Success with Herbaceous Perennials

Total Page:16

File Type:pdf, Size:1020Kb

Success with Herbaceous Perennials Landscape Basics: Success with Herbaceous Perennials Bodie Pennisi, Paul Thomas and Sheri Dorn Department of Horticulture hether in a commercial installation or residen- fewer pest problems, require less water and pruning, Wtial garden, perennial plants can be successfully and have an extended flowering time. This information used to offer more landscaping choices, distinguish can be acquired from books, annual trade confer- your firm from the competition and create a niche for ences and Extension publications (refer to “Additional your landscape business. Perennial plants are complex, Resources” at the end of this publication). An excellent and it is best to contract or hire a professional land- way to see how new selections perform is to visit the scape architect for the design phase and train knowl- University of Georgia Trial Gardens located on campus edgeable staff in proper maintenance later on. in Athens (http://ugatrial.hort.uga.edu/). With a rich history of plant introductions, hybridiza- This publication is intended to provide the basics of tion and selection, we can now enjoy plants in Georgia perennial plant biology, ideas on design and installa- that until recently were once found only on a remote tion, and information on cultivation and maintenance mountain slope in Asia or a tropical rainforest in South of perennial beds. It should also serve as a quick guide America. The perennial plant trade offers more than for the most common and recommended perennials 3,600 species and cultivated varieties, and more are for Georgia. Common-sense tips from a professional added each year. With such an extensive palette, how landscaper’s perspective are also included. do you choose the right plant? Our advice is to follow three simple rules: Biology of Perennials The term “perennial” in the broadest sense means 1. stick to species and selections that are known to do plants that live for more than two years. This broad well in the area; the “bread-and-butter” plants; definition includes shrubs and trees that retain woody 2. match the optimum growing requirements; plant stems above ground in winter, including ferns, cacti, the “right plant in the right place”; and succulents, grasses, bulbs, some herbs and groundcov- 3. spend the extra time to maintain each species at ers. For the home gardener, however, it means a plant its peak performance (e.g., additional nutrition, with stems that usually die back in winter and a root mid-summer trimming, regular deadheading); go system from which new foliage and flowers grow the the extra mile. following year. Most perennials have herbaceous stems (e.g., Coreopsis, Achillea, Echinacea, Liatris), but a few You don’t have to plant 50 species in one landscape, or develop woody stems (e.g., hardy hibiscus, rosemary). always use the latest and most fashionable cultivars – There is also a category of plants, called “tender peren- you could make a stunning display with five species or nials,” which are annuals that may over-winter occa- varieties that have been in the trade for 20 years. That sionally, depending on how severe the winter is (Figure said, you should strive to familiarize yourself with new 1). A true perennial has a capacity to survive from one hybrids that have improved tolerance to shade/sun, year to the next. Figure 1. Colocasia (elephant ear; front left, heart-shaped foliage), a tender perennial, is shown here emerging from the ground along with Hosta, a perennial (back of bed, gray foliage). Tender perennials have become very popular in the trade; however, you should be familiar with each spe- cies and cultivar, as their cold tolerance varies. Growing Perennials in Georgia Figure 2. Georgia Plant Hardiness Zone Map. Zones are based on average minimum winter temperatures. Many While perennial plants are found in almost every nat- other factors, such as summer heat, summer night tem- ural habitat around the world, not all plants can grow peratures, humidity and rainfall, affect plant performance well when they are cultivated in a different region of the and adaptability. globe. The best plant performance is achieved when the landscape habitat closely matches the plant’s endemic environment. Until recently, most plant catalogs listed only the cold hardiness zone when describing how far north a plant can be expected to survive (Figure 2). Georgia has a diverse range of cold hardiness zones, ranging from zone 6a in the northeast corner of the state to zone 9a in the southern coastal part of the state. Plants growing in Georgia also face a challenging com- bination of high summer temperatures and high hu- midity (Figure 3). Academicians, nurserymen, growers, and landscape professionals have found that growing Figure 3. Average days above 900 F (30-year annual aver- perennial plants in the