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Winter Blooming Shrubs by RICHARD E
Winter Blooming Shrubs by RICHARD E. WEAVER, JR. Winters in the eastern part of this country south of Washington, D.C. are seldom as unpleasant as they are here in the Northeast. Of course the temperatures there are less extreme, but for those of us who appreciate plants and flowers, the real difference is perhaps due to the Camellias. Blooming through the worst weather that January and February have to offer, these wonderful plants with their bright and showy blooms make winter something almost worth anticipating. Although there are some hopeful new developments through con- centrated breeding efforts, we in most of the Northeast still must do without Camellias in our gardens. Nevertheless, there are a sur- prising number of hardy shrubs, perhaps less showy but still charm- ing and attractive, that will bloom for us through the winter and the early days of spring. Some, such as the Witch Hazels, are foolproof; others present a challenge for they are susceptible to our capricious winters and may lose their opening flowers to a cold March. For those gardeners willing to take the chance, a few of the best early- flowering shrubs displayed in the border, or as the focal point in a winter garden, will help to soften the harshness of the season. Many plants that bloom in the early spring have their flowers per- fectly formed by the previous fall. Certain of these do not require a period of cold dormancy, and in mild climates will flower intermit- tently during the fall and winter. Most species, however, do require an environmental stimulus, usually a period of cold temperatures, before the buds will break and the flowers open. -
Wood Anatomy of Calycanthaceae Sherwin Carlquist
Aliso: A Journal of Systematic and Evolutionary Botany Volume 10 | Issue 3 Article 6 1983 Wood Anatomy of Calycanthaceae Sherwin Carlquist Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons Recommended Citation Carlquist, Sherwin (1983) "Wood Anatomy of Calycanthaceae," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 10: Iss. 3, Article 6. Available at: http://scholarship.claremont.edu/aliso/vol10/iss3/6 ALISO 10(3), 1983, pp. 427-441 WOOD ANATOMY OF CALYCANTHACEAE: ECOLOGICAL AND SYSTEMATIC IMPLICATIONS Sherwin Carlquist INTRODUCTION Wood anatomy of Calycanthaceae has not been studied as a unit. Wood features ofthe family have been summarized by Metcalfe and Chalk (1950); various authors have mentioned one or more traits in studies dealing with Calycanthaceae (e.g., Wilson 1979) or other families (e.g., Garratt 1934). In view of recent interest in Idiospermum australiense (Diels) Blake, a new comparative study is needed. One goal of the present study is clarification of relationships of Idiospermum to Calycanthus and Chimonanthus. Wood anatomy of Idiospermum was described by Blake ( 1972) and Wilson ( 1979); a new description is offered here to provide more quantitative data. De scriptions of the wood of Calycanthus and Chimonanthus provided here incorporate such quantitative data, but also modify earlier descriptions with respect to some important qualitative features. Material of the recently de scribed genus Sinocalycanthus (Cheng and Chan 1964) was not available, although the description of that genus suggests it is not strongly different from Calycanthus or Chimonanthus. The present study incorporates material of Calycanthus floridus L. var. floridus, C. -
Forest Structures, Composition, and Distribution on a Pacific Island, with Reference to Ecological Release and Speciation!
