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GASTROPOD CARE SOP# = Moll3 PURPOSE: to Describe Methods Of
GASTROPOD CARE SOP# = Moll3 PURPOSE: To describe methods of care for gastropods. POLICY: To provide optimum care for all animals. RESPONSIBILITY: Collector and user of the animals. If these are not the same person, the user takes over responsibility of the animals as soon as the animals have arrived on station. IDENTIFICATION: Common Name Scientific Name Identifying Characteristics Blue topsnail Calliostoma - Whorls are sculptured spirally with alternating ligatum light ridges and pinkish-brown furrows - Height reaches a little more than 2cm and is a bit greater than the width -There is no opening in the base of the shell near its center (umbilicus) Purple-ringed Calliostoma - Alternating whorls of orange and fluorescent topsnail annulatum purple make for spectacular colouration - The apex is sharply pointed - The foot is bright orange - They are often found amongst hydroids which are one of their food sources - These snails are up to 4cm across Leafy Ceratostoma - Spiral ridges on shell hornmouth foliatum - Three lengthwise frills - Frills vary, but are generally discontinuous and look unfinished - They reach a length of about 8cm Rough keyhole Diodora aspera - Likely to be found in the intertidal region limpet - Have a single apical aperture to allow water to exit - Reach a length of about 5 cm Limpet Lottia sp - This genus covers quite a few species of limpets, at least 4 of them are commonly found near BMSC - Different Lottia species vary greatly in appearance - See Eugene N. Kozloff’s book, “Seashore Life of the Northern Pacific Coast” for in depth descriptions of individual species Limpet Tectura sp. - This genus covers quite a few species of limpets, at least 6 of them are commonly found near BMSC - Different Tectura species vary greatly in appearance - See Eugene N. -
Global Seagrass Distribution and Diversity: a Bioregional Model ⁎ F
Journal of Experimental Marine Biology and Ecology 350 (2007) 3–20 www.elsevier.com/locate/jembe Global seagrass distribution and diversity: A bioregional model ⁎ F. Short a, , T. Carruthers b, W. Dennison b, M. Waycott c a Department of Natural Resources, University of New Hampshire, Jackson Estuarine Laboratory, Durham, NH 03824, USA b Integration and Application Network, University of Maryland Center for Environmental Science, Cambridge, MD 21613, USA c School of Marine and Tropical Biology, James Cook University, Townsville, 4811 Queensland, Australia Received 1 February 2007; received in revised form 31 May 2007; accepted 4 June 2007 Abstract Seagrasses, marine flowering plants, are widely distributed along temperate and tropical coastlines of the world. Seagrasses have key ecological roles in coastal ecosystems and can form extensive meadows supporting high biodiversity. The global species diversity of seagrasses is low (b60 species), but species can have ranges that extend for thousands of kilometers of coastline. Seagrass bioregions are defined here, based on species assemblages, species distributional ranges, and tropical and temperate influences. Six global bioregions are presented: four temperate and two tropical. The temperate bioregions include the Temperate North Atlantic, the Temperate North Pacific, the Mediterranean, and the Temperate Southern Oceans. The Temperate North Atlantic has low seagrass diversity, the major species being Zostera marina, typically occurring in estuaries and lagoons. The Temperate North Pacific has high seagrass diversity with Zostera spp. in estuaries and lagoons as well as Phyllospadix spp. in the surf zone. The Mediterranean region has clear water with vast meadows of moderate diversity of both temperate and tropical seagrasses, dominated by deep-growing Posidonia oceanica. -
BIO 313 ANIMAL ECOLOGY Corrected
NATIONAL OPEN UNIVERSITY OF NIGERIA SCHOOL OF SCIENCE AND TECHNOLOGY COURSE CODE: BIO 314 COURSE TITLE: ANIMAL ECOLOGY 1 BIO 314: ANIMAL ECOLOGY Team Writers: Dr O.A. Olajuyigbe Department of Biology Adeyemi Colledge of Education, P.M.B. 520, Ondo, Ondo State Nigeria. Miss F.C. Olakolu Nigerian Institute for Oceanography and Marine Research, No 3 Wilmot Point Road, Bar-beach Bus-stop, Victoria Island, Lagos, Nigeria. Mrs H.O. Omogoriola Nigerian Institute for Oceanography and Marine Research, No 3 Wilmot Point Road, Bar-beach Bus-stop, Victoria Island, Lagos, Nigeria. EDITOR: Mrs Ajetomobi School of Agricultural Sciences Lagos State Polytechnic Ikorodu, Lagos 2 BIO 313 COURSE GUIDE