Southeast justifies using a heat ages rounded to next highest class) in the state of Georgia. tolerance map as well -- one that describes a plant’s adaptability to summer heat. The American Horticul- to reduced vigor, small foliage and weak, stunted plants tural Society (www.ahs.org) provides information on and. The effects of heat damage are subtler than those plant heat tolerance in a detailed interactive map. of extreme cold, which can kill a plant quickly. Heat damage can first appear in many different parts of the Heat tolerance is more difficult to evaluate than cold plant -- flower buds may wither, leaves may droop or tolerance; plants respond to cold by dying, to heat by become susceptible to pests, chlorophyll may disap- slowly declining. High night temperatures increase res- pear so that leaves appear white or brown, or roots piration; for every 160F rise in temperature, respiration may cease growing. Plant death from heat is slow and rate doubles. Plants use up their stored reserves, leading lingering. UGA Extension Bulletin 1424 2 Landscape Basics: Success with Herbaceous Perennials Georgia also has a very diverse ecology, with more than tion in sandy soils causes plant stress from drought. In 100 distinct environments and at least eight major habi- fact, more perennials die from wet winter soils than from tats. These are generally grouped within three geological cold in Georgia. Amend perennial beds with organic regions: Mountains / Northwest, Piedmont and Coastal matter to improve soil conditions. Plain. Figure 3 shows that the Coastal Plains experience more days of temperatures over 900 F compared to more Research has shown that plant growth in Georgia cli- northern regions (Mountains), which plays a major role mates is typified by earlier-flowering, taller plants with in plant performance across the state. weaker stems due to accumulated heat – more so than in more northern climates. Tall forms tend to collapse with- The last two factors to consider when planting peren- out support; therefore, dwarf selections are usually more nials are rainfall and soil. The entire state, including the effective. Fertilizer may not need to be applied as gener- Mountains, receives moderate to heavy rain, which var- ously, particularly for tall plants as additional growth is ies from 45 inches in central Georgia to approximately not desired. Leggy, lanky growth and fewer flowers result 75 inches around the northeastern part of the state. The if too much nitrogen is applied. Coastal Plains experience hot, humid summers with fre- quent afternoon thunderstorms and mild, drier winters. Soil pH and Mineral Nutrients Wet, cool springs are typical. Since perennials are expected to live in a certain area for many years, it is very important to be sure the soil is Rainfall, when considered with various soils, affects amended to fit the plant’s needs. In addition to organic perennial plant survival and adaptability to Georgia’s matter, pay attention to soil pH. Too high or too low soil climate. The soils in Georgia are varied; generally, the pH may affect nutrient uptake, stress the plant and re- Coastal Plains have sandy, well-drained soils that are duce the plant’s chances of surviving more than one year. poor in organic matter, while the Piedmont and Moun- tain regions have heavier clay or clay-loam soils that Most perennials prefer a pH between 5.2 and 6.7 (slightly retain water well, with organic matter present in some acidic). Soil pH affects nutrient uptake (Figure 5). This areas (Figure 4). “Goldilocks” range assures that two important micro- nutrients -- iron (Fe) and manganese (Mn) -- are being Drainage becomes a very important factor in plant sur- absorbed by the plant roots in optimal concentrations. vival. Inadequate drainage in clay soils creates favorable If the pH is below 5, both of these elements can interfere conditions for root rots, and insufficient moisture reten- with the uptake of other nutrients. If the pH is higher than 7, Fe and Mn can become deficient. Symptoms of nutrient deficiency include yellow (chlorotic) leaf tips and young foliage (Figure 6). Figure 4. Georgia soil map (for detailed information, visit www.dot.ga.gov). Colors indicate different soil types. Figure 5. Nutrient availability chart. The red box shows the pH range in which mineral nutrient uptake is optimal. Landscape Basics: Success with Herbaceous Perennials 3 UGA Extension Bulletin 1424 sium [N-P-K]) in a 3-1-2 or 4-1-2 ratio. A 12-4-8 fertilizer, for instance, is a 3-1-2 ratio, and a 16-4-8 fertilizer is a 4-1-2 ratio. Research shows that phosphorus, the middle number in the analysis, is held by soils and does not leach with rains or irrigation as much as nitrogen or potas- sium do, so it is usually needed in lower amounts. The most common types of fertilizer ap- plied to perennial beds in the spring are general-purpose 10-10-10 (N-P-K), 8-8-8 or 16-4-8.