Pacific Science (1991), vol. 45, no. 1: 28-49 © 1991 by University of Hawaii Press. All rights reserved Forest Structures, Composition, and Distribution on a Pacific Island, with Reference to Ecological Release and Speciation! YOSHIKAZU SHIMIZU2 AND HIDEO TABATA 3 ABSTRACT: Native forest and scrub of Chichijima, the largest island in the Bonins, were classified into five types based on structural features: Elaeocarpus Ardisia mesic forest, 13-16 m high, dominated by Elaeocarpus photiniaefolius and Ardisia sieboldii; Pinus-Schima mesic forest, 12-16 m high, consisting of Schima mertensiana and an introduced pine , Pinus lutchuensis; Rhaphiolepis Livistonia dry forest, 2-6 m high, mainly occupied by Rhaphiolepis indica v. integerrima; Distylium-Schima dry forest, 3-8 m high, dominated by Distylium lepidotum and Schima mertensiana; and Distylium-Pouteria dry scrub, 0.3 1.5 m high, mainly composed of Distylium lepidotum. A vegetation map based on this classification was developed. Species composition and structural features of each type were analyzed in terms of habitat condition and mechanisms of regeneration. A group of species such as Pouteria obovata, Syzgygium buxifo lium, Hibiscus glaber, Rhaphiolepis indica v. integerrima, and Pandanus boninen sis, all with different growth forms from large trees to stunted shrubs, was subdominant in all vegetation types. Schima mertensiana , an endemic pioneer tree, occurred in both secondary forests and climax forests as a dominant canopy species and may be an indication of "ecological release," a characteristic of oceanic islands with poor floras and little competitive pressure. Some taxonomic groups (Callicarpa, Symplocos, Pittosporum, etc.) have speciated in the under story of Distylium-Schima dry forest and Distylium-Pouteria dry scrub. -
Pollination-Induced Gene Changes That Lead to Senescence in Petunia × Hybrida
Pollination-Induced Gene Changes That Lead to Senescence in Petunia × hybrida DISSERTATION Presented in Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy in the Graduate School of The Ohio State University By Shaun Robert Broderick, M.S. Graduate Program in Horticulture and Crop Science The Ohio State University 2014 Dissertation Committee: Michelle L. Jones, Advisor Feng Qu Eric J. Stockinger Esther van der Knaap Copyrighted by Shaun Robert Broderick 2014 Abstract Flower longevity is a genetically programmed event that ends in flower senescence. Flowers can last from several hours to several months, based on flower type and environmental factors. For many flowers, particularly those that are ethylene- sensitive, longevity is greatly reduced after pollination. Cellular components are disassembled and nutrients are remobilized during senescence, which reduces the net energy expenditures of floral structures. The goal of this research is to identify the genes that can be targeted to extent shelf life by inhibiting pollination-induced senescence. Identifying and characterizing regulatory shelf-life genes will enable breeders to incorporate specific alleles that improve post production quality into ethylene-sensitive crops. Petunia × hybrida is particularly amenable to flower longevity studies because of its large floral organs, predictable flower senescence timing, and importance in the greenhouse industry. A general approach to gene functional analysis involves reducing gene expression and observing the resulting phenotype. Viruses, such as tobacco rattle virus (TRV), can be used to induce gene silencing in plants like petunia. We optimized several parameters that improved virus-induced gene silencing (VIGS) in petunia by increasing the consistency and efficiency of silencing. -
Chimonanthus Praecox
Shang et al. Genome Biology (2020) 21:200 https://doi.org/10.1186/s13059-020-02088-y RESEARCH Open Access The chromosome-level wintersweet (Chimonanthus praecox) genome provides insights into floral scent biosynthesis and flowering in winter Junzhong Shang1†, Jingpu Tian1†, Huihui Cheng2†, Qiaomu Yan1, Lai Li1, Abbas Jamal1, Zhongping Xu3,4, Lin Xiang1, Christopher A. Saski5, Shuangxia Jin3,4* , Kaige Zhao1*, Xiuqun Liu1* and Longqing Chen6* * Correspondence: [email protected]. edu.cn; [email protected]; Abstract [email protected]; clqhzau@126. com Background: Wintersweet (Chimonanthus praecox), an important ornamental plant, †Junzhong Shang, Jingpu Tian and has evolved unique fragrant aroma and winter-flowering properties, which are critical Huihui Cheng contributed equally for its successful sexual reproduction. However, the molecular mechanisms underlying to this work. 3National Key Laboratory of Crop these traits are largely unknown in this species. In addition, wintersweet is also a typical Genetic Improvement, Huazhong representative species of the magnoliids, where the phylogenetic position of which Agricultural University, Wuhan, relative to eudicots and monocots has not been conclusively resolved. Hubei 430070, People’s Republic of China Results: Here, we present a chromosome-level wintersweet genome assembly with a 1Key Laboratory of Horticultural total size of 695.36 Mb and a draft genome assembly of Calycanthus chinensis. Plant Biology, Ministry of Education, Huazhong Agricultural University, Phylogenetic analyses of 17 representative angiosperm genomes suggest that Wuhan, Hubei 430070, People’s Magnoliids and eudicots are sister to monocots. Whole-genome duplication Republic of China signatures reveal two major duplication events in the evolutionary history of the 6Southwest Engineering Technology and Research Center of wintersweet genome, with an ancient one shared by Laurales, and a more recent Landscape Architecture, State one shared by the Calycantaceae. -
Establishment of in Vitro Plant Regeneration System for Chimonanthus Praecox (L.)
