Hardy Aroids in the Garden Judy Glattstein

Total Page:16

File Type:pdf, Size:1020Kb

Hardy Aroids in the Garden Judy Glattstein Hardy Aroids in the Garden Judy Glattstein Though not showy plants and with only a modest following among plant lovers, the hardy aroids are interesting, display many virtues in cultivation, and attract "a different class of gardeners" The Arum Family, or Araceae, consists of tion, especially in the United States. They about fifteen genera, most of them tropi- are, therefore, unusual and have the cal but of wide distribution. Some of the appeal of novelty. tropical members of the family have long Aroids contain a bitter substance, calci- been under cultivation, especially in east- um oxalate, and are little bothered by ern Asia and the Pacific Islands. Taro pests. Slugs, mice, rabbits, and deer find (Colocasia esculenta) and several species them decidedly unpalatable. When aroids of Xanthosoma (yautia), for example, are are used for food, the calcium oxalate grown for their edible tubers as staple first must be destroyed by heat. Garden- sources of starch. Other tropical species ers should be careful to wash their hands are handsome foliage plants used in the after handling berries or a bruised tuber. temperate zones for summer bedding (Ca- Once, after cleaning Aris~ma seeds, I ladium) or as houseplants (Aglaonema, inadvertently touched my mouth. The Dieffenbachia, Monstera, Philodendron). resulting unpleasant tingling and numb- Others are used by florists as cut flowers ness took several hours to wear off. (Anthurium, Calla~.I. My garden in Wilton, Connecticut, is Some members of the family are hardy, shaded by mature white oaks (Quercus notably Aris~ma, Arisarum, Arum, Lysi- alba). Understory trees are dogwood chiton, and Symplocarpus. The Araceae (Cornus florida) and black birch (Betula might seem a poor prospect for garden- lenta). The Araceae I raise are quite worthy plants to those familiar only with hardy in Wilton, which is situated in Ar- the skunk cabbage (Symplocarpus fc~tid- nold Arboretum Hardiness Zone 6 (-5 us) of New England’s swamps. I have en- Fahrenheit to 5 Fahrenheit). In fact, the joyed cultivating representatives of sever- temperature once dipped to -8 Fahren- al genera, some for their flowers, some for heit, and there were no losses. The soil in their foliage. the garden is a good loam, which I keep Aroids have a modest following, appear- mulched with leaves for a constant supply ing in an occasional article, mentioned of humus; as in most of Connecticut, the briefly in gardening books. Visitors to my pH is rather low (acid). Other plants I use garden have admired them; they have sev- in the garden include such American wild- eral points of appeal. Many of the aroids flowers as Trillium, Sanguinaria canaden- I discuss in this article are rare in cultiva- se (bloodroot), Hexastylis spp. (evergreen 28 gingers from the southeastern states), Northeast are low in phosphorus, and pot- Phlox stolonifera, Phlox divaricata, and ash is especially useful for tuberous many kinds of ferns. Other shade-tolerant plants. It is not safe to use bonemeal in plants, such as hostas, epimediums, and my garden because it attracts skunks, primroses, also do well under these condi- which dig up the tubers looking for bones. tions. They do not eat the tubers, but it is a nui- Since I have to obtain most of the a- sance to replant them. Nitrogen is ap- roids from abroad, I prefer to receive plied in the spring, in the form of dried them in the autumn. They are completely blood, cottonseed meal, or leather tank- dormant at this time, and the tubers travel age. Fertilization after the first year is usu- well and arrive in excellent condition. If ally not required. The constant mulch of they are shipped in the spring, there is leaves seems to keep the plants growing the risk that they will break dormancy in good condition. while in transit. New growth can be dam- An alternative way of obtaining these aged either by the confines of the ship- plants is to raise them from seed. I soak ping container, or by rot. As soon as the dried berries in a little tepid water for an tubers are received they are planted hour or so, until the coat softens. Then, I directly in the garden. The area is spad- rub the seeds gently between paper tow- ed over, and extra compost is added if els and separate the seed. Each berry necessary. I fertilize with muriate of has one to four seeds. I sow the seeds in potash and superphosphate. Soils in the a sterile mix of half potting soil and half The familiar skunk cabbage (Symplocarpus foetidus) Lysichiton americanum in flower in the wild, Washing- of New England’s swamps. This and all other ton, D. C. photographs accompanying this article were taken by the author. 