Reclassification of North American Haplopappus (Compositae: Astereae) Completed: Rayjacksonia Gen
AmericanJournal of Botany 83(3): 356-370. 1996. RECLASSIFICATION OF NORTH AMERICAN HAPLOPAPPUS (COMPOSITAE: ASTEREAE) COMPLETED: RAYJACKSONIA GEN. NOV.1 MEREDITH A. LANE2 AND RONALD L. HARTMAN R. L. McGregor Herbarium(University of Kansas NaturalHistory Museum Division of Botany) and Departmentof Botany,University of Kansas, Lawrence, Kansas 66047-3729; and Rocky MountainHerbarium, Department of Botany,University of Wyoming,Laramie, Wyoming82071-3165 Rayjacksonia R. L. Hartman& M. A. Lane, gen. nov. (Compositae: Astereae), is named to accommodate the "phyllo- cephalus complex," formerlyof Haplopappus Cass. sect. Blepharodon DC. The new combinationsare R. phyllocephalus (DC.) R. L. Hartman& M. A. Lane, R. annua (Rydb.) R. L. Hartman& M. A. Lane, and R. aurea (A. Gray) R. L. Hartman & M. A. Lane. This transfercompletes the reclassificationof the North American species of Haplopappus sensu Hall, leaving that genus exclusively South American.Rayjacksonia has a base chromosomenumber of x = 6. Furthermore,it shares abruptlyampliate disk corollas, deltatedisk style-branchappendages, and corolla epidermalcell type,among other features,with Grindelia, Isocoma, Olivaea, Prionopsis, Stephanodoria, and Xanthocephalum.Phylogenetic analyses of morphologicaland chloroplastDNA restrictionsite data, taken together,demonstrate that these genera are closely related but distinct. Key words: Astereae; Asteraceae; Compositae; Haplopappus; Rayjacksonia. During the past seven decades, taxonomic application lopappus sensu Hall (1928) are reclassifiedand are cur-
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