African Journal of Biotechnology Vol. 11(45), pp. 10358-10361, 5 June, 2012 Available online at http://www.academicjournals.org/AJB DOI:10.5897/AJB11.4278 ISSN 1684–5315 ©2012 Academic Journals Full Length Research Paper Establishment of in vitro plant regeneration system for Chimonanthus praecox (L.) Mingxiao Zhao, Guoqiang Fan* and Xiaoshen Zhang College of Forestry, Henan Agricultural University, Zhengzhou 450002, China. Accepted 10 May, 2012 An efficient protocol for plant regeneration of Chimonanthus praecox (L.) Link, was developed using leaves from seedlings of seeds. Shoots induction were influenced by cytokinins and nodal positions. The results show that the highest callus induction frequency was obtained on MS medium supplemented with 2.0 mg l−1 6-benzyladenine (6-BA) and 0.9 mg l−1 1-naphthalene-acetic acid (NAA) after 10 days incubation in darkness. The frequency of callus induction of different nodal leaves (from the first to the forth node) varied from 70.0 to 93.5%; the highest frequency of callus induction (93.5%) was observed in the second node group. Through many times subculture of callus, green and compact callus was selected and transferred to regeneration medium (MS, 1/2 MS and woody plant medium (WPM) supplemented with different concentrations of thidiazuron (TDZ) (2.37, 4.74 and 9.48 mg l−1) and NAA (0.74, 1.48, and 2.96 mg l−1). It was shown that the optimal combination of the three factors was WPM + NAA + TDZ. The highest adventitious shoot regeneration frequency of leaf explants (71.33%) and the highest mean number of shoots per explant (2.23) were obtained on WPM supplemented with 2.37 mg l−1 TDZ and 1.48 mg l−1 NAA. -
Analysis of Floral Fragrance Compounds of Chimonanthus Praecox with Different Floral Colors in Yunnan, China
separations Article Analysis of Floral Fragrance Compounds of Chimonanthus praecox with Different Floral Colors in Yunnan, China Liubei Meng 1,†, Rui Shi 2,† , Qiong Wang 1,* and Shu Wang 1,2,* 1 College of Landscape Architecture and Horticulture, Southwest Forestry University, Kunming 650224, China; [email protected] 2 Southwest Landscape Architecture Engineering Research Center of National Forestry and Grassland Administration, Kunming 650224, China; [email protected] * Correspondence: [email protected] (Q.W.); [email protected] (S.W.) † These authors contributed equally to this work and should be considered as co-first authors. Abstract: In order to better understand the floral fragrance compounds of Chimonanthus praecox belonging to genus Chimonanthus of Chimonanaceae in Yunnan, headspace solid-phase microextraction combined with gas chromatography-mass spectrometry was used to analyze these compounds from four C. praecox plants with different floral colors. Thirty-one types of floral fragrance compounds were identified, among which terpenes, alcohols, esters, phenols, and heterocyclic compounds were the main compounds. Interestingly, the floral fragrance compounds identified in the flowers of C. praecox var. concolor included benzyl acetate, α-ocimene, eugenol, indole, and benzyl alcohol. By contrast, the floral fragrance compounds β-ocimene, α-ocimene, and trans-β-ocimene were detected in C. praecox var. patens. Cluster analysis showed that C. praecox var. concolor H1, H2, and C. praecox var. patens H4 were clustered in one group, but C. praecox var. patens H3 was individually clustered in Citation: Meng, L.; Shi, R.; Wang, Q.; α Wang, S. Analysis of Floral Fragrance the other group. Additionally, principal component analysis showed that -ocimene, benzyl alcohol, Compounds of Chimonanthus praecox benzyl acetate, cinnamyl acetate, eugenol, and indole were the main floral fragrance compounds that with Different Floral Colors in could distinguish the four C. -
The Scent-Ual Garden: Creating Four Seasons of Olfactory Delight