29 Jiffy-mix® or Pro-mix@, with enough sharp gressed from an asexual to a male and sand for good drainage. (I sow them thin- finally to a female state, remaining in the ly enough that I won’t have to prick them last state. Many plants-Ilex and Myrica, out for a year.) I cover the the seeds well, for example-have single-sexed plants water them, and wait. Fresh seeds will ger- that are either male or female and that minate promptly under growth lights. remain so for the life of the individual Older seeds will germinate more slowly, plant, a condition called "dioecious." The and outdoor conditions slow the germina- transitional nature of the sexual state of tion process somewhat. Arisxma is referred to as "paradioe- My biggest problem has been to keep cious."" the plants through their dormant stages. While the garden site may be quite Aris~rna damp, pot-grown plants rot with the great- In North America there are two species est of ease. At the same time, small tu- of Arisa?ma, Arisxma triphyllum, which bers dry out quickly. It is difficult to find has four subspecies, and Arisxma dra- the correct balance. Second-year plants contium of the southeastern states. can go into a prepared site in the garden Aris~ma triphyllum (Linnxus) Torrey and should begin flowering in their third is found from the Gaspe Peninsula, or fourth year. I have used this method southern Quebec and Ontario, Wisconsin with several species of Arisxma and with and Minnesota south to eastern Texas Arum italicum. Aris~ma seeds do not and southern Florida, growing in moist, need a period of stratification but will ger- shady woodlands. There are four sub- minate during the autumn they ripen if specific populations, with widespread hy- they are sown indoors. Sown outdoors in brid swarms. the autumn they will, of course, germi- Arisxma triphyllum ssp. triphyllum is nate the following spring. The production the most widespread. Its height varies of seeds is generous, one spadix of Ari- with growing conditions. I have seen sxma sikokianum having from one to specimens that were dwarf in the wild four seeds in a berry, for a total of five reach two feet in height in the garden with hundred eighty-seven seeds. Plants of Ari- richer soil and ample water. Typically, it s~ma sikokianum often begin to flower in has one or two leaves, each bearing three their third year. Once established, the leaflets, which are glaucous beneath. The plants are most agreeable. I have dug spathe may vary in color from green to one up in full bloom, potted it for a rock- green-and-purple striped, to chocolate garden show, and replanted it in the gar- purple. The name ’Zebrinum’ is often ap- den without any difficulty or damage to plied to cultivars whose spathes are the plant. purple to bronze and have whitish longitu- The flowering of Arisxma follows an dinal stripes inside. An interesting variant unusual pattern. Immature corms, from has recently been discovered by Peggy either seeds or offsets, are asexual and French in Wilton, Connecticut. It has pro- have a single foliage leaf. As corms in- nouncedly white-veined leaves and crease in size after their first year, they comes true from seed. reach sexual maturity, producing two The second subspecies, which I have leaves and one scape. Smaller (lighter) seen in several gardens, is Arisxma tri- corms are male, heavier corms are invari- phyllum ssp. stewardsonii. This is a ably female, the sexual state having pro- northern variant in which the spathe is 30 green and strongly fluted with white site. The leaf is solitary, three-parted, and ridges on the outside. It tends to appear a glossy mid-green; it appears after later in the spring than the other sub- flowering, which occurs early in June. species and grows consistently in moist The spathe is very beautifully marked sites. Its leaves are never glaucous. with pink and white stripes. Mature tubers The third subspecies is Arisa?ma make numerous offsets, which form a triphyllum ssp. pusillum, which grows in good-sized clump in a few years. the same habitat as Arisa?ma triphyllum Aris~ma sikokianum Franchet and Sa- ssp. stewardsonii, although farther south vatier comes from Honshu, Shikoku, and and at lower elevations. Its leaves, too, Kyushu in Japan. Mature plants have two are never glaucous. There are no ridges three- to five-parted leaves that often have on the spathe, and the coloring is nearly attractive silver markings. Its Japanese always completely green or completely name, yuki-mochi-so, means "snow rice- purple, occasionally with thin, green cake plant," in reference to the pure- stripes. white, clublike spadix. The spathe is a The fourth subspecies, Arisa?ma tri- deep chocolate brown on the outside, phyllum var. quinatum, has a very re- green shading to white inside.