Recommended publications
  • Atoll Research Bulletin No. 503 the Vascular Plants Of
    ATOLL RESEARCH BULLETIN NO. 503 THE VASCULAR PLANTS OF MAJURO ATOLL, REPUBLIC OF THE MARSHALL ISLANDS BY NANCY VANDER VELDE ISSUED BY NATIONAL MUSEUM OF NATURAL HISTORY SMITHSONIAN INSTITUTION WASHINGTON, D.C., U.S.A. AUGUST 2003 Uliga Figure 1. Majuro Atoll THE VASCULAR PLANTS OF MAJURO ATOLL, REPUBLIC OF THE MARSHALL ISLANDS ABSTRACT Majuro Atoll has been a center of activity for the Marshall Islands since 1944 and is now the major population center and port of entry for the country. Previous to the accompanying study, no thorough documentation has been made of the vascular plants of Majuro Atoll. There were only reports that were either part of much larger discussions on the entire Micronesian region or the Marshall Islands as a whole, and were of a very limited scope. Previous reports by Fosberg, Sachet & Oliver (1979, 1982, 1987) presented only 115 vascular plants on Majuro Atoll. In this study, 563 vascular plants have been recorded on Majuro. INTRODUCTION The accompanying report presents a complete flora of Majuro Atoll, which has never been done before. It includes a listing of all species, notation as to origin (i.e. indigenous, aboriginal introduction, recent introduction), as well as the original range of each. The major synonyms are also listed. For almost all, English common names are presented. Marshallese names are given, where these were found, and spelled according to the current spelling system, aside from limitations in diacritic markings. A brief notation of location is given for many of the species. The entire list of 563 plants is provided to give the people a means of gaining a better understanding of the nature of the plants of Majuro Atoll.
    [Show full text]
  • Rapid in Vitro Micropropagation of Agapanthus Praecox
    View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Elsevier - Publisher Connector South African Journal of Botany 86 (2013) 46–50 Contents lists available at SciVerse ScienceDirect South African Journal of Botany journal homepage: www.elsevier.com/locate/sajb Rapid in vitro micropropagation of Agapanthus praecox Ponnusamy Baskaran, Johannes Van Staden ⁎ Research Centre for Plant Growth and Development, School of Life Sciences, University of KwaZulu-Natal Pietermaritzburg, Scottsville 3209, South Africa article info abstract Article history: In vitro micropropagation and acclimatization for the ornamental Agapanthus praecox, are reported. The influ- Received 6 March 2012 ence of different growth regulators on shoot multiplication from shoot-tip explants of A. praecox was inves- Received in revised form 19 November 2012 tigated. Prolific shoot multiplication (47.3±1.96 shoots per explant) was achieved on Murashige and Skoog Accepted 28 January 2013 (MS) medium supplemented with 22.2 μM benzyladenine (BA), 2.9 μM indole-3-acetic acid (IAA), and Available online 6 March 2013 4.5 μM thidiazuron (TDZ). Shoots were rooted on half-strength MS basal medium supplemented with μ μ Edited by NJ Taylor 5.7 M IAA and 2.5 M 2-isopentenyladenine (2iP) with 11.3±0.78 roots per shoot. The in vitro-raised plants were established successfully in a 1:1 (v/v) vermiculite:sand mixture when maintained in a greenhouse with Keywords: 100% survival. The elongated shoots (more than 5 cm in length) were treated for rooting and acclimatization Agapanthus praecox in a moistened (5.7 μM IAA and 2.5 μM 2iP) vermiculite:sand (1:1 v/v) mixture, first in the misthouse and In vitro regeneration then in the greenhouse.
