It Is Time for a New Classification of Anoles (Squamata: Dactyloidae)
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Zootaxa 3477: 1–108 (2012) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA Copyright © 2012 · Magnolia Press Monograph ISSN 1175-5334 (online edition) urn:lsid:zoobank.org:pub:32126D3A-04BC-4AAC-89C5-F407AE28021C ZOOTAXA 3477 It is time for a new classification of anoles (Squamata: Dactyloidae) KIRSTEN E. NICHOLSON1, BRIAN I. CROTHER2, CRAIG GUYER3 & JAY M. SAVAGE4 1Department of Biology, Central Michigan University, Mt. Pleasant, MI 48859, USA. E-mail: [email protected] 2Department of Biology, Southeastern Louisiana University, Hammond, LA 70402, USA. E-mail: [email protected] 3Department of Biological Sciences, Auburn University, Auburn, AL 36849, USA. E-mail: [email protected] 4Department of Biology, San Diego State University, San Diego, CA 92182, USA. E-mail: [email protected] Magnolia Press Auckland, New Zealand Accepted by S. Carranza: 17 May 2012; published: 11 Sept. 2012 KIRSTEN E. NICHOLSON, BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE It is time for a new classification of anoles (Squamata: Dactyloidae) (Zootaxa 3477) 108 pp.; 30 cm. 11 Sept. 2012 ISBN 978-1-77557-010-3 (paperback) ISBN 978-1-77557-011-0 (Online edition) FIRST PUBLISHED IN 2012 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/ © 2012 Magnolia Press All rights reserved. No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any means, without prior written permission from the publisher, to whom all requests to reproduce copyright material should be directed in writing. This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose other than private research use. ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition) 2 · Zootaxa 3477 © 2012 Magnolia Press NICHOLSON ET AL. Table of contents Abstract . 3 Introduction . 4 Current Systematic Status . 4 Phylogenetic Analyses . 9 Phylogenetic Inference . 10 Systematic Accounts . 13 Family Status of Anolis (sensu lato) . 13 Family Dactyloidae Fitzinger, 1843 . 16 Genus Dactyloa Wagler, 1830 . 16 Dactyloa latifrons Species Group . 20 Dactyloa punctata Species Group . 21 Dactyloa heteroderma Species Group . 22 Dactyloa roquet Species Group . 23 Genus Deiroptyx Fitzinger, 1843 . 23 Deiroptyx occulta Species Group . 25 Deiroptyx vermiculata Species Group . 26 Deiroptyx chlorocyana Species Group . 26 Deiroptyx equestris Species Group . 27 Deiroptyx hendersoni Species Group . 28 Genus Xiphosurus Fitzinger, 1826 . 28 Xiphosurus chamaeleonides Species Group . 29 Xiphosurus cuvieri Species Group . 30 Genus Chamaelinorops Schmidt, 1919 . 30 Genus Audantia Cochran, 1934 . 31 Genus Anolis Daudin, 1802 . 32 Anolis lucius Species Group . 34 Anolis alutaceus Species Group . 35 Anolis angusticeps Species Group . 36 Anolis loysianus Species Group . 37 Anolis carolinensis Species Group . 37 Genus Ctenonotus Fitzinger, 1843 . 38 Ctenonotus bimaculatus Species Group . 40 Ctenonotus distichus Species Group . 41 Ctenonotus cristatellus Species Group . 41 Genus Norops . 41 Norops sagrei Species Group . 44 Norops valencienni Species Group . 44 Norops auratus Species Group . 45 Evolution of ecomodes in the family Dactyloidae . 50 Ancestral Node Reconstruction - Ecomodes . 51 Biogeography of the Dactyloidae . 55 The fossil record of Dactyloidae and related families . ..