The Scent-ual Garden: Creating Four Seasons of Olfactory Delight Winter: Edgeworthia chrysantha (rice paper plant) Edgeworthia chrysantha ‘Akebono’ –orange-red flowered form, blooms more frost tender Daphne odora (winter daphne) o many forms from pure white flowered to pink tinged o solid green leaves, pale yellow margined o Daphne odora ‘Nakafu’ – interesting green & pale green form Sarcococca orientalis (sweet box) – best species for the SE o S. hookeriana var. humilis (Himalayan sweet box) o 'SarSid1' Fragrant Valley™ - 18” x 36” size o 'SarSid2' Fragrant Mountain™ - 24” x 36” size Prunus mume (Japanese flowering apricot) o First of the cherries to begin flowering, sometimes in December o Great for forcing and bringing inside o ‘Bridal Veil’ – white weeping form o ‘Hokkai Bungo’ – earliest to flower, deep red, cinnamon scent o ‘Peggy Clark’ – most common, pink form Mahonia ×media (hybrid grapeholly) o ‘Buckland’, ‘Charity’, ‘Winter Sun’ – most common forms o ×lindsayae ‘Cantab’ – lax form but one of the most fragrant o Self-infertile Chimonanthus praecox (wintersweet) o Flower colors range from near translucent to deep yellow to deep yellow with red center o Great for forcing and bringing inside Spring: Hyacinthus orientalis (common or Dutch hyacinth) o Not particularly long lived o Remove spent flower stalks Phlox divaricata (woodland phlox) o Comes in pinks, whites and shades of blue to purple o Seeds around lightly Dianthus (pinks) o Need good drainage o Look for heat tolerant cultivars o ‘Baths Pink’ o ‘Feuerhexe’ (Firewitch) o ‘Wicked Witch’ o Star series Daphne ‘Blafra’ Eternal Fragrance o Amazingly resilient for a daphne o Big flush of flowers, then re-flowers sporadically through the summer Rhododendron sp. -
Chimonanthus Praecoxpraecox Winterwinter Sweetsweet
ChimonanthusChimonanthus praecoxpraecox WinterWinter sweetsweet Chimonanthus praecox (Winter sweet, melon tree) is native to China, in the mountains of east Sichuan, north-east Chongqing, north-west Hunan and north-west Hubei, up to a height of 3,000 metres. This plant was introduced in Europe by the English in 1766, but they were not the first. The Arabs had taken the shrub with them from China long before that. This can be seen from the Chimonanthus shrubs one finds in almost all Moresque gardens in Spain, like the Alhambra. Winter sweet grows into a wide, vase-shaped, multi-stem shrub 2 to 3 metres high. From December onwards, Chimonanthus praecox blooms light-yellow flowers that stand out with their deliciously sweet scent. In the rest of the year, the rough long leaves offer a lovely background for e.g. permanent plant borders. In autumn, the winter sweet has a bright-yellow autumn colour. From the age of 5, it bears large, urn-shaped fruit with coffee bean-like seeds. For a rich bloom, the shrub prefers a sunny environment, sheltered from wind, but Chimonanthus praecox can also tolerate light shade. A dry summer will make for better buds for the flowers. The Winter sweet prefers moist, nutrient- rich and permeable soil, but also tolerates dryness and heat. The species is an unknown, but easily usable and entirely winter-hardy, multi-stem shrub for gardens. But Chimonanthus will also enrich the range in parks with its winter blooms and delicious accompanying fragrance. TYPES OF PLANTING Tree types: characteristic trees, solitary -
High Line Plant List Stay Connected @Highlinenyc
BROUGHT TO YOU BY HIGH LINE PLANT LIST STAY CONNECTED @HIGHLINENYC Trees & Shrubs Acer triflorum three-flowered maple Indigofera amblyantha pink-flowered indigo Aesculus parviflora bottlebrush buckeye Indigofera heterantha Himalayan indigo Amelanchier arborea common serviceberry Juniperus virginiana ‘Corcorcor’ Emerald Sentinel® eastern red cedar Amelanchier laevis Allegheny serviceberry Emerald Sentinel ™ Amorpha canescens leadplant Lespedeza thunbergii ‘Gibraltar’ Gibraltar bushclover Amorpha fruticosa desert false indigo Magnolia macrophylla bigleaf magnolia Aronia melanocarpa ‘Viking’ Viking black chokeberry Magnolia tripetala umbrella tree Betula nigra river birch Magnolia virginiana var. australis Green Shadow sweetbay magnolia Betula populifolia grey birch ‘Green Shadow’ Betula populifolia ‘Whitespire’ Whitespire grey birch Mahonia x media ‘Winter Sun’ Winter Sun mahonia Callicarpa dichotoma beautyberry Malus domestica ‘Golden Russet’ Golden Russet apple Calycanthus floridus sweetshrub Malus floribunda crabapple Calycanthus floridus ‘Michael Lindsey’ Michael Lindsey sweetshrub Nyssa sylvatica black gum Carpinus betulus ‘Fastigiata’ upright European hornbeam Nyssa sylvatica ‘Wildfire’ Wildfire black gum Carpinus caroliniana American hornbeam Philadelphus ‘Natchez’ Natchez sweet mock orange Cercis canadensis eastern redbud Populus tremuloides quaking aspen Cercis canadensis ‘Ace of Hearts’ Ace of Hearts redbud Prunus virginiana chokecherry Cercis canadensis ‘Appalachian Red’ Appalachian Red redbud Ptelea trifoliata hoptree Cercis -
