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Phan H9 Chu Dang - Periplocoideae Thuqc Hq Thien Ly - Asclepiadaceae 0 Viet Nam

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Phan H9 Chu Dang - Periplocoideae Thuqc Hq Thien Ly - Asclepiadaceae 0 Viet Nam HQI NGHI KHOA HQC TOAN QU6C V£ SINH THAI VA TAI NGUYgN SINH VAT tAN THlJ BA PHAN H9 CHU DANG - PERIPLOCOIDEAE THUQC HQ THIEN LY - ASCLEPIADACEAE 0 VIET NAM TRAN THE BACH, VU TIEN CHINH DO VAN HAI, BUI HONG QUANG Vien Sinh thdi vd Tdi nguyen sinh vgt JOONGKU LEE Vi^n Nghien ciht sinh hgc vd Cong nghe sinh hgc Hdn Quoc Hifn nay vi fri phan loai cua phan hp Chu ding - Periplocoideae van dang li van de dang quan tam ciia cic nha phan loai hpc. Nhung dan lifu khoa hpc ve mat hinh thii va sinh hpc phan tir ngay cang dugc bd sung de xic dinh mpt each diing dan vj tri phan loai ciia phan hp Chu ding va tir do gdp phan lam sing td moi quan hf hp hang giira 2 hp Tnic dao - Apocynaceae vi hp Thien ly - Aselepiadaceae, Nhihig thdng tin khoa hpc ciia phan hp Chu dang d Ddng Duong ciing nhu d Vift Nam se gdp phan khdng nhd de giai quyet van de tren. Chinh vi vay chiing tdi da tien hanh nghien cuu phan hp Chu dang d Vift Nam vdi de tai: "Phan hp Chu dang - Periplocoideae thupc hp Thien ly - Aselepiadaceae d Vift Nam". L PHirONG PHAP NGHIEN CUtT Phuang phap so sinh hmh thii dugc ip dyng frong nghien cuu nay. Day la phuong phap tmyen thdng trong phan loai hgc thye vat. n. KET QUA NGHIEN CUtJ 1. Subfam. Periplocoideae R. Br. ex Endl. Phan hg Chu dSng Endl. 1838, Gen, PI. 2: 587; S, Liede & F, Albert, 1994, Taxon 43: 203. Bao phan 4 d, hat phan hgp thanh nhdm 4 hoac dinh thanh khoi phan nhung khdng cd sip bao ben ngoai vich khoi phan. Co quan tmyen phan hinh thia cd day md rpng thanh dia dinh mem, khdng cd gdt dinh, Typus: Periploca L, Phan hp Chu ding gdm 1 tdng Chu ding (Trib. Periploceae) 2. Khoa djnh loai cac chi ciia phan hp Chu dSng (Periplocoideae) thupc hg Thien ly (Aselepiadaceae) d Vift Nam 1 A. Cay byi dung 1. Telectadium IB, Cay leo hay trudn, 2A, Trang phy dinh vdi trang. 3A, Tiln khai hoa van 2. Hemidesmus 3B, Tien khai hoa van. 4A, Vay trang phy khdng xe thiiy,. 3. Cryptolepis 4B. Viy trang phy xe sau thanh 2 thiiy 4. Zygoslelma 2B, Tring phy khdng dinh vdi tring. 5A. Giira cac chi nhj khdng cd tuyen. 