The GREAT LAKES ENTOMOLOGIST Vol. 11, No. 3 Fall 1978 THE GREAT LAKES ENTOMOLOGIST Published by the Michigan Entomological Society Volume 11 No. 3 TABLE OF CONTENTS Baetis (Ephemeroptera: Baetidae) of Wisconsin Edward A. Ber-man and William L. Hilsenhoff. ................... 125 A Natural Subgeneric Classification of Ephemerelh bartoni and Related Species (Ephemeroptera: Ephemerellidae) W. P. SlcCafferty .................................... .I37 Annotated List of Stoneflies ~Plecoptera)from Stebbins Gulch in Northeastern Ohio Martin A. Tkac and B. A. Foote ............................ 139 Annotated List of Caddistlies tTrichoptera) Occurring along the Upper Portion of the West Branch of the Mahoning River in Northeastern Ohio Eric P. McElra~~and B. .L Foote ........................... 143 Effects of Various Phoropsiids on Color and Pubescence in Thyanta calceata (Hemiptera: Penratomidae:~ J. E. McPherson. .................................... .I55 A List of the Scutellerojdea IHemiptera) of Southern Illinois J. E. YcPherwn ..................................... ,159 Neu- World Limnichinae IV. Eutimnichus Casey. A. Synonymies, Lectotype Designations. and Redexfiptions icoleoptera: Limnichidae) David P. U-ooldridge ................................... 163 Behavior at a 14;esting Site adPrey of Crabro cribrellifer (Hymenoptera: Sphecidae) Edward M. Banoa-s. PamiLG L. Lebau. and Cathlyn E. Eckstein ......... 175 CO\ER ILLUSTRATION Dorsal and ventral iien- of Pelrod~ressexmacuhtus Roberts (Coleoptera: Haliplidae). Drawing by Ms. Peninnah Sni-A. Illinoh yamral History Society. Vol. 11, No. 2 of The Great U-es Entomologist was mailed on 11 July, 1978. THE MICHIGAN ENTOMOLOGICAL SOCIETY 1977-76 OFFICERS President Roger G. Bland President-Elect ~anielK. Young Executive Secretary M. C. Nielsen Editor D. C. L. Gosling The Michigan Entomological Society traces its origins to the old Detroit Entomo- logical Society and was organized on 4 November 1954 to ". promote the science of entomology in all its branches and by all feasible means, and to advance cooperation and good fellowship among persons interested in entomology." The Society attempts to facilitate the exchange of ideas and information in both amateur and professional circles, and encourages the study of insects by youth. Membership in the Society, which serves the North Central States and adjacent Canada, is open to all persons interested in entomology. There are four paying classes of membership: Student (including those currently enrolled in college or graduate programs)-annual dues $2.00 Active-annual dues $4.00 Institutional-annual dues $10.00 Sustaining-annual contribution $25.00 or more Dues are paid on a calendar year basis (Jan. 1-Dec. 31). Memberships accepted before July 1 shall begin on the preceding January 1; member- ships accepted at a later date shall begin the following January 1 unless the earlier date is requested and the required dues are paid. All members in good standing receive the Newsletter of the Society, published quarterly. All Active and Sustaining Members may vote in Society affairs. All dues and contributions to the Society are deductible for Federal income tax purposes. SUBSCRIPTION INFORMATION Institutions and organizations, as well as individuals not desiring the benefits of membership, may subscribe to The Great Lakes Entomologist at the rate of $10.00 per volume. The journal is published quarterly; subscriptions are accepted only on a volume (4 issue) basis. Single copies of The Great Lakes Entomologist are available at $2.75 each, with a 20 per cent discount for 25 or more copies sent to a single address. MICROFILM EDITION: Positive microfilm copies of the current volume of The Great Lakes Entomologist will be available at nominal cost, to members and bona fide sub- scribers of the paper edition only, at the end of each volume year. Please address all orders and inquiries to University Microfilms, Inc., 300 North Zeeb Road, Ann Arbor, Michigan 48106, USA. Inquiries about back numbers, subscriptions and Society business should be directed to the Executive Secretary, Michigan Entomological Society, Department of Entomology, Michigan State University, East Lansing, Michigan 48824, USA. Manuscripts and related correspondence should be directed to the Editor (see inside back cover). Copyright @ 1978, The Michigan Entomological Society THE GREAT LAKES ENTOMOLOGIST BAETlS (EPHEMEROPTERA: BAETIDAE) OF WISCONSIN^ Edward A. Bergman and William L. ~ilsenhoff~ Data on life histories and environmental requirements for species in many mayfly genera remains sketchy at best This is certainly true of Baetis, which is one of the most common components of Wisconsin's lotic fauna. Most Wisconsin streams that are not grossly pobted contain one or more species