1. CHIMONANTHUS Lindley, Bot. Reg. 5: T. 404. 1819, Nom. Cons. 蜡梅属 La Mei Shu Butneria Duhamel (1755), Not Byttneria Loefling (1758), Nom
Flora of China 7: 92–94. 2008. 1. CHIMONANTHUS Lindley, Bot. Reg. 5: t. 404. 1819, nom. cons. 蜡梅属 la mei shu Butneria Duhamel (1755), not Byttneria Loefling (1758), nom. cons.; Meratia Loiseleur-Deslongchamps. Shrubs or small trees, erect, deciduous or evergreen. Branchlets dichotomous, quadrangular to subterete; winter buds with im- bricate scales but exposed in summer. Leaf blade papery or subleathery, adaxially scabrous or ± smooth. Flowers axillary, fragrant, subsessile to very shortly pedicellate. Tepals numerous, yellow, yellowish white, or white and sometimes with purple markings, membranous, varying in size and shape from outer to inner but not distinctly dimorphic. Stamens 5–8, arranged on cuplike recep- tacle; filaments filamentous but basally broad and connate, usually puberulent; staminodes few to numerous, puberulous, arranged inside stamens on receptacle. Carpels 5–15, distinct; ovules 2 per carpel but 1 ovule usually abortive. Pseudocarp urceolate, ovoid- ellipsoid, obovoid-ellipsoid, or campanulate, pubescent. Achenes oblong, oblong-ellipsoid, ellipsoid, oblong-ovoid, or reniform. ● Six species: China. It is estimated that the Chinese species diverged from each other perhaps as recently as 1–2 million years ago, and the presently available molecular evidence distinguishes all six species but groups Chimonanthus campanulatus and C. praecox separately (with a bootstrap support of 100) from the other four species (S. L. Zhou et al., Molec. Phylogenetic. Evol. 39: 1–15. 2006). However, the molecular evidence is based on a limited number of samples, mostly from botanical gardens. It is difficult to morphologically circumscribe differences to distinguish all six species. Because the molecular evidence does distinguish all six species, it seems best to treat all six in this account but to point out that additional research may well change this interpretation. -
Chimonanthus Praecox: a Tions to Circulation Manager, Arnoldia, Arnold Arboretum, 125 Arborway, Boston, MA 02130- Redolence of China 3500
The Magazine of the Arnold Arboretum VOLUME 71 • NUMBER 3 The Magazine of the Arnold Arboretum VOLUME 71 • NUMBER 3 • 2014 CONTENTS Arnoldia (ISSN 0004–2633; USPS 866–100) 2 Untangling the Twisted Tale of is published quarterly by the Arnold Arboretum Oriental Bittersweet of Harvard University. Periodicals postage paid Peter Del Tredici at Boston, Massachusetts. Subscriptions are $20.00 per calendar year 19 Magnolia virginiana: Ephemeral domestic, $25.00 foreign, payable in advance. Courting for Millions of Years Remittances may be made in U.S. dollars, by Juan M. Losada check drawn on a U.S. bank; by international money order; or by Visa, Mastercard, or American 28 Wish You Were Here Express. Send orders, remittances, requests to Nancy Rose purchase back issues, change-of-address notices, and all other subscription-related communica- 36 Chimonanthus praecox: A tions to Circulation Manager, Arnoldia, Arnold Arboretum, 125 Arborway, Boston, MA 02130- Redolence of China 3500. Telephone 617.524.1718; fax 617.524.1418; David Yih e-mail [email protected] Arnold Arboretum members receive a subscrip- Front cover: Fruit of Oriental bittersweet (Celas- tion to Arnoldia as a membership benefit. To trus orbiculatus), a notoriously invasive vine. become a member or receive more information, Photo by Peter Del Tredici. please call Wendy Krauss at 617.384.5766 or email [email protected] Inside front cover: Read about the complex flowering and pollination process of sweetbay Postmaster: Send address changes to magnolia (Magnolia virginiana) starting on page Arnoldia Circulation Manager 19. Photo of M. virginiana × virginiana ‘Milton’, The Arnold Arboretum accession 779-87-A, by Michael S.