    [Show full text]
  • Agapanthus Praecox Subsp. Orientalis African Lily, Blue Lily, Lily of the Nile
    Agapanthus praecox subsp. orientalis African Lily, Blue Lily, Lily of the Nile There is some controversy about the number of species of Agapanthus, depending on the source this can vary from six to ten. And in spite of the various common names (African Lily, Lily of the Nile, Blue Lily), Agapanthus isn’t a lily at all, that is, not in the family Liliaceae, nor does it come from anywhere near the Nile. In fact, all Agapanthus species come from South Africa, between the Limpopo River (‘all set about with Fever Trees’) and the Cape of Good Hope. Agapanthus praecox subsp. orientalis, the most commonly grown species, comes from the Eastern Cape and southern KwaZulu-Natal1. Amaryllidaceae – Liliaceae Ovary sits below the petals Ovary sits above the petals The genus Agapanthus has been placed in the Amaryllidaceae family which includes Daffodils, Nerines and Naked Ladies, although some consider it may sit better in its own family, the Agapanthaceae, or even with garlic and onions in the Alliaceae2. Flowers of Amaryllidaceae typically have the petals attached above the ovary; those of lilies (Liliaceae) have the petals attached below the ovary. In Sydney the blue and white flowers of Agapanthus signal the approach of summer and Christmas festivities. Agapanthus flowers are loved by some, loathed by others. In recent years, Agapanthus has yet again become a fashionable garden plant. Agapanthus sets seed readily and is an invasive weed in the Blue Mountains, on Lord Howe Island and in parts of Victoria and Western Australia. If you live near bushland and grow Agapanthus in your garden, local councils and bushland regenerators will be appreciative if you remove old flower heads before they have a chance to set seed.
    [Show full text]
  • FULL ACCOUNT FOR: Agapanthus Praecox Global Invasive Species Database (GISD) 2021. Species Profile Agapanthus Praecox. Available
    FULL ACCOUNT FOR: Agapanthus praecox Agapanthus praecox System: Terrestrial Kingdom Phylum Class Order Family Plantae Magnoliophyta Liliopsida Liliales Liliaceae Common name African-lily (English), lirio africano (Spanish, Argentina), agapanthus (English), bloulelie (Afrikaans), lily-of-the-Nile (English), agapanto (Spanish, Argentina), common agapanthus (English), flor de Navidad (Spanish, Argentina), blue lily (English) Synonym Tulbaghia praecox , (Willd.) Kuntze Similar species Agapanthus africanus, Arthropodium cirratum Summary Agapanthus praecox is a rhizomatous herb that is native to South Africa, along with its subspecies, A. praecox subsp. minimus, A. praecox subsp. orientalis and A. praecox subsp. praecox. Most cultivated agapanthus are cultivars or hybrids of A. praecox, and the subspecies readily hybridise. Agapanthus has characteristic composite florescences that are white or blue and tubular and is commonly planted as an ornamental. view this species on IUCN Red List Species Description Agapanthus praecox is a rhizomatous, perennial herb. Its leaves are robust, strap-like and evergreen and there are 6 - 20 leaves per individual plant. The leaves grow in dense clumps from bulb up to 60 cm high. The composite inflorescences are large and round, made up of tubular flowers. These are either coloured white or light blue. Inflorescences can grow up to 1.2 m in summer. (ARC 2009; FloraBase 2010; Notten 2004; Weeds of Blue Mountains Bushland 2010). \n Notes Known subspecies of Agapanthus praecox include A. praecox ssp. minimus, A. praecox ssp. orientalis and A. praecox ssp. praecox. Subspecies readily hybridise, especially when grown in close proximity. In the Auckland region of New Zealand, the sale, propogation, planting and distribution of A. praecox is prohibited.
