NEW GUINEA. East. Milne Bay Dist.: Uga, 1949, Cruttwell 173 (K). Central Kapa to Kaili, Port Goldie E. Tovobada 1889, MacGregor s.n. (MEL); Moresby, s.d., s.n. (MEL); footslopes Hills, 12 m. Port Kairo & Streimann N. of Port Moresby, 1965, Heyligers 1229 (L); Moresby Subdist., Tupuseleia, 1967, —MorobeDist.: Clemens NGF30803 (L); Hisiu, 1935, Can 11451 (A, BM,L, NY). Kajabit, 1939. 10519(A); NGF Markham valley, Erap near Lea, 1958, Henly 9841 (BO, BRI, L, LAE), 1959, 10684 (LAE). Collectors' annotations: aromatic, maladorous; leaves dull green above, corolla fait white red and white. grey green below; pink, white, tinged pale mauve, or savannah Ecological notes: roadside weed; in open grassland; in open land; bare of with of in woodland. Alti- in strand vegetation; on patch ground plenty humus; tude: 0—300 m. Pi. 2. Pterocaulon sphacelatum (Labill.) F. v. M., Syst. Census Austr. 1 (1882) 79; Fl. Austr. Banks and Sol. Bailey, Queensl. 3 (1900) 824; Britten, 111. Pi. 2 (1901) 49 t. 156; Backer Domin, Bibl. Bot. Heft 89 (1929) 1218; Koster, Fedde Repert. 34 (1933) 14; in & Fl. — Monenteles Sert. Austro-caled. 2 Bakh., Java 2 (1965) 392. sphacelous Labill., (1825) t. Fl. Benth. Hook. Gen. Pi. 2 43, 44; Benth., Austr. 3 (1866) 523; & /, (1873) 295. Herbs stem 2—6 with entire 25—60 cm tall, upper part branched, mm thick, together wing densely lanate-tomentose, hairs long, crispy, hyaline, inflated at the base; internodes rounded and apiculate at the repand or subentire, 1—3 cm long. Leaves oblong, apex, chartaceous, thinly greyish more or less crispy-tomentose above, densely so below, smaller. Glomerules of heads globose 1—4.5 cm long, 0.5—2 cm broad, higher ones or sessile ellipsoid, I—1.5 cm wide, terminal, or on 0.5 —1.5 cm long winged peduncles; inner linear- heads ellipsoid-oblong, involucre ovoid, 4 mm long, phyllaries 4-seriate, whitish lanceolate, acuminate and acute, rigid, stramineous, shining, glabrous, outer obtuse narrowly spathulate, or rounded, densely white crispy-lanate. Marginal flowers corolla achene 10—20; finely 2- or 3-dentate, purplish, 3 mm long; pilose, 1 mm long, corolla mm flower mm long, lobes deltoid-ovate, glandular pappus 3 long. Disc one; 3.5 the achene mm at upper part; 0.75 mm long, empty, pappus 3 long. Distribution: SE. Malesia (Kangean I., Buton I., Lesser Sunda Is, New Guinea), Australia, New Caledonia. NEW GUINEA. East. Milne Bay Dist.: Milne Bay, 1889, MacGregor s.n. (MEL). — Central Dist.: ofFRed Kairuhu Subdist., Rogers Airstrip, 8 m. W. ofKanosia Plantation, 1962, Darbyshire 647 (L); island Scar Chalmers s.n. (MEL). Point, 1849, MacGillavry 473 (K); Fisherman L, 1880, s' herb to corolla Collector annotations: erect up 90 cm tall; faintly purple. Ecological notes: open savannah. Altitude: 9 m. 2. SPHAERANTHUS Gen. L., Sp. Pi. (1753) 927; Lessing, Syn. Gen. Comp. (1832) 201; Benth. & Hook./, Pi. Ind. Fl. Ind. (1881) Hoffmann, 2 (1873) 294; Clarke, Comp. (1876) 96; Hook./., Br. 3 274; E. & Pfl. Kew Bull. Hook. P., Nat. Fam. 4, 5 (1894) 178; Robijns, (1924) 177; Ross-Craig, 6 & Ic. Pl. V, (1954) 1; Koster in Backer Bakh., Fl. Java 2 (1965) 391. Herbs. Leaves alternate, nearly always decurrent along the stem. Heads minute, densely clustered ovoid in terminal or axillary, nearly always pedunculate, globular or glomerules, heterogamous; involucre narrowly campanulate, phyllaries about equal in length. less Marginal flowers many, female; corolla tubular, minutely 3-dentate, more or swollen, achene subcartilagineous; style shortly bifid; oblong, pappus wanting. Disc flowers one or anthers few, bisexual or male; corolla tubular-campanulate, 5-lobed, subcartilaginous; No. BLUMEA VOL. XX, I, 1972 196 with acute basal obtuse at the style undivided or sagittate or subcaudate, auricles, apex; minutely bifid, papillose; achene oblong, pappus wanting; receptacle small, naked, variable in shape. Distribution: tropical Asia, Africa, Australia. 37 species in Iraq, Iran, Br.
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