Introduction Animal Ecology (313) is a first semester course. It is a two credit unit elective course which all students offering Bachelor of Science (BSc) in Biology can take. Animal ecology is an important area of study for scientists. It is the study of animals and how they related to each other as well as their environment. It can also be defined as the scientific study of interactions that determine the distribution and abundance of organisms. Since this is a course in animal ecology, we will focus on animals, which we will define fairly generally as organisms that can move around during some stages of their life and that must feed on other organisms or their products. There are various forms of animal ecology. This includes: • Behavioral ecology, the study of the behavior of the animals with relation to their environment and others • Population ecology, the study of the effects on the population of these animals • Marine ecology is the scientific study of marine-life habitat, populations, and interactions among organisms and the surrounding environment including their abiotic (non-living physical and chemical factors that affect the ability of organisms to survive and reproduce) and biotic factors (living things or the materials that directly or indirectly affect an organism in its environment). -
The Biology of Seashores - Image Bank Guide All Images and Text ©2006 Biomedia ASSOCIATES
The Biology of Seashores - Image Bank Guide All Images And Text ©2006 BioMEDIA ASSOCIATES Shore Types Low tide, sandy beach, clam diggers. Knowing the Low tide, rocky shore, sandstone shelves ,The time and extent of low tides is important for people amount of beach exposed at low tide depends both on who collect intertidal organisms for food. the level the tide will reach, and on the gradient of the beach. Low tide, Salt Point, CA, mixed sandstone and hard Low tide, granite boulders, The geology of intertidal rock boulders. A rocky beach at low tide. Rocks in the areas varies widely. Here, vertical faces of exposure background are about 15 ft. (4 meters) high. are mixed with gentle slopes, providing much variation in rocky intertidal habitat. Split frame, showing low tide and high tide from same view, Salt Point, California. Identical views Low tide, muddy bay, Bodega Bay, California. of a rocky intertidal area at a moderate low tide (left) Bays protected from winds, currents, and waves tend and moderate high tide (right). Tidal variation between to be shallow and muddy as sediments from rivers these two times was about 9 feet (2.7 m). accumulate in the basin. The receding tide leaves mudflats. High tide, Salt Point, mixed sandstone and hard rock boulders. Same beach as previous two slides, Low tide, muddy bay. In some bays, low tides expose note the absence of exposed algae on the rocks. vast areas of mudflats. The sea may recede several kilometers from the shoreline of high tide Tides Low tide, sandy beach. -
The Behavioral Ecology and Territoriality of the Owl Limpet, Lottia Gigantea
THE BEHAVIORAL ECOLOGY AND TERRITORIALITY OF THE OWL LIMPET, LOTTIA GIGANTEA by STEPHANIE LYNN SCHROEDER A DISSERTATION Presented to the Department of Biology and the Graduate School of the University of Oregon in partial fulfillment of the requirements for the degree of Doctor of Philosophy March 2011 DISSERTATION APPROVAL PAGE Student: Stephanie Lynn Schroeder Title: The Behavioral Ecology and Territoriality of the Owl Limpet, Lottia gigantea This dissertation has been accepted and approved in partial fulfillment of the requirements for the Doctor of Philosophy degree in the Department of Biology by: Barbara (“Bitty”) Roy Chairperson Alan Shanks Advisor Craig Young Member Mark Hixon Member Frances White Outside Member and Richard Linton Vice President for Research and Graduate Studies/Dean of the Graduate School Original approval signatures are on file with the University of Oregon Graduate School. Degree awarded March 2011 ii © 2011 Stephanie Lynn Schroeder iii DISSERTATION ABSTRACT Stephanie Lynn Schroeder Doctor of Philosophy Department of Biology March 2011 Title: The Behavioral Ecology and Territoriality of the Owl Limpet, Lottia gigantea Approved: _______________________________________________ Dr. Alan Shanks Territoriality, defined as an animal or group of animals defending an area, is thought to have evolved as a means to acquire limited resources such as food, nest sites, or mates. Most studies of territoriality have focused on vertebrates, which have large territories and even larger home ranges. While there are many models used to examine territories and territorial interactions, testing the models is limited by the logistics of working with the typical model organisms, vertebrates, and their large territories. An ideal organism for the experimental examination of territoriality would exhibit clear territorial behavior in the field and laboratory, would be easy to maintain in the laboratory, defend a small territory, and have movements and social interactions that were easily followed. -