    [Show full text]
  • The Naturalized Vascular Plants of Western Australia 1
    12 Plant Protection Quarterly Vol.19(1) 2004 Distribution in IBRA Regions Western Australia is divided into 26 The naturalized vascular plants of Western Australia natural regions (Figure 1) that are used for 1: Checklist, environmental weeds and distribution in bioregional planning. Weeds are unevenly distributed in these regions, generally IBRA regions those with the greatest amount of land disturbance and population have the high- Greg Keighery and Vanda Longman, Department of Conservation and Land est number of weeds (Table 4). For exam- Management, WA Wildlife Research Centre, PO Box 51, Wanneroo, Western ple in the tropical Kimberley, VB, which Australia 6946, Australia. contains the Ord irrigation area, the major cropping area, has the greatest number of weeds. However, the ‘weediest regions’ are the Swan Coastal Plain (801) and the Abstract naturalized, but are no longer considered adjacent Jarrah Forest (705) which contain There are 1233 naturalized vascular plant naturalized and those taxa recorded as the capital Perth, several other large towns taxa recorded for Western Australia, com- garden escapes. and most of the intensive horticulture of posed of 12 Ferns, 15 Gymnosperms, 345 A second paper will rank the impor- the State. Monocotyledons and 861 Dicotyledons. tance of environmental weeds in each Most of the desert has low numbers of Of these, 677 taxa (55%) are environmen- IBRA region. weeds, ranging from five recorded for the tal weeds, recorded from natural bush- Gibson Desert to 135 for the Carnarvon land areas. Another 94 taxa are listed as Results (containing the horticultural centre of semi-naturalized garden escapes. Most Total naturalized flora Carnarvon).
    [Show full text]
  • Inventory of Vascular Plants of the Kahuku Addition, Hawai'i
    CORE Metadata, citation and similar papers at core.ac.uk Provided by ScholarSpace at University of Hawai'i at Manoa PACIFIC COOPERATIVE STUDIES UNIT UNIVERSITY OF HAWAI`I AT MĀNOA David C. Duffy, Unit Leader Department of Botany 3190 Maile Way, St. John #408 Honolulu, Hawai’i 96822 Technical Report 157 INVENTORY OF VASCULAR PLANTS OF THE KAHUKU ADDITION, HAWAI`I VOLCANOES NATIONAL PARK June 2008 David M. Benitez1, Thomas Belfield1, Rhonda Loh2, Linda Pratt3 and Andrew D. Christie1 1 Pacific Cooperative Studies Unit (University of Hawai`i at Mānoa), Hawai`i Volcanoes National Park, Resources Management Division, PO Box 52, Hawai`i National Park, HI 96718 2 National Park Service, Hawai`i Volcanoes National Park, Resources Management Division, PO Box 52, Hawai`i National Park, HI 96718 3 U.S. Geological Survey, Pacific Island Ecosystems Research Center, PO Box 44, Hawai`i National Park, HI 96718 TABLE OF CONTENTS ABSTRACT.......................................................................................................................1 INTRODUCTION...............................................................................................................1 THE SURVEY AREA ........................................................................................................2 Recent History- Ranching and Resource Extraction .....................................................3 Recent History- Introduced Ungulates...........................................................................4 Climate ..........................................................................................................................4
    [Show full text]
  • Lindfield Substation Review of Environmental Factors Technical