6A. Viy trang phy hinh trimg 5. Gymnanthera Ltn cam om: Cac tac gi4 xin chSn thanh c4m on sv giup (15 cua de tai "Tiem nang sinh hpc cua nguyen ii?u sinh hoc a Vi?Viet Nam'" ^ 28 HQI NGHI KHOA HQC TOAN QUOC V£ SINH THAI VA TAI NGUYEN SINH VAT LAN THU' BA 6B. Vay trang phu hinh sgi 6, Finlaysonia 5B. Giira cac chi nhi cd tuyen, 7A. Mat trong cua dai khdng cd tuyen d diy, 8A, Viy trang phy ngin, hinh tam giie 7, Alherolepis 8B. Viy trang phu dai, hinh sgi %. Atherandra IB. Mat trong cua dai cd tuyen d diy, 9A, Bao phan cd ldng, Thiiy trang phu chia 3 phan, phan d giira dang sgi, 2 phan ben rdng ra dang cii nip tiii 9, Periploca 9B. Bao phan nhan. Thiiy tring phu khdng chia thanh 3 phan nhu tren lOA. Qui khdng cd gd nhu cinh, cd long day 10. Streptocaulon lOB. Quacd gd nhu canh, nhin 11. Myriopleron 3. Mo ta cac chi cua phan hp Chu dang d Vift Nam 3.1. Telectadium Baill.-Vf tuyen Baill. 1889. Bull. Soc. Linn. Paris 2: 801; Cost. 1912. Fl. Gen. hidoch. 4: 36; Endress & Bmyns, 2000. Bot. Rew. 66(1): 44, - Ten Vift khic: Phao ludi. Cay byi dung. Li mpc ddi hay ddi khi mpc vdng, thudng thudn hfp. Cym hoa cd li bic rat phat trien, Ny rdng d dudi hem tren, Trang hinh binh xe; cinh hoa vin, phu nhau phii. Trang phy gom 5 viy day, Chi nhj rdi; bao phan 4 d; hat phan hop thinh nhdm 4; co quan truyen phan hinh thia cd day md rpng thanh dia dinh mem. Typus: Telectadium edule Baill. Cd khoang 3-4 loii phan bd d nhift ddi Chau A. 6 Vift Nam cd 3 loai: Telectadium edule Baill.-Vf tuyen ngpt, Telectadium linearicarpum Pierre ex Cost,-Vf tuyen la hpp, Telectadium dongnaiense Pierre ex Cost,-Vf tuyen dong nai Khoa djnh loai cac loai ciia chi Telectadium is Vift Nam 1 A, Co quan truyen phan cao hon hay bing 2mm, ong tring khdng chia thinh 2 phan 2A, Qui hinh biu dye /. T. edule 2B. Qua hinh dii 2, T. linearicarpum IB, Co quan truyen phan cao 1,1-1,4mm, dng trang chia thanh 2 phan rd (phan gde phong to len dang cau, phan tren thu hpp lai thanh dang hmh try) 3. T. dongnaiense 3.2, Hemidesmus R, Br.-Bin frang R, Br 1810. Asclep. 45; id, 1811, Mem, Werner Nat. Hist. Soc. 1: 56; Hook, f 1883. Fl, Brit, hid, 4: 4; K. Schum, in Engl, & Pranti, 1895, Nat, Pflanzenfam, 4(2): 218; Decne, m A, DC, 1844, Prodr. 8:494; Cost. 1912. Fl. Gen. hidoch. 4: 15; Endress & Bmyns, 2000. Bot. Rew, 66 (1): 43. Cay byi trudn. Li mpc doi, hiem khi mpc chym 4. Cum hoa ngin, nhanh ngin, it hoa, Ny hmh triimg-ndn, Dai ngin, hinh tning, gde dai cd 5 tuyen, Tring hinh binh xe; tien khai hoa van, dng tring ngin hon thiiy trang, Viy trang phy 5, ngin, nac, dinh vdi trang, Chi nhj rdi hay gan rdi, chi nhj dinh d goc trang, thudng dai, uon cong ra phia ngoai; bao phan 4 d, gin vdi dau nhyy, bao phan cd phin phu dang la ngin; hat phan hpp thanh nhdm 4; co quan tmyen phan hinh thia cd diy md rpng thanh dia dinh mem. Qua thudng gdm 2 dai. Typus: Hemidesmus indicus (L.) R. Br. ex Schult. 29 HQI NGHI KHOA HQC TOAN QUOC V£ SINH THAI VA TAI NGUY£N SINH VAT LAN THLf BA Cd khoang 2 loii phan bd d Chau A tir An Dp din Vift Nam, Thai Lan, Malaixia, Xrilanca. Q Vift Nam cd 1 loai: Hemidesmus indicus (L.) R. Br, ex SchuU.