Vol. 3, No. 1, March 1989 41
A Revision of the genus Octotoma
C.L. Staines, Jr. 3302 Decker Place Edgewater, MD 21037
Abstract There are three other species names connected The New World species of Octotoma are revised with Octotoma: O. tessellata Maulik which IS a and keys, diagnoses, and descriptions are provided synonym ofPhysocoryna scabra Guerin-Meneville for the Ulne known speCIes. The authorshIp of the (Ohmann, 1932); O. daguerrz Pic which is now in genus belongs to Chevrolat rather than Dejean. Uroplata (Uhmann, 1938); and O.germaini Pic Octotoma intermedia, from Mexico, is described as which is now in Acritispa (Staines, 1988). new. The biology and life history of several Octotoma species have been fairly well studied Octotoma scabrzpennzs IS the best studIed speCIes because of Introduction its wide use in the biologieal eontrol ofLantana sp. The name Octotoma was first published in (Verbenaceae). Other species are known to a lesser Dejean (1837). In this catalogue there was no gener extent and seveml have been used, but unsuccessful ic description, but the name was validated by the ly, against Lantana sp The life history will be dis- transfer of Hispa plicatula Fabricius to this new cussed further under each species. genus (Article 12 b (5) Internat. Code Zool. Nomen. There has been no comprehensive study of 1985). Though authorship of the genus has often Octotoma. Uhmann (1927) presented a key to the been attributed to Dejean, it actually belongs to described species. Uhmann (1950) discussed the Chevrolat. Dejean (1837), in the introduction, gave distinguishing elytral sculpture and shape for seven responsibility to Chevrolat for genera produced out species. Other characters used in distlllgmshlllg of the old genus Hispa (Barber & Bridwell, 1940) species have been the presence of metallic sheen on One other worker has been credIted With authorship the elytra; the sculpture of the clypeus; and the of Octotoma, namely Suffrian (1868). Full citations thiekness, length, and shape of the antennal seg- are presented below. ments. All of these characters have been re-exam- Guerin-M€meville (1844) described O. scabri ined as well as the male genitalia, I have found the pennis from Mexico Suffrian (1868) described 0 latter not to be useful in determining species. gundlachll from Cuba. Chapms (1875) gave the first The speCIes of Octotoma fall into two distinct generie deseription. Horn (1883) deseribed O. groups called "typical" and "atypical" by Uhmann marginicollis from Arizona and redescribed O. plica (1950). The "typical" species: plicatula, scabripennis, tuta. Baly (885) described O. championi from gundlachii, marginicollis, and championi have 10 Y2 Guatemala and redescribed 0 scabripennis and 0 rows of punctures, well developed elytral carinae, margznzeoills. Welse (1910) deSCribed O. crasszeor and asymmetrical elytraI sculpture. The "atypical" nis from Brazil and later described O. brasiliensis speeies: brasiliensis, crassicornis, nigra, and varie- from the same country (Weise, 1921). Uhmann gata have 8 Y2 rows of punctures, recognizable elytral (1940) described O. nigJa from Brazil and later costae, symmetrical elytral sculpture, and are para]- described O. variegata from Brazil (Uhmann, 1954). leI-sided. Since many of the genera of Uroplatini are Uctotoma IS III the tribe Oroplatllli (WeIse, dlstmgmshed primarily by the antennal structure, leIla) which is characterized by having the last all speeies should be retained in Octotoma for they antennal segments very closely united and the are uniform in this regard. antennae appearing as three- to eight-segmented. In the "typical" species the elytral sculpture is The tribe is represented by thirty-two genera. almost impossible to describe. The various longitu l Ictotoma is distinguished hy the follOWing combina- dinal and transverse carinae are highly variable in tion of characters: antennae short, 8-segmented, placement and degree of development among indi- ending in a club, segment I subglobular, II viduals of the same species. Uhmann (1950) at cylindrical, VII as wide as VIII; antennal club not tempted to describe the elytral sculpture by dividing longer than the four preceding segments combined. 42 Insecta MundI
each elytron into quarters and establishing a series midline. Elytra: each elytron at base slightly expla- of "honeyeombs" (Honig.....abe). nate, margin more or less irregularly arcuate, slight Types were examined for all species studied ly overlapping pronotal base; with 8 Y2 or 10 Y2 rows except O. charnpioni, O. gundlachji, and O. plicatu of punctures; costae may be developed, some species lao Lectotypes are designated for 0 brasiliensis and with transverse and lateral carinae, these species O. craSSlCornzs. In recordmg the label data from havmg asymmetrIcal sculpture on each elytron. type specimens, a slash (/) divides data on different Legs: tarsal segment I small; II twice as long and labels. wide; III longer than II, barely surpassing lobes of Measurements were taken with an ocular IV, all with setae beneath, claws divergent, empodi micrometer. Pronotal length and width were taken urn between claws along the midlmes. EIytral WIdth was measured at the humeri. Elytrallength was measured from the Key to the species of Octotoma Chevrolat base to apex. Total length was measured from the base of the frontal sulcus to the apex ofthe elytra. 1. Elytra with distinguishable costae , 2 1'. Elytra without distinguishable costae 5 Octotoma Cllevl olat 2(1). Elytral costa I interrupted 3 Octotorna Chevrolat 1837:390. 2'. ElytraI costa I entire; BrazIl . · variegata Uhmann Octoto rna Dejean: Melsheimer 1853:119; Chapuis 1875.311, Monras & Bechyne 1956:1135; 3(2). Elytral costa I interrupted Illore than IIhmann 1957'116; Balsbaugh & Hays 1972'187; once 4 WIlcox 1975:144; RIley & Enns 1979:80; Seeno 3'. Elytral costa I mterrupted one time; BrazIl & 'Nilcox 1982:161. · nigra Uhmann
Octotoma Suffrian. Hom 1883.299, Baly 1885.87, 4(3). Elytral costa I inten upted three times, Blatchley 1910:1225; Weise 1911a'31, 1911b'45; elytra expanded to apex; Brazil . Leng & Mutchler 1914:458; Leng 1920:303; · brasiliensis Weise Blatchley 1924:44; Uhmann 1927:135, 4'. Elytral costa I interrupted fuur times (Fig 1937b:336; Brimley 1938:230; Blackwelder 1); elytra parallel- sided; Brazil .