    Transport for NSW Ecological Assessment for proposed substation at Lindfield 22 July 2014 Document information Client: Transport for NSW Title: Ecological Assessment for proposed substation at Lindfield Document No: 2176502A-ECO-RPT-001 Date: 22 July 2014 Rev Date Details 18/12/2013 First Draft A 16/04/2014 Second Draft B 09/07/2014 Amendments of draft for survey of additional proposed works C 22/07/2014 Amendments following TfNSW review Author, Reviewer and Approver details Prepared by: Tanya Bangel Date: 22/07/2014 Signature: Reviewed by: Alex Cockerill Date: 22/07/2014 Signature: Approved by: Paul Greenhalgh Date: 22/07/2014 Signature: Distribution Transport for NSW, Parsons Brinckerhoff file, Parsons Brinckerhoff Library ©Parsons Brinckerhoff Australia Pty Limited 2013 Copyright in the drawings, information and data recorded in this document (the information) is the property of Parsons Brinckerhoff. This document and the information are solely for the use of the authorised recipient and this document may not be used, copied or reproduced in whole or part for any purpose other than that for which it was supplied by Parsons Brinckerhoff. Parsons Brinckerhoff makes no representation, undertakes no duty and accepts no responsibility to any third party who may use or rely upon this document or the information. Document owner Parsons Brinckerhoff Australia Pty Limited ABN 80 078 004 798 Level 27 Ernst & Young Centre 680 George Street, Sydney NSW 2000 GPO Box 5394 Sydney NSW 2001 Australia Tel: +61 2 9272 5100 Fax: +61 2 9272 5101 Email: [email protected] www.pbworld.com Certified to ISO 9001, ISO 14001, AS/NZS 4801 A GRI Rating: Sustainability Report 2011 Transport for NSW Ecological Assessment for proposed substation at Lindfield Contents Page number Executive summary v 1.
    [Show full text]
  • BAWSCA Turf Replacement Program Plant List Page 1 Species Or
    BAWSCA Turf Replacement Program Plant List Page 1 Species or Cultivar Common name Irrigation Irrigation (1) Requirement Type (2) Native Coastal Peninsula Bay East Salinity (3) Tolerance Abutilon palmeri INDIAN MALLOW 1 S √ √ √ √ Acer buergerianum TRIDENT MAPLE 2 T √ H Acer buergerianum var. formosanum TRIDENT MAPLE 2 T √ Acer circinatum VINE MAPLE 2 S √ √ √ √ Acer macrophyllum BIG LEAF MAPLE 2 T √ √ L Acer negundo var. californicum BOX ELDER 2 T √ √ Achillea clavennae SILVERY YARROW 1 P √ √ √ M Achillea millefolium COMMON YARROW 1 P √ √ √ M Achillea millefolium 'Borealis' COMMON YARROW 1 P √ √ √ M Achillea millefolium 'Colorado' COMMON YARROW 1 P √ √ √ M Achillea millefolium 'Paprika' COMMON YARROW 1 P √ √ √ M Achillea millefolium 'Red Beauty' COMMON YARROW 1 P √ √ √ M Achillea millefolium 'Summer Pastels' COMMON YARROW 1 P √ √ √ M Achillea 'Salmon Beauty' 1 P √ √ √ M Achillea taygetea 1 P √ √ √ Achillea 'Terracotta' 1 P √ √ √ Achillea tomentosa 'King George' WOLLY YARROW 1 P √ √ √ Achillea tomentosa 'Maynard's Gold' WOLLY YARROW 1 P √ √ √ Achillea x kellereri 1 P √ √ √ Achnatherum hymenoides INDIAN RICEGRASS 1 P √ √ √ √ Adenanthos sericeus WOOLYBUSH 1 S √ √ √ Adenostoma fasciculatum CHAMISE 1 S √ √ √ √ Adenostoma fasciculatum 'Black Diamond' CHAMISE 1 S √ √ √ √ Key (1) 1=Least 2=Intermediate 3=Most (2) P=Perennial; S=Shrub; T=Tree (3) L=Low; M=Medium; H=High 1/31/2012 BAWSCA Turf Replacement Program Plant List Page 2 Species or Cultivar Common name Irrigation Irrigation (1) Requirement Type (2) Native Coastal Peninsula Bay East Salinity (3) Tolerance Adenostoma fasciculatum 'Santa Cruz Island' CHAMISE 1 S √ √ √ √ Adiantum jordnaii CALIFORNIA MAIDENHAIR 1 P √ √ √ √ FIVE -FINGER FERN, WESTERN Adiantum pedatum MAIDENHAIR 2 P √ √ √ √ FIVE -FINGER FERN, WESTERN Adiantum pedatum var.