Recommended publications
  • Size Variations of Flowering Characters in Arum Italicum (Araceae)
    M. GIBERNAU,]. ALBRE, 2008 101 Size Variations of Flowering Characters in Arum italicum (Araceae) Marc Gibernau· and Jerome Albre Universite Paul Sabatier Laboratoire d'Evolution & Diversite Biologique (UMR 5174) Bat.4R3-B2 31062 Toulouse cedex 9 France *e-mail: [email protected] ABSTRACT INTRODUCTION In Arum, bigger individuals should An extreme form of flowering character proportionally invest more in the female variations according to the size is gender function (number or weight of female modification, which occurs in several flowers) than the male. The aim of this species of Arisaema (Clay, 1993). Individ­ paper is to quantify variations in repro­ ual plant gender changes from pure male, ductive characters (size of the spadix when small, to monoecious (A. dracon­ parts, number of inflorescences) in rela­ tium) or pure female (A. ringens) when tion to plant and inflorescence sizes. The large (Gusman & Gusman, 2003). This appendix represents 44% of the spadix gender change is reversible, damaged length. The female zone length represents female individuals will flower as male the 16.5% of the spadix length and is much following year (Lovett Doust & Cavers, longer than the male zone (6%). Moreover 1982). These changes are related to change these three spadix zones increase with in plant size and are explained by the plant vigour indicating an increasing size-advantage model. The size-advantage investment into reproduction and pollina­ model postulates a sex change when an tor attraction. It appears that the length of increase in body size is related to differen­ appendix increased proportionally more tial abilities to produce or sire offspring than the lengths of the fertile zones.
    [Show full text]
  • The Evolution of Pollinator–Plant Interaction Types in the Araceae
    BRIEF COMMUNICATION doi:10.1111/evo.12318 THE EVOLUTION OF POLLINATOR–PLANT INTERACTION TYPES IN THE ARACEAE Marion Chartier,1,2 Marc Gibernau,3 and Susanne S. Renner4 1Department of Structural and Functional Botany, University of Vienna, 1030 Vienna, Austria 2E-mail: [email protected] 3Centre National de Recherche Scientifique, Ecologie des Foretsˆ de Guyane, 97379 Kourou, France 4Department of Biology, University of Munich, 80638 Munich, Germany Received August 6, 2013 Accepted November 17, 2013 Most plant–pollinator interactions are mutualistic, involving rewards provided by flowers or inflorescences to pollinators. An- tagonistic plant–pollinator interactions, in which flowers offer no rewards, are rare and concentrated in a few families including Araceae. In the latter, they involve trapping of pollinators, which are released loaded with pollen but unrewarded. To understand the evolution of such systems, we compiled data on the pollinators and types of interactions, and coded 21 characters, including interaction type, pollinator order, and 19 floral traits. A phylogenetic framework comes from a matrix of plastid and new nuclear DNA sequences for 135 species from 119 genera (5342 nucleotides). The ancestral pollination interaction in Araceae was recon- structed as probably rewarding albeit with low confidence because information is available for only 56 of the 120–130 genera. Bayesian stochastic trait mapping showed that spadix zonation, presence of an appendix, and flower sexuality were correlated with pollination interaction type. In the Araceae, having unisexual flowers appears to have provided the morphological precon- dition for the evolution of traps. Compared with the frequency of shifts between deceptive and rewarding pollination systems in orchids, our results indicate less lability in the Araceae, probably because of morphologically and sexually more specialized inflorescences.