•• 'tXT 4 1946.729; Uhmann 1947.113, 1950.270; Papp · .....•..•.•...... crasszCOfflZS'f else 1953:92; Guerin 1953:102; Ford & Cavey 1985:53. 5(1'). Carmae over entire elytra ...... 7 5'. Carinae only on apical half of elytra . . . 6 Type species: Octotoma plicatula (Fabricius), type species by monotypy (Article 68d Internat. Code 6(5'). Elytra not strongly expanded at apex (Fig. Zoo1. Nomen.). Uhmann (1950) designated O. sca 2); larger species, 4.8-6.0 mm; southwest bripennis as the type species under Article 67a Of the II S and Mexico marginicollis Horn Code While Dejean (1837) listed. both plicatula and scabrzpennzs Dejean, there was no species descrip 6'. Elytra strongly expanded at apex (F'Ig. 5); tion ofscabripennis thus making it a nomen nudum. smaller species, 3.6 mm., Mexico . Uhmann's (1950) designatioll of O. scabripennis as · intermedia n.sp. the type species is not vaiid ooeause it does not meet the requirements of Article 69a (Inter. Code Zoo1. 7(5). Pronotum black; body with metallic sheen; Nomen.). Cuba ., gundlachzz Suffrran 7'. Pronotum predominately red, body 'Nithout Description: Color variable, either black, black with metallic sheen 8 reddish pronotum, brown, or hrown with yellow Head: median sulcus present or not; antennae 8 8(7'). Apex of elytra with large flattened lamina segmented, as long as head and pronotum com (Fig. 3) 9 bined; segment I subglobose; VII as wide as VIII; VIII 8'. Apex of elytra without large flattened lam- clavate. Pronotum: transverse; anterior angles ina (Fig. 4); Mexico to Nicaragua . produced into tooth 011 either side of head; margined .... scabripennis Gu€nin-M€meville laterally; four depressions, two before and two after Vol. 3, No. 1, March 1989 43
9(8). Antennal segment II longer than I, III longer than II; lateral margin of elytra distinetly Distribution: Brazil. bisinuate; head without median sulcus; south Texas to Panama. ... championi Daly Spechllells exalllhied. BRAZIL (DEIC). Total.i. 9'. Antennal segment I and II equal in length, III barely longer than II; lateral marglD of Octotoma crassicornis Weise eIytra weaklY bisinuate; eastern half of the Figure 1 United States; head with median sulcus ...... plicatula (Fabricius) Octotoma crassicornis Weise 1910.146 (Lectotype (here designated): 28381/ Brasilien/ 244/ Octotoma blasiliensis vVeise Typusj J. Weise det.j Oetotoma erassieornisj 231 Brasilien/28381 Zool Mus Berlinl Lecto- Oetotoma braSILlenSIS WeIse 1921:270 (Lectotype type, Oetotoma eraSSlcornzs Weise, det. CL (here designated): Brasil, Sehaumj brasiliensis Staines, Jr. Paraleetotype: Typusj J. Weise det.j n. sp./ call. Haag/ typus/ Weise det.j Oetotoma Brasilien, 28381/ Zool. Mus. Berlin/ Paralecto- blUsiliensis, typel syntypej Colt DEI, Eberswa- type, Octotoma crassicornis Weise, det. CL hIe (DEIC» IIbmann 1927'135; Blackwelder Staines, Jr (ZMHB» Weise 1911 a'31 , 1946:29; Ohmann 1950:260; Papp 1953:93; 1911b:45; Ohmann 1927:136; Blackwelder Uhmann 1957:116; Gaedike & DObler 1971:344. 1946:729; Uhmann 1950:261; Papp 1953:93; Uhmann 1957:116. Desel iptioll. Dody flat, black variegated with dark reddish-brown. Head' vertex micropunctate; Description' Body color yellowish-brown; head and medIan sulcus deep; antennae mostly 11ght reddIsh pronotum lIghter. Head: vertex micropunctate; brown; segments I II reddish; V VI broader than median sulcus present, deep; antennae thick, preceding; front light reddish-brown; clypeus dark segments I-II subequal, as long as wide; IV-VI trans er. Prortotum. COvered with large, coarse punc- verse, decreasing in length, V-VI wider than IV. tures; lateral margins evenly curved; basal margin Pronotum: margined laterally, four clusters of bisinuate; disc highly elevated; median callus dense, large punctures on disc; midline impunctate, present; anterior margin straight; surface between rest with scattered large punctures; raised areas punctures micropunctate; pronotal length 0.6 mm. micropunctate; pronotal length 0.6-0.7 mm. (avg. (0 1); ,...idth 0.9. Seutellum: yeHovlish; mieropune 0.65, n 2); vlidth 0.8 0.9 (avg. 0.85). Elytra: bro...... n tate; orbicular. Elytra: lateral margins dentate, except for apical fifth which is yellow; almost paral- expanded to apex; narrowed after humeri; dilated at lel-sided; lateral margins dentate, most pronounced apex; apical margins dentate; 8 rows deep, coarse at exterior apical angles; four straight costae; costa I punctures; tncostate; costa I mterrupted four times weakly developed at base, Interrupted three times, so that it forms five seetions, one medially and two seetion before and after middle, last section, the preceding and following; costa II interrupted three longest beginning at elytral declivity and ending times, on basal fourth, just beyond middle, near before reaching apical margin, section behind apex; costa III entire, depressed in middle; puncture middle curved; costa II interrupted four times, basal rows VII and VIII reduced to smgle row m mIddle; sectIOn longest, sectIOn two weak and low, section base reddish, remainder blaek with reddish mark three in the gap between seetions two and three of ings; elytrallength 3.1 mm; width 1.4. Legs: red costa I, section four ends at elytral declivity, section dish; micr op unctate. "v'enteI. pIO-, meso·, and five short, slightly curved; costa III short, most visi- metasterna punctate in middle, rugose at sides; ble at humeri; IV fine, weak, interrupted once, abdommal sterna micropunctate, black m mIddle, strongest at apex; all costae umted by transverse reddish at sides. Total length 3.8 mm. earinae, all raised seetions of eostae strongly pro duced; elytrallength 2.5-2.7 mm (avg. 2.69); width Discussioll. Life history unknown. Immature stages 1.3. VenteI. pro-, meso-, and metasterna black; undescribed. This species is most similar to O. metasternum punctate in middle; metasternum crassicornis It can be distinguished bY the expand· p1mctate; abdomen yellow Total length 3.2-3.3 mm ed elytral apices and elytral costa I being interrupted (avg. 3.25). three times. Discussioll. Life history unknown. Immature stages Larval hosts: Unknown. undescribed. This species is most similar to O. 44 Insecta MundI
nigra. It can be distinguished by elytral costa I being 4'. .~ •• v ....~u, v ..~ ~vuv~. "'''''''6 strongly produced and the pronotum havine- de- ~ . lJ~<:U- LlJ~ ua~aJ margIn. \ I Larval nosts: Unknown.
Distribution: Brazil.
Sm'ldm • Rl'A7i1 ".M HK 'Rr<>C!i1 ~"r""" r ~/1A11OO4 . TOtal: is. ( "- UCIOIoma nlara unmann ------1 ~ ? Octotoma nil!ra Uhmann 1940'4fi4 (Holotvne~ Reenl ~ Dl:llJil:l1 .lloJOLypusl VClOlOma nlgra un., TTl- (I<>t <.tSl./ • ~ ... / rf'\ (l\ Paratypus I KM N H». Blackwelder 1946:729; !1mm ••• n Tn \ ~vvv._vv, ~ ~t't' ~vv~.v~, l 1957:116' Gaedike & Dobler 1971:355.