    [Show full text]
  • WUCOLS List S Abelia Chinensis Chinese Abelia M ? ? M / / Copyright © UC Regents, Davis Campus
    Ba Bu G Gc P Pm S Su T V N Botanical Name Common Name 1 2 3 4 5 6 Symbol Vegetation Used in Type WUCOLS List S Abelia chinensis Chinese abelia M ? ? M / / Copyright © UC Regents, Davis campus. All rights reserved. bamboo Ba S Abelia floribunda Mexican abelia M ? M M / / S Abelia mosanensis 'Fragrant Abelia' fragrant abelia ? ? ? ? ? ? bulb Bu S Abelia parvifolia (A. longituba) Schuman abelia ? ? ? M ? ? grass G groundcover GC Gc S Abelia x grandiflora and cvs. glossy abelia M M M M M / perennial* P S Abeliophyllum distichum forsythia M M ? ? ? ? palm and cycad Pm S Abelmoschus manihot (Hibiscus manihot) sunset muskmallow ? ? ? L ? ? T Abies pinsapo Spanish fir L L L / / / shrub S succulent Su T N Abies spp. (CA native and non-native) fir M M M M / / P N Abronia latifolia yellow sand verbena VL VL VL / ? ? tree T P N Abronia maritima sand verbena VL VL VL / ? ? vine V California N native S N Abutilon palmeri Indian mallow L L L L M M S Abutilon pictum thompsonii variegated Chinese lantern M H M M ? ? Sunset WUCOLS CIMIS ET Representative Number climate 0 Region zones** Cities zones* S Abutilon vitifolium flowering maple M M M / ? ? Healdsburg, Napa, North- San Jose, Salinas, Central 14, 15, 16, 17 1, 2, 3, 4, 6, 8 San Francisco, Coastal San Luis Obispo S Abutilon x hybridum & cvs. flowering maple M H M M / / 1 Auburn, Central Bakersfield, Chico, 8, 9, 14 12, 14, 15, 16 Valley Fresno, Modesto, Sacramento S T Acacia abyssinica Abyssinian acacia / ? / ? / L 2 Irvine, Los South Angeles, Santa 22, 23, 24 1, 2, 4, 6 Coastal Barbara, Ventura,
    [Show full text]
  • Class B Gene Expression and the Modified ABC Model in Nongrass Monocots
    Review Article Special Issue: Evolution of MADS-box genes in Monocots TheScientificWorldJOURNAL (2007) 7, 268–279 TSW Development & Embryology ISSN 1537-744X; DOI 10.1100/tsw.2007.86 Class B Gene Expression and the Modified ABC Model in Nongrass Monocots Akira Kanno*, Mutsumi Nakada, Yusuke Akita, and Masayo Hirai Graduate School of Life Sciences, Tohoku University, Katahira 2-1-1, Aoba-ku, Sendai 980-8577, Japan E-mail: [email protected] Received August 9, 2006; Revised December 28, 2006; Accepted January 4, 2007; Published February 19, 2007 The discovery of the MADS-box genes and the study of model plants such as Arabidopsis thaliana and Antirrhinum majus have greatly improved our understanding of the molecular mechanisms driving the diversity in floral development. The class B genes, which belong to the MADS-box gene family, are important regulators of the development of petals and stamens in flowering plants. Many nongrass monocot flowers have two whorls of petaloid organs, which are called tepals. To explain this floral morphology, the modified ABC model was proposed. This model was exemplified by the tulip, in which expansion and restriction of class B gene expression is linked to the transition of floral morphologies in whorl 1. The expression patterns of class B genes from many monocot species nicely fit this model; however, those from some species, such as asparagus, do not. In this review, we summarize the relationship between class B gene expression and floral morphology in nongrass monocots, such as Liliales (Liliaceae) and Asparagales species, and discuss the applicability of the modified ABC model to monocot flowers.