OREGON ESTUARINE INVERTEBRATES an Illustrated Guide to the Common and Important Invertebrate Animals
OREGON ESTUARINE INVERTEBRATES An Illustrated Guide to the Common and Important Invertebrate Animals By Paul Rudy, Jr. Lynn Hay Rudy Oregon Institute of Marine Biology University of Oregon Charleston, Oregon 97420 Contract No. 79-111 Project Officer Jay F. Watson U.S. Fish and Wildlife Service 500 N.E. Multnomah Street Portland, Oregon 97232 Performed for National Coastal Ecosystems Team Office of Biological Services Fish and Wildlife Service U.S. Department of Interior Washington, D.C. 20240 Table of Contents Introduction CNIDARIA Hydrozoa Aequorea aequorea ................................................................ 6 Obelia longissima .................................................................. 8 Polyorchis penicillatus 10 Tubularia crocea ................................................................. 12 Anthozoa Anthopleura artemisia ................................. 14 Anthopleura elegantissima .................................................. 16 Haliplanella luciae .................................................................. 18 Nematostella vectensis ......................................................... 20 Metridium senile .................................................................... 22 NEMERTEA Amphiporus imparispinosus ................................................ 24 Carinoma mutabilis ................................................................ 26 Cerebratulus californiensis .................................................. 28 Lineus ruber ......................................................................... -
The Diet and Predator-Prey Relationships of the Sea Star Pycnopodia Helianthoides (Brandt) from a Central California Kelp Forest
THE DIET AND PREDATOR-PREY RELATIONSHIPS OF THE SEA STAR PYCNOPODIA HELIANTHOIDES (BRANDT) FROM A CENTRAL CALIFORNIA KELP FOREST A Thesis Presented to The Faculty of Moss Landing Marine Laboratories San Jose State University In Partial Fulfillment of the Requirements for the Degree Master of Arts by Timothy John Herrlinger December 1983 TABLE OF CONTENTS Acknowledgments iv Abstract vi List of Tables viii List of Figures ix INTRODUCTION 1 MATERIALS AND METHODS Site Description 4 Diet 5 Prey Densities and Defensive Responses 8 Prey-Size Selection 9 Prey Handling Times 9 Prey Adhesion 9 Tethering of Calliostoma ligatum 10 Microhabitat Distribution of Prey 12 OBSERVATIONS AND RESULTS Diet 14 Prey Densities 16 Prey Defensive Responses 17 Prey-Size Selection 18 Prey Handling Times 18 Prey Adhesion 19 Tethering of Calliostoma ligatum 19 Microhabitat Distribution of Prey 20 DISCUSSION Diet 21 Prey Densities 24 Prey Defensive Responses 25 Prey-Size Selection 27 Prey Handling Times 27 Prey Adhesion 28 Tethering of Calliostoma ligatum and Prey Refugia 29 Microhabitat Distribution of Prey 32 Chemoreception vs. a Chemotactile Response 36 Foraging Strategy 38 LITERATURE CITED 41 TABLES 48 FIGURES 56 iii ACKNOWLEDGMENTS My span at Moss Landing Marine Laboratories has been a wonderful experience. So many people have contributed in one way or another to the outcome. My diving buddies perse- vered through a lot and I cherish our camaraderie: Todd Anderson, Joel Thompson, Allan Fukuyama, Val Breda, John Heine, Mike Denega, Bruce Welden, Becky Herrlinger, Al Solonsky, Ellen Faurot, Gilbert Van Dykhuizen, Ralph Larson, Guy Hoelzer, Mickey Singer, and Jerry Kashiwada. Kevin Lohman and Richard Reaves spent many hours repairing com puter programs for me. -
Genetic Evidence for the Cryptic Species Pair, Lottia Digitalis and Lottia Austrodigitalis and Microhabitat Partitioning in Sympatry