    [Show full text]
  • Italian Arum Flyer
    PUBLIC ANNOUNCEMENT- WESTERN INVASIVES NETWORK Revised April, 2020 847 NW Monroe Ave. Corvallis, OR 97330 (541) 910-8769 Photo: Travis Johnson 1Photo: Liz West************ NOXIOUS WEED ALERT************ Italian Arum Italian arum (Arum italicum), also known as Digging and disposing of the small “orange candleflower” and “Italian lords and underground corms in the garbage can help. ladies,” is a non-native perennial that was However, manual removal is only introduced as an ornamental plant. It has recommended on small patches, because if now naturalized and appears to be the plant is not completely eradicated, the spreading. Arum italicum is a woodland soil disturbance can increase its spread. If species and prefers moist, well-shaded you have it on your land, do not let it go to environments. It’s extremely difficult to seed. Wearing gloves, cut and bag all seed eradicate once it becomes established and heads, and dispose of them in the garbage. may spread from residential gardens into The variegated, exotic leaves and showy red fruits woodland areas. For more info on Italian arum: make it an attractive garden plant. Unfortunately, Western Oregon is the perfect environment for Why is it a problem? WA Noxious Weed Control Board unwanted invasions. (Photo: East Multnomah SWCD) This plant is problematic because of its ability Lower Hudson PRISM to form a dense cover in open sites. It can out-compete native plants and prevent their The Western Invasives Network encourages establishment. you to report Italian arum, in an effort to inform future management priorities across Italian arum is also toxic to humans and western Oregon.
    [Show full text]
  • Italian Arum What Is Italian Arum ? Italian Arum, Arum Italicum, Is Also Known As Orange Candleflower, Cuckoo’S Pint, and Italian Lords-And-Ladies
    February Invasive Plant Highlight Italian Arum What is Italian Arum? Italian Arum, Arum italicum, is also known as Orange Candleflower, Cuckoo’s Pint, and Italian Lords-and-Ladies. It is native to the Mediterranean region of Europe. It has been cultivated as an ornamental plant for traditional and shade gardens in the U.S. Italian arum is a perennial, herbaceous plant that grows from small pea-sized white bulbs. Many bulbs are produced during the growing season and they make it difficult to eradicate the plant. Italian Arum leaves Italian Arum is classified as a toxic weed. It is poisonous to humans if ingested and can cause an allergic reaction if touched without using gloves. Italian Arum can grow 12 to 18 inches. It blooms in the spring with white flowers that make bright orangish-red fruit. It spreads when its seed disperses by water or by birds consuming the berries. The plant invades riparian forest areas, threatening native plant diversity and also can cause increased erosion and slope instability along streambanks. Italian Arum berries Better Groundcovers Alumroot, Wild Ginger, Mayapple, Jack-in-the-pulpit, and Allegheny Spurge Please Remove It! If you have Italian Arum, please remove it before it spreads into other adjacent properties. Italian Arum emerges in the Fall and will stay green during mild winters making it easy to spot. Herbicides are not effective since death of foliage above ground does not indicate that the bulbs have died. To remove small patches with a trowel, dig down to root end (about 6 inches) and sift the soil to extract all the small bulbs.
    [Show full text]
  • History and Current Status of Systematic Research with Araceae
    HISTORY AND CURRENT STATUS OF SYSTEMATIC RESEARCH WITH ARACEAE Thomas B. Croat Missouri Botanical Garden P. O. Box 299 St. Louis, MO 63166 U.S.A. Note: This paper, originally published in Aroideana Vol. 21, pp. 26–145 in 1998, is periodically updated onto the IAS web page with current additions. Any mistakes, proposed changes, or new publications that deal with the systematics of Araceae should be brought to my attention. Mail to me at the address listed above, or e-mail me at [email protected]. Last revised November 2004 INTRODUCTION The history of systematic work with Araceae has been previously covered by Nicolson (1987b), and was the subject of a chapter in the Genera of Araceae by Mayo, Bogner & Boyce (1997) and in Curtis's Botanical Magazine new series (Mayo et al., 1995). In addition to covering many of the principal players in the field of aroid research, Nicolson's paper dealt with the evolution of family concepts and gave a comparison of the then current modern systems of classification. The papers by Mayo, Bogner and Boyce were more comprehensive in scope than that of Nicolson, but still did not cover in great detail many of the participants in Araceae research. In contrast, this paper will cover all systematic and floristic work that deals with Araceae, which is known to me. It will not, in general, deal with agronomic papers on Araceae such as the rich literature on taro and its cultivation, nor will it deal with smaller papers of a technical nature or those dealing with pollination biology.