. 'Roilv hl"t'1r .. •_C!;(I<>iI H ....i1· hl",,1r· vertex mlCropunctate; median sulcus present; two 1. ',1. ·.J.4'.1 • ~ '-'0' v .. ~.v••~. u.~~ v. u~."'~u ..~~. ~. "u, antennae in pit; antennae light reddish-yellow; • OY .. • • / .1.- , .l.l _" ,• , l:li:l JUll~ as I' III '.J : tn "n<>y lnna<>r th"n TT· TV_VT tr"nC!_ \, ~ "- ~ verse, punctate; v11 wan rows or setae; v111 elllpu- ~ nnl ..';Tl. .~"Tn ~4' n .•Tn...... ,;, .n 1. 'un no. , , _u tum: covered with larg,e coarse punctures' , disc with longitudinal furrow; area between punctures micro- punctate; basal area around disc depressed; basal Figs. 1-4. Left elytron of Octotoma species. 1. crassicomis. 2. margin sinuate; lateral margins weakly convergent; marglnlcolLLs. 3. pLLcatula. 4. scabnpenms. pronotallength 0.7 mm. (0-3); width 1.2. Scutel· Ium: quadrate; reddish-brown. Elytra: eight rows Larval hosts: Unknown. of punctures; scuteBar row of two punctures; rows VII & VIII united at middle; costa I interrupted in Distribution: Brazil. middle, more raised on apical balf; II raised at base, in middle, and apex; III weak, most visible at apex; Specimens examined' BRAZIl,' no further data elytra narrows after humeri; apices individually (ZMHB). BahIa (BMNH); BraSIlIa (BMNH). Total: rounded; lateral margin serrate, apical margin 3. irregularly toothed; costae II & III united at apex; elytral length 3.8-4.1 mm (avg 39); width 19. Octotoma valiegafa Uhmann
Venter: pro- meso-, and metasterna black; abdo Octotoma varzegata Ohmann 1954:499 (Holotype: men yellow. Legs: reddish brov/n. Total length 4.5 2683/ BRAZIL, Dr. G. Bondar! Holotypusj E. 4.9 mm (avg. 4.7). Uhmann det. 1952, Octotoma variegata Uh. (BMNH); type locality Bahia (Ullmann, 1954». Discussion: Life history unknown. Immature stages Gaedike & Dobler 1971:362. undescribed. 'this species is most similar to O. crassicornis. It can be distinguished by elytral costa Description: Body color reddish-brown with black I being interrupted one time, the weak costae, and markings, parallel-sided. Head: reddish-brown; the lack of depr essions near the base of the pr ono- median sulcus present; frons dark, transverse; row tum. of punctures on inner margin of each eye; eyes Vol. 3, No. 1, March 1989 45
coarsely faceted; vertex micropunctate; antennae longer than II; IV-VI transverse; VIII elliptical, yello'N brown; segment I II globular, subequaI; III hirsute. Pronotum: reddish; anterior margin black; smaller, conical; IV-V conical, shorter than III; VI row of punctures on anterior margin; six black transverse, VII wider than VI, as long as II. Prono maculae, one on each lateral margin in middle, two tum: twice as long as wide; covered with large, transverse on anterior half, two large cirClllar ones coarse punctures, except dISC WhICh IS Impunctate; on posterIOr half; dISC smooth, except for several lateral margins almost parallel anteriorly, weakly large punctures; pronotallength 0.7 0.8 mm (avg. convex after middle; front angles acute; pronotal 0.7; n= 15); width 1.3-1.4 (avg. 1.4). Scutellum: length 0.8 mrn. (n 1), width 1.1. Scutellum. red alutaceous, reddish-yellow , transver se. ElytIa. dish-brown; quadrate. Elytra: with eight rows of black; carinae over entire surface; lateral margins punctures; with scutellar row of three punctures; dentate; apical margins more dentate; exterior apical narrows slightly after humeri; tricostate- I and III margin with flattened lamina; elytrallength 3.6-4.4 entIre, II Interrupted In mIddle, II and III united at mm (avg. 3.9); width 1.7-2.0 (avg. 1.9). Legs: red- apex, none strongly produced; lateral margin finely dish; pro and rnesofemur with tooth, punctate; serrate; suture carinate; apical margin finely den- protibia flattened. Venter: alutaceous at sides. Total tate; apices individually weakly rounded; elytral length 4.3-5.3 lIUll (avg. 4.7). length 3 ] mm; width 1 5 Venter' dark Legs' femur clavate. Total length 4.0 mm. DISCUSSion: LIfe hIstory unknown. Larva and pupa described by Uhmann (1949). The larval mine is Discussion: Life history unknown. Immature stages very narrow, with a very faint fecal trail. Pupation undescribed. This species is most similar to O. occurs in the mine Ilear the starting point (Ulllnann, nigra. It can be distinguished by elytral costa I being 1937a) Adults feed on foliage by destroying the entire. upper epIdermIS of leaves (DIatloff, 19'1'1). ThIS species is most similar to O. plicatula. It can be Larval hosts: Unknown. distinguished by antennal segment II being longer than segment I, elytral carinae sharper and more Distribution: Known only from the type locality. elevated; sinuation of lateral elytral margin anterior to apical margin more distinct; apical elytral angle Specimens examined' BRAZIL (BMNH) Total' 1 more angular and more coarsely serrate; and the vertex of the head without a median sulcus. Octotoma champiani Baly Map 1 Larval host plants: Lantana camara L., L. hispida H. B. & K., and L. trifolia L. (Diatloff, 1977). Diatloff Octotoma championi Baly 1885'88 (type not seen; (1977) tested feeding and