-Bin trang 3.3, Cryptolepis R, Br,- An lan R, Br, 1810, Asclep, 58; id, 1811, Mem, Wemer, Nat, Hist, Soc, 1: 69; Hook, f 1883, FI, Brit, hid, 4: 5; K. Schum. in Engl. & Pranfl, 1895, Nat, Pflanzenfam, 4(2): 219; Cost, 1912, Fl, Gen. Indoeh. 4: 23; Back. & Bakh. f 1965. Fl. Java 2: 250; Tsiang & Li, 1977. Fl. Reip. Pop. Sin, 63: 262; P, T, Li & al, 1995, Fl. China 16: 193; Endress & Bmyns, 2000, Bot, Rew, 66 (1): 43, _ Ten Vift khic: Cang cua. Cay bui trudn. La mpc ddi, hmh biu due din thudn, cd nhieu gan nam ngang. Cum hoa xim, it hoa. Nu hoa keo dai thinh hinh cii khoan, Thuy dai nhd, dau gan tii; gde dai cd 5 hay nhieu tuyen (vay), Trang hinh binh xe, tiln khai hoa van, phii nhau phai; dng trang ngin hon thiiy trang, Trang phu cd 5 vay nac, khdng xe thiiy, gin hinh friing, dinh d giua dng trang. Chi nhj ngin, rdng d dudi, hpp d tren, dinh nhau mdt phan d diy; bao phan 4 d, dinh vio dau nhuy tao thinh dang ndn nhpn; hat phin hpp thanh nhdm 4; co quan truyen phan hinh thia cd day md rdng thanh dTa dinh mem. Dau nhyy hinh ndn rdng. Qua gdm 2 dai, tda rpng, nhan. Typus: Cryptolepis buchananii Roem. & Schult. (vide S. Liede & F. Albers, 1994). Cd 12 loai d Chau A va nhiet ddi Chau Phi. 6 Vift Nam cd 2 loai: Cryptolepis buchananii Roem. & Schult. -An lan buchana, Cryptolepis sinensis (Lour.) Merr. Day li bac. Khda djnh loai cac loai cua chi Cryptolepis a Vift Nam IA, Gin ben 25-30, dai nhin 1, C, buchananii IB. Gan ben 5-9, dai cd long 2. C. sinensis 3.4, Zygostelma Benth, (Day) Dy gdt Benth, in Benth, & Hook, f 1876. Gen. PI. 2: 740; K. Schum, m Engl. & Pranti, 1895. Nat. Pflanzenfam. 4(2): 221; Cost. 1912. Fl. Gen. fridoch. 4: 18; Endress & Bmyns, 2000. Bot, Rew. 66 (1): 44, -Ten Vift khic: Xy got. Cay byi leo hay tmdn. Li mpc doi, thudn. Cum hoa xim, ngan, it hoa. Ny rdng ra d dudi, hinh chiiy dai, vin d tren. Thuy dai ngin, hinh bau dye hay hinh friing; d gde dai cd 5 tuyin. Trang hinh binh xe, nac; cinh hoa hinh tning, tiln khai hoa van, phu nhau phai; dng trang ngin hem thiiy trang. Trang phy gom 5 viy, mdi viy xe sau thanh 2 thiiy, dinh vdi trang. Chi nhj ngin, dinh nhau d phan dudi (gin rdi); bao phin 4 d; bao phin keo dii thanh phin phy mdng va ndi vdi dau nhuy phdng len tao thanh hinh ndn, hat phan hpp thanh nhdm 4; co quan tmyin phan hinh thia cd diy md rpng thinh dTa dinh mem. Typus: Zygostelma benlhamii Baill, (vide S, Liede & F, Albers, 1994). Cd 1 loii phan bo d vimg Ddng Nam A (Lio, Vift Nam, Campuchia, Thai Lan): Zygoslelma benlhamii Baill. -(day) Dy gdt 3.5, Gymnanthera R. Br, - Loa ti gia R, Br. 1810. Asclep, 47; id, 1811. Mem, Werner, Nat. Hist, Soc. 1: 58; K. Schum. in Engl.
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