Mar Biol (2007) 152:1–13 DOI 10.1007/s00227-007-0621-4 RESEARCH ARTICLE Genetic evidence for the cryptic species pair, Lottia digitalis and Lottia austrodigitalis and microhabitat partitioning in sympatry Lisa T. Crummett · Douglas J. Eernisse Received: 8 June 2005 / Accepted: 3 January 2007 / Published online: 14 April 2007 © Springer-Verlag 2007 Abstract It has been proposed that the common West rey Peninsula, CA to near Pigeon Point, CA, where L. digi- Coast limpet, Lottia digitalis, is actually the northern coun- talis previously dominated. terpart of a cryptic species duo including, Lottia austrodigi- talis. Allele frequency diVerences between southern and northern populations at two polymorphic enzyme loci pro- Introduction vided the basis for this claim. Due to lack of further evi- dence, L. austrodigitalis is still largely unrecognized in the Sibling species are deWned as sister species that are impos- literature. Seven additional enzyme loci were examined sible or extremely diYcult to distinguish based on morpho- from populations in proposed zones of allopatry and symp- logical characters alone (Mayr and Ashlock 1991). Marine atry to determine the existence of L. austrodigitalis as a sibling species are ubiquitous, found from the poles to the sibling species to L. digitalis. SigniWcant allele frequency tropics, in most known habitats, and at depths ranging from diVerences were found at Wve enzyme loci between popula- intertidal to abyssal (Knowlton 1993). We will refer to spe- tions in Laguna Beach, southern California, and Bodega cies that are indistinguishable morphologically, whether or Bay, northern California; strongly supporting the existence not they are sister species, as “cryptic species” and will of separate species. -
Intertidal Organisms of Point Reyes National Seashore
Intertidal Organisms of Point Reyes National Seashore PORIFERA: sea sponges. CRUSTACEANS: barnacles, shrimp, crabs, and allies. CNIDERIANS: sea anemones and allies. MOLLUSKS : abalones, limpets, snails, BRYOZOANS: moss animals. clams, nudibranchs, chitons, and octopi. ECHINODERMS: sea stars, sea cucumbers, MARINE WORMS: flatworms, ribbon brittle stars, sea urchins. worms, peanut worms, segmented worms. UROCHORDATES: tunicates. Genus/Species Common Name Porifera Prosuberites spp. Cork sponge Leucosolenia eleanor Calcareous sponge Leucilla nuttingi Little white sponge Aplysilla glacialis Karatose sponge Lissodendoryx spp. Skunk sponge Ophlitaspongia pennata Red star sponge Haliclona spp. Purple haliclona Leuconia heathi Sharp-spined leuconia Cliona celata Yellow-boring sponge Plocarnia karykina Red encrusting sponge Hymeniacidon spp. Yellow nipple sponge Polymastia pachymastia Polymastia Cniderians Tubularia marina Tubularia hydroid Garveia annulata Orange-colored hydroid Ovelia spp. Obelia Sertularia spp. Sertularia Abientinaria greenii Green's bushy hydroid Aglaophenia struthionides Giant ostrich-plume hydroid Aglaophenia latirostris Dainty ostrich-plume hydroid Plumularia spp. Plumularia Pleurobrachia bachei Cat's eye Polyorchis spp. Bell-shaped jellyfish Chrysaora melanaster Striped jellyfish Velella velella By-the-wind-sailor Aurelia auria Moon jelly Epiactus prolifera Proliferating anemone Anthopleura xanthogrammica Giant green anemone Anthopleura artemissia Aggregated anemone Anthopleura elegantissima Burrowing anemone Tealia lofotensis -
The Global Distribution and Status of Seagrass Ecosystems
The global distribution and status of seagrass ecosystems ^^ ^^^H Discussion paper prepared for tlie UNEP-WCWIC Global Seagrass Workshop St Pete's Beach, Florida, 9 November, 2001 Prepared by: Mark D. Spalding, Michelle L. Taylor, Sergio Martins, Edmund P. Green, and Mary Edwards WA.. WORLD CONSERVATION MONITORING CENTRE Digitized by tine Internet Archive in 2010 witii funding from UNEP-WCIVIC, Cambridge Iittp://www.archive.org/details/globaldistributi01spal The global distribution and status of seagrass ecosystems Discussion paper prepared for tlie UNEP-WCIVIC Global Seagrass Workshop St Pete's Beach, Florida, 9 November, 2001 Prepared by: Mark D. Spalding, Michelle L. Taylor, Sergio Martins, Edmund P. Green, and Mary Edwards With assistance from: Mark Taylor and Corinna Ravilious Table of Contents Introduction to the workshop 2 The global distribution and status of seagrass ecosystems 3 Introduction 3 Definitions 3 The diversity of seagrasses 3 Species distribution 4 Associated Species 6 Productivity and biomass 7 The distribution and area of seagrass habitat 8 The value of seagrasses 13 Threats to seagrasses 13 Management Interventions 14 Bibliography; 16 29 Annex 1 : Seagrass Species Lists by Country Annex 2 - Species distribution maps 34 Annex 3 - Seagrass distribution maps 68 74 Annex 4 -Full list of MPAs by country ; /4^ ] UNEP WCMC Introduction to the workshop The Global Seagrass Workshop of 9 November 2001 has been set up with the expressed aim to develop a global synthesis on the distribution and status of seagrasses world-wide. Approximately 20 seagrass experts from 14 counu-ies, representing all of the major seagrass regions of the world have been invited to share their knowledge and expertise. -