    [Show full text]
  • PLANTS Agapanthus (Agapanthus Praecox) an Attractive Blue Or White Flowered Lily That Forms Robust Clumps with Long, Strap-Shape
    PLANTS Agapanthus (Agapanthus praecox) An attractive blue or white flowered lily that forms robust clumps with long, strap-shaped perennial leaves. They are long-lived prolific seeders, very tolerant to temperature, rainfall, salt, poor soils and can survive immersion in the sea. Agapanthus is particularly invasive in the coastal environment outcompeting native coastal plants. Aluminium plant (Lamium galeobdolon ‘Variegatum’) Sometimes known as Artillery plant, Aluminium plant is an erect, stoloniferous, mat-forming, invasive perennial herb, up to 50 cm tall. The leaves are mint-like with silvery-grey patches on the upper surface, seldom reverting to a plain green-leaved form. Often found in gardens growing in the cooler areas around trees and shrubs. From December to May tubular, hairy, lemon-yellow flowers (20-25 mm long) are produced in dense axillary whorls, but no seed is set. They grow rapidly and cover large areas of ground with a thick mat that stops seedlings of other plants from establishing. It is shade tolerant so can be an issue deep into the bush as well as along margins. PLANTS Arum lily (Zantedeschia aethiopica) A robust, evergreen, clump-forming perennial up to 1.5 m tall, with large, leathery, dark green, arrow-shaped leaves. Large, white flower with a yellow spike and yellow-green berries. Long-lived and persists under regenerating canopy, forming dense patches excluding other vegetation. Tolerates excessive water, wind, salt, hot to cold, most soil types, moderate shade, and is drought-resistant once established. Stock avoid it as it is poisonous, allowing it to gradually dominate grazed sites. Seeds drop near to parent plants, and are occasionally spread by birds and water.
    [Show full text]
  • Verslag Van Een KNNV-Reis Door ZW-Australië
    Verslag KNNV-reis Azoren, aug. 2014 Verslag van KNNV-reis naar De Azoren, aug. 2014 1 Verslag KNNV-reis Azoren, aug. 2014 Colofon Het rapport “Verslag van KNNV-reis naar De Azoren, Het rapport is als PDF-bestand aan te vragen bij Ad. aug. 2014” is door Ad. Littel in eigen beheer uitgege- Littel, e-mail: [email protected]. ven in okt. 2014 te Leiden (NL). Aan dit rapport is door alle deelnemers aan deze reis een bijdrage geleverd; dit is bij de betreffende delen aangegeven. Het rapport in zijn geheel is samengesteld en geredi- geerd door Ad. Littel. Wandeling naar Ribeiro Quente op São Miguel (foto Gea Warringa) Op voorblad zijn afgebeeld: landschap bij Capelinho op Faial, De inzet betreft 2 Azorenendemen: Azorina vidalii (links) en Vaccinium cylindraceum (rechts) 1 Verslag KNNV-reis Azoren, aug. 2014 Verslag van een KNNV-reis naar enkele eilanden van De Azoren (Portugal) Algemene inleiding De Algemene ReisCommissie van de Koninklijke Nederlandse Natuurhistorische Vereniging (ARC-KNNV) organiseerde en begeleidde in augustus 2014 een natuurreis naar enkele eilanden van De Azoren (NB: het lidwoord ‘de’ is onderdeel van de eigennaam). Het programma kondigde aandacht aan voor planten en geologie. De flora van deze tot de staat Portugal behorende maar ver in de Atlantische Oceaan gelegen eilandengroep is belangwekkend vanwege een groot aantal endemische soorten en de aanwezigheid van bijzondere exotische soorten die hier door de zeer zachte winters en vochtige zomers goed kunnen overleven. De geologie is op veel plaatsen zichtbaar aanwezig in de vorm van allerlei restanten van vulkanische activiteiten. De eilanden zijn geheel van vulkanische oorsprong.
    [Show full text]