oviposition of 0 champio- type locahty Guatemala). Donckier 1899:571; nz on 52 plant speCIes In 27 familIes. Minor adult Vleise 1911a:31, 1911b:45, 1921:270; Uhmann feeding was noted on },{entha sp., Oreganum sp. 1927~135, 1934:274, 1937a:64; Maulik 1937:139; (Labitae), and Sesamum sp. (Pedaliaceae). Blackwelder 1946.729, Uhmann 1949.25, 1950'266; Papp 1953'93; Uhmann 1957'116, Distribution' South Texas to Panama 1961:20; Harley 1969:836; WIlcox 1975:144; Diatloff 1977:165; Julien 1982:50; Riley & Specimens examined: COSTA RICA. Cartago, 2flII/1974 Balsbaugh 1988:150. (DSNM); San Jose (DSNM); Turnalba, IX/1956 (HDOA). GUATEMALA: (USNM). HONDURAS: Sta. B, 13 km SE EI Mllooitll, 16/VII/lS77 (EGRC). MEXICO: DF (USNM). Ghia Octotoma sp. probablyplicatula: Krauss 1962:136, pas- 25 mi S. Malpaso, 25/V/1969 (CNC). Colima- Vuleano 1964:92. (USNM), Valkan (USNM). MOlelos- Caernavaca, V/1945, VIII/1955, IX/1955 (USNM). Nayarit- Tepic, 20-24/VII/1952 lUAIC). San LUIS Potosi- 13 mi W. EI Naranjo, 3200 It , Octotoma sp.: Fullaway 1958:73. 30/VI/1965 (TAMU); EI Saito de Agua, 28-30/VII/1960 (CNC). Tabasco Campo Exp CSAT, 20 km W Cardllnas, 22JVII/19110 (TAMU). Tamauhpas- Bacatoma, 7 km SSE Gomez Fanas, 25 Description: Head and elytrablack, pronotum reddish 30/II1/1978 (EGRC); Mun. Gomez Farias, La Florida, "POlO Witb black maculae Head, vertex black; micropunc- Azul", 2fTV/1987 (PRJC) Veracruz- SD Rafael, Jicaltllpec tate; median sulcus absent; row of punctures along (MCZC); Veracruz, 6/X/1962 (USNM). NICARAGUA: Jinote ga S mi SE JiflBtega, 11100 ft., HI/VII/ISH (EGRG). PANAMA. inner margin of each eye; front reddish; antennae Chiriqui- Las Lagunas, 4 km W Hato de Vulcan, 1360 m, di vided bJ keel; clypeus punctate; antennae reddish; 13/X/1915 (STRl). UNITED STATES. Texas. Cameron Co.- segment II transverse, longer than I; III cylindrical, Brownsviile, 12jV/1919 (USNM); 6 mi NW Brownsville on HWY 46 Insecta MundI
281, 6-7/X/1984: Sabal Palm Grove Sanct., 28·29/III/1986 Octotoma gundlachi' Gemminger & Harold . onesT , c 0 oma In ermedza 1876:3610; Donckier 1899:571; Weise 1911a:31, (depos~t~~n 19116:45; Leng & Mutabler 1914:458; Uhmann Staines, det. CL Staines, Jr., 1987. PRJC). MEXICO: Tampaulipas, Nacimiento e 0 1927:185, 1982:~66i Blackwelder 1946.729; Papp Frio, 5 mi. S. Gomez Farias, 400ft., 5-6....i 88, R. 1953:93; Vaurle 1956:80; Fullaway 1958:73; A d r"on ical ecid. forest PARATYPE Octo- oma in erme ia nes, e. .. Bruner, et. al. 1975:185. (deposited in 1.8 Askevold collection) Description: Body color black with metallic sheen. Description. Body color yellow with brown markings. Head: vertex alutaceous; row of punctures on inner malgin of eye; median sulcus present; keel present between antennae, extended onto vertex; antennae reddish-brown, darker at apex; I-VI punctate. t lllner t an~il ongest;r~nsverse;~rga: Pronotum: margins black, disc reddish-brown' brous; VII VIII hirsute. Pronotum: yellov.' ...lith two aoyered with coarse, dense punctures' anterio; black maculae; anterior margin with strong tooth on margm. prolonged into tooth on either side' of head; either side of head, lateIal rnaIgins con'Vergent to pronotallength 0.7-0.9 mm (avg. 0.76, n= 10); width mar~ bisinu~~; c~~ered 1.1-1.6 (avg 1 4) Scute])um' black, alutaceons base; basal with large Elytra: carinae over entire surface; lateral margin sparsely dentate; apiaal margin dentate; elytral n 2); width 1.0-1.1 (avg. 1.0). Scutellum: yel ow; length 3.6-4.3 mm (av~. 4.0); width 1.9-2.3 (av~. 2.1). guadute; micIopunctate. Elytra. yellow with Legs. black, femUI pu lctate at apex, pwfemu with r wn arkin S' ex nded to a ex' exterior a ical ang es grea y expan e ,s ope; black. Total length 4.3-5.1 mm (avg. 4.7). dentate, more so at base, sinuate; carinae on apical half of elytra, most distinct in middle; sutural Discussi~n: h~story u~known Life ..Immature sta;es malgin carinate; elytrallength 2.9 8.0 mm (avg. undescIlbed. ThIS speCIes was lnttoduced h to 2.9); width 1.5-1.6 (avg. 1.5). Legs: tibiae widened to Hawaii as a biolo~cal contr: =i~stLa:~ middle, narrowed to apex; base yellow, apex black, sp., but has not ~come esb¥0 aus~ 0 It is similar to O. ckampioni, O. pliootula, and O. scabripennis, and can be distinguished by the black pronotum, the metallic sheen of the bodrand by its yellow in center and sides, rest black; mesosternum black, micropunctate; metasternum black, punctate at sides; abdominal sterna dark medially, yellow at Larval hosts' Lantana sp. (Vaurie, 1956). sides. Total length: 3.6 mm. Distribution: Cuba. Etymology: Combination of inter between and medius middle, Latin; for the unifying chmacteI of Specimens examined: CUBA: no further data the elytral sculpture. (IISNM); 14/IV1953, IV/1953 (HD01\). Loma de Camoa, Habana, 14711171952 (CLS). Havana, Discussion: Life history unknown. Immature stages IV/1952 (HDOA). Paro del Niedis, 2/'11/1933 undescribed. This species is most similar to O. marginicollis. It can be distinguished by its smalleI size and the expanded elytral apices. Oriente, ]918 (USNM) Total' 27 Vol. 3, No.1, March 1989 47 1953:93; Uhmann 1957:116; Gibson & Carrillo "''''''''"jJ''VII• .uuu,Y l;UiUi ,Yt::llUW W iLll Ui UW 11 Ui Uii1\;.... •. or black. Head~ vellow: eves dark' anten- nal segment 11 larger ana WlOer tnan 1, transverse; , , punctate; row of parallel punctures on each side of "'Ui"U", "'.Y"''' l<1""'L"'U. .. .