1 Phylogenetic Regionalization of Marine Plants Reveals Close Evolutionary Affinities Among Disjunct Temperate Assemblages Barna
Phylogenetic regionalization of marine plants reveals close evolutionary affinities among disjunct temperate assemblages Barnabas H. Darua,b,*, Ben G. Holtc, Jean-Philippe Lessardd,e, Kowiyou Yessoufouf and T. Jonathan Daviesg,h aDepartment of Organismic and Evolutionary Biology and Harvard University Herbaria, Harvard University, Cambridge, MA 02138, USA bDepartment of Plant Science, University of Pretoria, Private Bag X20, Hatfield 0028, Pretoria, South Africa cDepartment of Life Sciences, Imperial College London, Silwood Park Campus, Ascot SL5 7PY, United Kingdom dQuebec Centre for Biodiversity Science, Department of Biology, McGill University, Montreal, QC H3A 0G4, Canada eDepartment of Biology, Concordia University, Montreal, QC, H4B 1R6, Canada; fDepartment of Environmental Sciences, University of South Africa, Florida campus, Florida 1710, South Africa gDepartment of Biology, McGill University, Montreal, QC H3A 0G4, Canada hAfrican Centre for DNA Barcoding, University of Johannesburg, PO Box 524, Auckland Park, Johannesburg 2006, South Africa *Corresponding author Email: [email protected] (B.H. Daru) Running head: Phylogenetic regionalization of seagrasses 1 Abstract While our knowledge of species distributions and diversity in the terrestrial biosphere has increased sharply over the last decades, we lack equivalent knowledge of the marine world. Here, we use the phylogenetic tree of seagrasses along with their global distributions and a metric of phylogenetic beta diversity to generate a phylogenetically-based delimitation of marine phytoregions (phyloregions). We then evaluate their evolutionary affinities and explore environmental correlates of phylogenetic turnover between them. We identified 11 phyloregions based on the clustering of phylogenetic beta diversity values. Most phyloregions can be classified as either temperate or tropical, and even geographically disjunct temperate regions can harbor closely related species assemblages. -
Evolutionary Consequences of Food Chain Length in Kelp Forest Communities (Biogeography/Coevolution/Herbivory/Phlorotannins/Predation) PETER D
Proc. Natl. Acad. Sci. USA Vol. 92, pp. 8145-8148, August 1995 Ecology Evolutionary consequences of food chain length in kelp forest communities (biogeography/coevolution/herbivory/phlorotannins/predation) PETER D. STEINBERG*, JAMES A. ESTEStt, AND FRANK C. WINTER§ *School of Biological Sciences, University of New South Wales, P.O. Box 1, Kensington, New South Wales, 2033, Australia; tNational Biological Service, A-316 Earth and Marine Sciences Building, University of California, Santa Cruz, CA 95064; and §University of Auckland, Leigh Marine Laboratory, P.O. Box 349, Warkworth, New Zealand Communicated by Robert T. Paine, University of Washington, Seattle, WA, May 12, 1995 ABSTRACT Kelp forests are strongly influenced by mac- consistently important structuring processes throughout the roinvertebrate grazing on fleshy macroalgae. In the North food web. Under these conditions, we would predict that Pacific Ocean, sea otter predation on macroinvertebrates top-level consumers are resource limited. Consequently, the substantially reduces the intensity of herbivory on macroal- next lower trophic level should be consumer limited, in turn gae. Temperate Australasia, in contrast, has no known pred- causing the level below that (if one exists) to again be resource ator of comparable influence. These ecological and biogeo- limited. Looking downward through the food web from this graphic patterns led us to predict that (i) the intensity of very generalized perspective, a pattern emerges of strongly herbivory should be greater in temperate Australasia than in interacting couplets of adjacent trophic levels. Given these the North Pacific Ocean; thus (ii) Australasian seaweeds have circumstances, the interactive coupling between plants and been under stronger selection to evolve chemical defenses and herbivores should be strong in even-numbered systems and (iii) Australasian herbivores have been more strongly selected weak in odd-numbered systems, a prediction recently substan- to tolerate these compounds.