of the minnow-like nymphs, which are usually found clinging to surfaces of rocks or aquatic plants. Biological studies of Baetis in North America have been neglected primarily because of their enigmatic taxonomy. Even keys of Needham et al. (1935) and Burks (1953), which are considered standard referenas, are either incomplete or difficult to use when identifying Baetis. Before a study of the biology of Baetis in Wisconsin could be completed, a study of their taxonomy was necessary, and numerous unexpected problems were encountered. Most were the occurrence of maleless parthenogenetic species, seasonal size and mlor variations in male imagoes of several species, difficulties in associating reared specimens with existing descriptions, and physical deterioration of important nymphal specimens collected by early taxonomists. Despite these problems, we collected valuable data on Baetis and this paper reports our knowledge of Baetis in Wisconsin today. In Wxonsin, 14 species of Baetis have been collected. Adult males and mature nymphs can be separated using keys that follow. Supplemental characters useful in identification and synonomies are presented in a discussion of each species, and developmental cycles, ecology, and known distribution of Wisconsin Baetis are sum- marized in Table 1. KEY TO WISCONSIN BAETZS-ADULT MALES~ Hind wing with 3 longitudinal veins and with a costal projection near base. .. 2 Hind wing with 2 longitudinal veins, with or without a costal projection .... 3 Second vein of hind wing simple. ........................... 6 Second vein of hind wing forked (males and females). ............ hageni Costal projection of hind wing absent; first segment of genital forceps with a prominent apicomesal tubercle (Fig. 1B) .................propinquus Costal projection present, although sometimes reduced .............. 4 A spine between bases of genital forceps (Figs. lD,E); costal projection reduced ............................................... 5 No spine between bases of genital forceps; first segment of forceps approxi- mately 113 as long as second, third segment about 2-112 times longer than wide (Fig 1C); costal projection relatively well developed ........... pygmaeus First segment of genital forceps with a prominent, pointed mesa1 projection (Fig. ID) ......................................... spinosus First segment of genital forceps cylindrical (Fig. 1E) ............frondalis First segment of genital forceps cylindrical (Fig. IF), no tubercle on inner apical margin of basal segment. ............................. vagans First segment of genital forceps conical (Fig. IG, H, I, J), a tubercle present on inner apical margin of basal segment ......................... 7 Tubercle on apical margin of basal segment of genital forceps poorly developed (Fig 1G); eyes (live) lemon yellow .......................levitans l~esearchsupported by the College of Agricultural and Life Sciences, University of Wficonsin. Madison, and by a grant from the Department of Natural Resources. l~e~artrnentof Entomology, University of Wisconsin, Madison, WI 53706 3~ale mocdunnarghi unknown, females difficult to separate from female B. pygmaeus and not included in this key. 126 THE GREAT LAKES ENTOMOLOGIST Vol. 11, No. 3 1978 THE GREAT LAKES ENTOMOLOGIST 127 7'. Tubercle on apical margin of basal segment of genital forceps well developed . 8 8(7'). Abdominal terga 2-6 brown (amount variable) ..............brunneicolor 8'. Abdominal terga 2-6 hyaline-white or yellowish-white, often with a narrow reddish-range transverse stripe (variable). ...................... 9 9(8'). Thorax shiny black; abdominal terga 2-6 hyaline-white; turbinate eyes reddish- brown (live) ...................................intercalaris 9'. Thorax olive-brown, never shiny black; abdominal terga 2-6 yellow-white, often with a reddish-range transverse stripe (intensity of stripe variable); turbinate eyes reddish-orange (live) ............................phoebus KEY TO WISCONSIN BAETIS-MATURE NYMPHS 1. Two tails. ...................................Baetis sp. B 1'. Threetails ......................................... 2 (1. Tails without crossbands at or near middle. ..................... 3 2'. Tails with dark crossbands at or near middle. .................... 6 3(2). Tails without crossbands. ................................4 3'. Tails with white or dark apical crossbands ...................... 5 4(3). Abdominal terga brown, often with a pale median stripe (Fig. 2A); abdominal terga 10 and 5 sometimes paler than other terga ........... .brunneicolor 4'. Abdominal terga without a pale median stripe; abdominal tergum 9 paler than other terga, 5 also usually pale (Fig. 2B) ....................vagans 5(3'). Tails tan, with a dark brown apical band on cerci. .............