• marP"in with deen nit on each side of midline' cov- ereu wnn coarse, uense punCtures; completelY . '." . ,,. _l"."u... ' . ".. entirely dark;'. length 0.7-1.1 mm (avg. 0.9; n=26); 1A1,., ">\ ,.~~•• ~.~-~. '~'E>' ~.~. • J~"~ '''1~~~ I rate' microDunctate. Elvtra: base yellow aDical half , , , ~ ~ . Uli1l;J\. Ui Ui1iJ\.' .lU iUW;:; Ui PUll\;- tllr.,,, "t h""., nl11" rnur nf ~ lateral and apICal margms dentate; costa 11 distinctly / _ft:ft,,~ ~~._ ~. I,'~ 1.. .."" •• ,ft! '" , , 'r carinae difficult to follow puncture rows' apical Legs: nOla Witn tutt or setae on mner apical margin; ~ft_., , '.~ •.ftl...... ,I,:_~ft Venter: prosternum reddish-brown anteriorly, ", ., '" Vla".... • ,lV' Vl. C:UVU5 ._ , anex of nrocess rUl1'Ose' mesosternum medially transverse, punctate, rugose laterauy; metasternum punctate, more dense laterally; abdominal sterna- 1- Fig. 5. Habitus ofOctotoma intermedia, n.sp. II punctate in middle, rest rugose; III rugose later8.1 anteriOr margin, with 3 rows of punctures; IV-V Larval hosts: Unknown. with 3 rows of punctures; VI with 4 rows of punc- tures. Total length 4.8-6.0 mm (avg. 5.3). Distribution' Tamalllipas, Mexico Discussion: Life history unknown. Immature stages Specimens examined. MEXICO: Tamaulipas Mun. undescribed. This species is most similar to O. Cd. Victoria, Canon La Libertad, 7/111/1986 intermedia. It can be distinguished by the elytral (IISNM); Mun Gomez Farias, La Florida, "Pozo apices not oeing expanded and its larger size It is Azul", 2/IV/1987 (PRJC); Nacimiento del Rio Frio, 5 variable in color, with a bicolored yellow and brown ml. S. Gomez Farias, 400ft., 5-6JVI/1983 (Askevold). form and an entirely black form. The placement of Total: 3. the depressions and carinae on the elytra are quite variable. Octotol17B I17Brginicoffls HOI n Figure 2, Map 1 Larval hosts: Perezia thurberi Gray (Compositae) (Jones & Bnsley, 1925). Adults have been collected Oetotoma marginieollis Horn 1883:300 (Lectotype: offFraxinus sp. (Oleaceae) (Chittenden, 1902). Ari[zona], type 3880, here designated; Paralecto types- AtiFzonal, Paratype (numbered 3880.2 Distribution: Arizona, southern California, New through 3880.10), MCZC). Baly 1885:88; Mexico, Texas, and Mexico. Champion 1894:240; Donckier 1899:571; Chit tenden 1902:89; Weise 1911a:31, 1911b:45; Leng Specimens examined: MEXICO: Serrino, Buena Vista, 1920:303; Weise 1921:270; Frost 1924:465; Jones 6000 ft., Sierra del Carmen, Coah., 18/VII/1938 (TAMU). & Brisley 1925.174; Ulunann 1927.136; Black- Durango- VII/l896 lUSNM!; 5 ml W. Durango, l/VIII/I964 welder 1946:729; Uhmann 1950:262; Papp (CNC); 24 mi W. La Ciudad, 20/VI/1964 (CNC). Jaliscoo mts N 48 Insecta MundI 115 '", 'm '0 0< 00 A' AO 75 70 65 60 .- ') .. I 1---1 I ~y ~ .. • i-----\ \ 'f .. 30 -\ ~ '\ -:='? )~\ \.1 \'\ ( y,\ / y \ c\~) VI '" • 1 ../ '" " U~ 25r- '(~ \~ " i\ 'II - " '\. "f-.. ...---. ~----...-..v 4{.•• '\\ '\"\ AI" /\ ~ •\ ~ _~O·· v r-. ~ (.l .~ ,./ ...... ~ . , \.\D:'}~ ~\ ~ 5 ... c7f <; <:> ---- .ur-- }J ,.....Lv, r 'l -v-J- u ~ 7Jl:::'_~~ \Ji . , C> ~'\ 10 ~;::(.?~ FJh /~ 15_ ~f: /()\ t~~/ \~ .-. _5 lOt-- ~ '\ \ l "- ~ ~_O ( 5 t-- r t )~\ ~. o t-- VJi t I II III \. v I'" '( \ 110 '0< ,nn 0< Map 1. Distribution ofDc/aloma species. champia/li (squmes), gU/ldtaehii (ell des), IIIwgi/licaliis (tIiangles). vI lOSNM), 15Jvli931 (CNC), I4JVIJI926 (CNC), VIIJ1936 17fVIIT/1953, VI/1959, VII/1965 (USNM), VI (CASC). S~n Luis (CASC), 9/VII!1940 (SEMC), 14fVI/1928 (MCZC); Huachuca Potosi- Ciudad del Maiz, 3000 ft , 23jVIU/1954 (CNC). Sonora 9 Mts., Carr Can., 5000 ft, 4/VI/1986 (TTSNM), 15/VII/1932 mi S Conanea, 16/IX/1970 (FSCA). UNITED STATES: (CASC); Garden Can., 23/VI/1950 (CNC); Miller Cyn., ARIZONA: state label onI, (CUIC, MCZC), S. E. AI iz. (CUIC), 19f1II/1987 (CLB), Sawmi:ll Can., 22/VI/1986 (USNM); PalmeF Atascasa Mts., 2/X/1933 (CUIC); Ft. Grant, Pinaleno Mts., 15 lee, VI (USNM), 19JVII/1987 (CLS); 2 nu. SW Portal, 5000 ft, 19/VII/1917 (CUIC); Madero (MCZC); Morrison 31N/1973 (CHAH), 4 mi SW Portal, 20/VII/1963 (CNC), 3 mi (MCZC,USNM). Pena Blanea, 14/IX/1947 (USNM). Goehise Go. W Portal, 115flIII/1979 (GRAH), 4 mi BE Portal, 4fVIII/1959 4fVIII/1966 (PMNH); Bisbee, 5/VI/1915 (CASC); Cave Ck., (CUIC); Ramsey Can., 15-19/VII/1912 (CASC), 15fVII/1969 Chiricahua Mts., 22/VI/1967 (CDFA), 7/VII/1908, 27/VII 1908 (MCZC); Southwest Res. Sta.. 5 mi W Portal, 18/IX/1966 (CASC). Coconino Co.- Grand Can., 11/VII/1912 (USNM); Wil liams 8NU (USNM) Gila Co Pine Creek Globe 8lVIl1953 (CUIe): Graham Co.- HospItal Flat, 2jViIIj1965' (CASC). Greenlee Co.- Clifton, 29/VII/1946 (USNM). Pima Co.- Catalina Vol. 3, No.1, March 1989 49 f 'H 1 t~\ \ \ \ ( \ "-/ [)- '" 25 1\ .... \. u ." - 15f-- 10",- > - - , I I 1 I 1 (~j J 110 105 100 95 90 80 75 70 Map 2. Distribution ofOctotoma species: scahripennis (squares); intermedia (circles). Mts., 22/VII/1942, 15/VI/1933 (CASC); Josephine Can., VII/1913 (CNC,MCZC). Hidalgo Co.- 2 mi N Rodeo, 8/VIII/1968 4fVII/1972 (TISNM); Mt Lemlln, 17fVI/1912 (C4.SC)j Madera (FSCA.) TEXAS: Bexar CII. San Antonio, Il/IV/1907 1853:119; Gemminger & Harold 1876:3610; Horn 1883:300; Donckier 1899:572: Schwarz Distribution: Maryland and Dclav,rare west to south 1890:232: Chittenden 1902:88; Ulke 1902:29, 52; ern Michigan and south to Texas. Blatchley 1910.1225, Weise 19Ua:31, 1911b:4o; Leng ]920'303; Dozier ]922']23; Frost 1924'464; Specimens examined' I INITED STATES: ALABAMA: Lee Blatchley 1924:44; Ohmann 1927:135; Mauhk H137:139; Brimley 1938:230; Lading 1945:136; au ner (;o.-a ower, 01 7 E ~ emp ~ Uhmann 1950:266' Pa 1953:93' Wilcox no localit , 29 V/1922 (UADE). Johnson Co.- no locality, , . , Fayetteville (UADE). DISTRICT OF COLUMBIA: no data; Uhmann 1957:116; Krauss 1962'136; ]964'92; '1/1918 (CASC), a/VII, 25/VII/1922 (USNM). FLORIDA: DQ Harley 1969:836; Kirk 1969:98, 19'70:95; Bals further data (FSCA). Alachua Co.- Gamesvllle, 247 V7 1961, baugh & Hays 1972:187; Wilcox 1975:144, 13/YI/1967, 23/YI/1984, 25/YII/1984, 1959 (FSCA). Leon Co.- no loeality, 7/VIII/198 t (FSCA). Seminole Co Sanford, 1979:8,14; Riley & Enns 1979:80; Ford & Cavey 5,011/1927 (CNC) GEORGIA- Clarke Co.- Athens, 23N/1972, 1985:53: Lago & Mann 1987.6. 14/V/1915, 8-14fyI'1916, 4fl/1984 (UGCA), Whitehall Forest, 21/V/1976 (FSCA). Thomas Co.- Thomasville, 29/111/1938 Description: Herod: vertex h1aek, alutaeeous; median suleus lUGCA). ILLINOIS: state label only (MCZC). Jackson Co.- prese.nt; front testaceous; two rows of punctures along inner Carbondale, 27/IV/1908 (USNM). Pulaski Co.- Pulaski, Ilungui of eye, alitellilae separated by keel, anleJlIlae 8..segInent- =I~;~: l~;~no~~- r~~~%~N~G?H:RyOe;~~};,l}~i4 ed, testaceous; segment I thick; II cylindrical, equal to II; III barely longer than II; III-VI transverse; VII hirsute; VIII round- (FSCA). Brown Co.- no locality, 25/V/1985, 16/VI/1982 (FSCA). ed, hirsute. Pronotum: reddish; six black maculae present, one Clarke Co.- HenryVllle, 25/V/1963 (F'SCA). MarlOn Co.- Indian ...... a olis 30 1963, 28 I 1975 (FSCA). Monroe Co.- Blooming- , (CASC). KANSAS: Douglas Co.- Kaw River, 6/V/1979 (SEM9)· , Scutellum: testaceous; alutaceous; transverse. Elytra: black; Co.- Berheim Forest, 9 1/1978 ( ). Istlan 0.- op carinae over entire smfaee; lateral and apieal margins dentate; 8 kinsville. 7 /VI/ 1963 (CNC). Daviess Co.- Owensboro, r?ws of punctures VISible at base, row I VISible to apex; parallel 24/IX/1976 (CDFA). Madison Ca. Riehmond, 2/V/1964, Sided for basal half, then divergent, dilated at apex; apical margin 17/V/1964 (CNC). Ohio Co.- 18/VI/1961 (CNC). LOUISIANA: flattened and expanded; elytral length 3.3-4.0 mm (avg. 3.6); Forbing, 24/1II/WOS (USNM). Baton Rouge Pm.- Baton Rouge, width 1.4-2.0 av . 1.7 . Le s: trochanter testaceous' femur ba e 1O/III/1985, 8/III/1984 (EGRC). Natchioches Par.- Campti, . . : on . elongate punctures ~ear apex; tibia dark reddish-br~wn,with Anne Arundel Co.- Annapolis, 9/IX/1987 (CLS); Chesapeake rior mar n WI rows 0 se e on mner margin, setose at apex; protibia with small tooth. Venter: black, ~l;~~~;:~:':':;Z~lsr;,~=:;:l~:~~;~n~~;t:~,!~~;:'o~;s:~ Sides; metasternum alutaeeous, basal margin rugose in eenter, 311VIII/1982 (EJF); Great Falls, 21/VI/1912 (USNM), center punctate; abdommal sterna aIutaceous m middle rugose 31/VIII/1982 (JFC); Marshall Hall, 20/IV/11l911 (USNM); at sides. Total length 3.8-4.8 mm (avg. 4.3). ' Plummers Island, numerous 24/IV- 19/IX (CASC,USNM). Prince George's Co.- Blandywine, glffIII/198? (CLS), College Discussion: This species forms tentiform mines. Excrement is accumulated as a black spot near the land, 18/IX/1986 (CLS). Somelset Co.- Hopewell, 17fVII/1983; base of the leaf. Pu ation occurs in the mine near Marion, 15/VI/1977; Pokomoke, 16/VII/1982; Shelltown, ar ., e u m e 0 ong 0 es In eaves. e Easton, 18/VIII/1982 (JFC), 19/VII/1982 (EJF). Worcester Co.- larva was described by Ford & Cavey (1985). This species is most similar to O. championi and can be distinguished by antenna! segments I and II being subequal in length, elytral carinae less sharp and less elevated, sinuation of lateral elytral margin less Co. Delta N. F., 8015, 6. t km E of Rolling Fork, 1/X/19110 (CNC). Wilkinson Co.- Woodville, 7/VI/1921 (CASC). MIS- distinct, apical elytral angle less angular and more SOURI. Boone Co.- 8/IX/1915 (EGRC). Cole Co.- Jeffersofl finely serrate, and the vertex of the head with a City, 17/V (USNM). Jefferson Co.- Arnold, 16/VII/1975 median suleus. (EGRCt Randolph Co.- 1 ml E Moberly, 23/VI/1974, 26/VI/1972, 22/VI/1972, 11/VI/1972, 30/IV/1974 (EGRC). St. Francis Co St Francis St Pk, 21fV1/1978 (EGRC) St Louis Co.- Sf: LOUIS, 197 V (USNM). NORTH CAROLINA. state label only (CASC); 26/V/1952, 17/V/1961; Morrow Mt., 21fVII/1959 (breSIl) Duplin Co - Faison 41Xl1963 (NCg,,) Durham Co- Duke Forest, 1/VII/1973 (PMNH). Edgecomb Co.- 24/VI/1927 (EDNG). Gates Co. VIII/1963 (NCSU). Hyde Co. SVlliR Quar and Chionanthus vir inica L. (Oleaceae) (Blatchle ter, 2/V/1952 (FSCA). Johnston Co.- 1/X/1959 (NCSU). Lenior 0.- ...... t ,, 19/IV/1960 (NCSU). Pender Co.- Willard, 16/V/1958, adults from Daucus carom L (Apiaceae) flowers 22/VIII/1957 (NCSII) Wake Co - Cary, 22/1V/1974 (NCSU): Vol. 3, No. 1, March 1989 51 Fiaure 4, Map 2 Y' • nllt>rin_..... ' '" Hl.44~279 ~type not exammeu; type JOcallty MeXICO). -, -01 •• , Champion 1894:240; Donckier 1899:572; Weise ~.,~~... u~, ~v~~u.&V, ~ "UU.." ... V ~V_&.vv, Swezey 1925:376' Uhmann 1927:136' Black- weluer 1::1'1:0: I ~::1; unmann 1::10V:~OO; rapp 10I;:q.OQ. TTY 101;:7.117. UTaha.. 1 0r;:a.1 aA. -1'10 r \-' / "/i/2.'~ -~d\ /' ~'ullaway 1958:73; Gibson & CarrilJo 1959:128; '7 1 nl>...... 1> • nl> A .n... T\ 0 '7 ~vv_.~vv, ~vv~.vv, ~_ '"u ~ 1965:89 1966:204' Uhmann 1966:273' Harlev - . . I• i~ ~" >;~.~ ~ Yadav & Pillal 1!n4:75; WlIcox 1975:144; :~ .~... _~/?r- !/ \ .' - _... TT' , 'O'7a.,o., ,.,.". '0'7'7.')..1,)., ,." 1978:405; Forno & Harley 1979:152; Kamath ~::" v;uo, ~~i:Ult::.Y, t::L. <11. ~VIV;Uil; H'IRO·10')· Julien H'IR')·!'\O· (!illi",rc:l 1QR~'1~7 with black maculae. Head: vertex black, alutaceous; ,. , ,. . ., , "...... " ''''''., ''''0'' ..,,'u Map. 3. Distribution ofOctotoma plicatula. eyes; front reddish; clypeus reddish, coarsely punc- tate; antennae divided by keel; segment II trans Raleigh, 13jVf1951, 2fVIIf1982, 15/X11968 (NCSTT), V/1922, verse; III cylindrical, longer than II; IV-VI trans- IX/1924, 21/VIII/1903, 16/VII/19212, 28/VI/1933 (EDNC), verse; VIII ellIptical, hIrsute, longer than III; 1-VII 5fVIIf1950 (CNC), 21JV/1952 (FSCA). OHIO: state label only (cASC) Clinton Co - no 10C1llity, lfrlvI11961 (FSCA! Hamilton vlith some setae. Pl'onotum: reddish vRth six blaek Co.- Cincinnati, VI/1913 (USNM):I4j'VI/I922 (MCZC). High- maculae, one on each lateral margin at middle, two Ianel Co. noloeaHty, HlfVIl/1961 (FSCA). HoGking Co. no loeal transverse spots OIl anterior half on either side of ity, 17fVI/1961 (FSCA). ~KLAHOMA: Latimer Co.- no locality, , , SOUTH CAROLINA: Anderson Co.- Anderson, 20fVIII/1939 on elt er Sl e 0 ongl u ma carma; covere Wit CUSNMl. Clarendon Co.- Summerton, 23/VI/1984 (EJF,JFCl. large, eoarse punetures; basal margin bisinuate; HOllY Co.- 5 mi W. Myl tie Beach, 15/'1111/1980 (CNC). TENNESSEE: Shelby Co.- Memphis (USNM). Smith Co. front angles acute; pronotallength 0.8-1.1 mm. (avg. Elmwood ICASC). TEXAS: state label only (MCZC!. Belfrage 0.9; Il-36); width 1.8-1.'7 (avg. 1.5). Scutellum: (MCZC). Anderson Co.- lO/IV/1960 (TAMU). Brazos Co.- Col- 28/III/1969 (TAMU). Colorado Co.- 20/VII/1959 (TAMU); ac ; carmae over en re su ace; a r an apl Columbus, 23 25/V (USNM), 24/V (SEMC), 25/V (MCZC). margins dentate; gradually expanding to apex; apieal Gonzales Co.- Palmetto St. Pk., Ottme, 21/IX/I979 ('I'AMO). margin flattened; lateral margin more flattened Harris Co.- Houston, 1892 (USNM), 1886 (MCZC). Harrison Co. Marshall, 25/111/1907 (USNM). Karnes Co. 23fVII/192!l apically; elytrallength 4.2-5.5 nun (avg. 4.9); width (SEMC). Marion Co.- Jefferson, 27/III/1908 (USNM). Polk Co. 2.0-2.3 (avg. 2.1). Legs: pro- and mesofemura with Ace, Menald Cleek, 4 mi N. Romayol, 22/'/1/1984 ('l'AMU). tooth, femur reddIsh at base and apex, black m Victoria Co.- Victoria, 23/III (USNM). Wharton Co.- Wharton, 97 \7/1905 (USNM). VIRGINIA: state label only lCASC,MCZC); middle, with coarse punctures at apex, rugose; tibia expands to apex, protibia flattened apically. Venter: black; I ugose at sides. Total length 5.1-6.4 mm (avg. (USNM). Chesterfield Co.- Richmond (USNM). Fairfax Co. 5.8). 21l/VI/1912; Chain Bridge, ll/lX/1921 mSNMI Spotsylvania Co.- Fredericksburg, 28/IX/1891 (MCZC). WEST VIRGINIA: Discussion: Immature stages undescribed. Life histo- state label only (USNM). Kanawha Co. Kanawha Stn., 29fVII/1919 (USNM). Total: 593. ry was investigated by Harley (1969). This species has been widely released as a biological control agent (Julien, 1982). This species is similar to both O. 52 Insecta MundI championi and O. plicatula. It can be distinguished (CNC), L. LeSage; J.F. Cavey (JFC); S.M. Clark; by the laek of an expanded lamina at the elytral Cornell University (CUlC), R.E. Hoebeke; Florida apices. State Collection of Arthropods (FSCA), RE. Wood- ruff; E.J. Ford (EJF); University of Georgia Larval hosts' Lantana spp (Perkins & Swezey, 1924) (ITGCA), CL Smith; Hawaii Department of Agri- Harley (1969) noted adult feedmg on 1'ectona gran- culture (HDOA), B.R. KumashIro; H.A. Hespenhe dis L., Ltppia umhel/ata (Vernenaeeae), Oreganum ide (CHAH); Humboldt Universitt zu Berlin, F. sp., Mentha spicata L., Salvia occidentalis Hieke (ZMHB); Institut fur Pflanzenschutzfor- (Labiatae), Sesamam indicum L. (Pedaliaceae), schung (DEIC), H. Morge; R. Jones (PRJC), Vigna unguiculata (1.) Walp , Phaseolus lIulagris L, Museum of Comparative Zoology (MCZC), S.R and Stizolobzum aterrzmum O"abaceae). Shaw; North Carohna Department of AgrIculture (EDNC), J.H. Greene; North Carolina State Univer- Distribution: Central America. sity (NCSU), L.L. Deitz; Ohio State University (OSUC), C.A. Triplehorn, Peabody Museum of Specimens examined: EL SALVADOR: San Salvador, Natural History (PMNH), D.G. Furth; E.G. Riley Xj1!l1l5 (USNM), 1953 (TAMU), USll!lltan, 4/VIII/1964 (EGRC); H.P. Stockwell (STRl); Texas A& M (TAMU). GUATEMALA: Sacapulas, 1500 m, 12/VIII/1947 (USNM). HONDURAS. Olancho- 5 m) SE Catacamas, University (TAMU), H,R. Burke; II S National 13/VI/1974, 11 mi NE Catacamas, 1200 ft., 15/VI/1974 Museum (USNM), RE. White. (EGRCt MEXICO' DO further data (MCZC,USNM) Cbiapas- 10 I would like to thank the staff of the Annapolis mi N Chiapa de Corzo, 12/VII/1955 (SEMC); EI Ocotol, 13jVIlIj1969 (CNC); Oaxaco Border, 21 km W Rize de Oro, lela Area Library, Anne Arundel County, Maryland, for m, 6-8/X/1972 (CASC). Chihuahua- Sierra Madre MtS, La Bula, their assistance in locating literature. Also I would 900 m, 7/VII/1972 (CASC).Colima- 10 mi NE Comala, 17- like to thank my wife for her constant support and 19jVIU/1983 (TAMU); Vulcano (USNM). DUFango 24 mi W La Ciudad, 15/VI/1964 (CNC). Guananjuato- DO further data assistance. (MCZC); Salamanea, 4 mi B, 16/VII/1974 (FSCA). Guellelo- Hwy 134, 34-36 kIri NE Jet. 200, 14-16fVI1/1985 (EGRC); 10.3 mi S Iguala, 23/VII/1981 (TAMU). Hidalgo- Minera Autlan, Literature Cited Andres, LA Alitlan, 7/VII/1984 ('FAMU), Chap~la,j l'i'tv'II/l!l66 (EGRC), 6 1973. The bIOlogIcal control of weeds with in mi W Chapala, 30/VI/1963 (CNC); 6 mi E Lagos de Morino, 29!VII/1978 (TAMU), 15.5 ml NE Lagos de Mormo, 621111lt, sects in the United States. in P.H. Dunn (ed,). 26/VII/1952 (SEMC); 12 mi W Plan de Barrancas, 25/VII/1966 Proc. 2nd Internat. Symp. BioI. Control Weeds. (TAW!); 5 5 mj NE Tecolotlan, 13jVII/1982 (CDFA); Hwy 80, COIlllllW. lust. Bioi. Control Misc. Pub. 6.11-28. 10 Illi NE Tecolotlan, MOO ft., UrVIIT71982 (EGRC). MlChoa can 7 mi. W Sall'llaZe, 21/VII/19GB (OBUC); Uruapan, VII/1912 (CASC). Morelos- Cuernavaca, V/1945, VI/1945, VII/1953, Balsbaugh, E.U., & K.L. Hays VI/1959 (USNM), VI (MCZC), 1902 (CASC). 5/IX/1982 (EGRC); 1972. The leaf beetles of Alabama. Auburn Morlllos, IX (CASC). Nayarit 12.3 mi W Comllostela, 7/VII/1982 (CDFA). Oaxaca- no further data (MCZC). 2 mi N Agric. Expt. Stn. Bull. 441:1-223. • t • • n 16 1974 (TAMU); Oaxaca, 18fVII/1987; Oaxaca City, 15- Baly, J.S. Mro~~~3177~Uk~Cg g~':a:: :t~a~;~,~: J;~'?iWR:;~ 1885 Bio]ogia Centrali-Americana, Insecta, (CNC) Sinaloa- 38 mi NE Concordia, nr. LobllFaS, 3/'111/1982 Coleoptera, Hispidae. 6(2):1-124. (CDFA), SO Ion E Mazatlan, 267IVJ1974, 277IVJI9/4 (CASC); 36.6 mi NE Villa Union, nr. La Capilla Detaxte, 2 VII 1982 GUA: Chontales (USNM). Total: 218. 1940. Dejean catalogue names (Coleoptera). Bu]] Brooklyn Entomol Soc 3t'H-12. AcknOWledgments Blackwelder, R.E. 1946. Checklist oBhe eoleopterous inseets of lent materIa or t IS S U y, t €I assistance 0 t €I Mexico, Central America, the West Indies, and ~~ators responsible is gratefully acknowledged South America II S Nat Mus. Bull. 185.551- 763. University, (AUEM), W.E. Clark; British Museum of Blatchley, W.S. Natural History (BMNH), S.L. Shute; Cahforma 1910. The Coleoptera of Indiana. Ind. Dept. ~c~~m~ o;;ciences (CASC), D.H. Kavanaugh; Geol & N, R. Bull. 1:1-1386, a 1 orllla epartment of Food & Agriculture (CDFA), T.N. Seeno, Canadian National Collection Vol. 3, No. 1, March 1989 53 1924. The Chrysomelidae of Florida. Florida Davis, C.J. EntomOl. 8:39 46. 1972. Reeent introduetions for biologieal eontrol in Hawaii XVII. Proc. Hawaiian Entomol. Soc. Brimley, C.S. 21.18'7-190. 1938. The insects of North Carolina N. C Dept Agric., Dlv. Entomol. 560 pp. DaVIS, C.J., & N.L.H. Krauss 1965. Recent introductions for biOlogical control Bruner, S.C., et. al. in Hawaii- X. Proc. Hawaiian Entomol. Soc. 1975. Catalogo de los insectos que mcan a los 19.87-90. plantas economicas de Cuba. Acad. Ciencias de Cuba, Inst. Zoot Habana. 399 pp. 1966. Recent introductions for biolOgical control in Hawaii- XI. Proc. Hawaiian Entomol. Soc. Champion, G.C. 19:201-207. 1894. Biologia Centrali l~merieana, Inseeta, Dejean, P.F.M. Coleoptera (Cassididae and supplement; appen 1837. Catalogue des CoUmpteres de la collection dix to Hispinae) 6(2)'125-242 de M ]e comte Dejean 3rd ed , pp 1-503 Chapuis, F. Diatloff, G. 1875. in Lacordaire, T. Historie naturelle des 1977. Control biologico de la mala hierba Lanta- insects. Genera des CoIeopteres. Vol. 11. Fa na camUt a pOI Octotoma championi y V,oplata mille des phytophages Paris 420 p sp (cere bilineata) Agron Costar J.J 65- 167 Chevrolat, LA.A. Donckier, H. 1837. in Dejean, Catalogue des CoIeopreres de 1a 1899. Catalogue systematique des Hispides. collection de M. Ie comte Dejean. 3rd Ed., pp. 1- Ann. Entomol. France 68.540-615. 503. Dozier, H.t. Chittenden, FH 1922 An annotated list of Mississippi Chry 1902. The leaf-mInIng locust beetIe, With notes somelidae. Ohio Jour. Sci. 22:117-124. on related species. USDA Div. Entomol. Bull. 38:70-89. 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