    [Show full text]
  • Total Phenolics, Antioxidant, Antibacterial and Cytotoxic Activity Studies of Ethanolic Extracts Arisarum Vulgare O.Targ.Tozz
    Int. J. Sec. Metabolite, Vol. 4, Issue 2 (2017) pp. 114-122 Research Article ISSN: 2148-6905 online Journal homepage: http://www.ijate.net/index.php/ijsm Total Phenolics, Antioxidant, Antibacterial and Cytotoxic Activity Studies of Ethanolic Extracts Arisarum vulgare O.Targ.Tozz. and Dracunculus vulgaris Schott. Çiğdem AYDIN1,*, Cennet ÖZAY1, Olcay DÜŞEN1, Ramazan MAMMADOV1, Figen ORHAN1 1Pamukkale University, Science & Art Faculty, Department of Biology, 20070, Denizli, Turkey Received: 08 February 2017 - Revised: 20 March 2017 - Accepted: 18 April 2017 Abstract: The present study reports the antioxidant, total phenolics, antibacterial and cytotoxic activities of Arisarum vulgare and Dracunculus vulgaris of aerial and under ground ethanol extracts. The antioxidant activity was determined by DPPH radical scavenging activity and β-carotene linoleic acid assays. Folin-Ciocalteu assay was used to detect total phenolic contents in the extract. In vitro cytotoxic activity was determined by the Brine shrimp lethality test. Antibacterial activity was investigated with the microdilution method. As a result, under ground parts of the extract showed higher antioxidant activity than aerial parts of the extract. Total phenolics ranged from 14.5±2.02 to 53.4±2.01 mg GAE/g, and the antioxidant activity according to the β-carotene/linoleic acid assay ranged from 70.76±1.08 % to 85.43±1.05 and according to the DPPH assay IC50 values ranged from 0.089±1.02 to 1.095±1.07mg/ml. The A. vulgare under ground extract was tested against 2 and showed a good antibacterial activity at a concentration of <50 g/mL Minimum inhibitory concentration (MIC) values for the bacteria S.aureus.
    [Show full text]
  • Pollinators and Visitors of Aroid Inflorescences
    66 AROJDEANA, Vol. 26 Pollinators and Visitors of Aroid Inflorescences Marc Gibernau Laboratoire d'Evolution & Diversite Biologique Universite de Toulouse m 118 Route de Narbonne, B§t. IV R 3-B 2 31062 Toulouse Cedex 4 France e-mail: [email protected] ABSTRACf view of this subject, as Thomas Croat (2000) did in his history and current status Data on aroid pollinators was first sum­ of systematic research with Araceae, but to marized by Grayum (1984) who docu­ give, in the first place, a statement of the mented 35 genera and about 90 species. A subject and to develop some remarks on second summary was published in 1997 in aroid pollination. The Genera ofAraceae (Mayo et al., 1997) with 38 genera and less than 100 species listed including data from Grayum 0986, RESULTS 1990). This paper brings the reference list Summarizing data from Grayum (1984, up to date since 1997, documenting the 1990) and Mayo et al. (997), and includ­ pollinators of 49 genera and about 125 ing omitted and new publications, the pol­ species. These numbers are still very low linators of 49 genera and about 125 spe­ in comparison with the diversity of the Ar­ cies are documented in Table 1. These aceae family which contains 105 genera numbers are still very low in comparison and about 3,300 species. Some questions with the diversity of the Araceae family on aroid pollination are developed in the which contains 105 genera and about discussion. 3,300 species (Mayo et at., 1997). Thus, KEYWORDS pollinators are cited for only 47% of the genera.
    [Show full text]
  • Araceae) from the Upper Maastrichtian of South Dakota
    Int. J. Plant Sci. 177(8):706–725. 2016. q 2016 by The University of Chicago. All rights reserved. 1058-5893/2016/17708-0006$15.00 DOI: 10.1086/688285 EVALUATING RELATIONSHIPS AMONG FLOATING AQUATIC MONOCOTS: A NEW SPECIES OF COBBANIA (ARACEAE) FROM THE UPPER MAASTRICHTIAN OF SOUTH DAKOTA Ruth A. Stockey,1,* Gar W. Rothwell,*,† and Kirk R. Johnson‡ *Department of Botany and Plant Pathology, Oregon State University, 2082 Cordley Hall, Corvallis, Oregon 97331, USA; †Department of Environmental and Plant Biology, Ohio University, Athens, Ohio 45701, USA; and ‡National Museum of Natural History, Smithsonian Institution, MRC 106, PO Box 37012, Washington, DC 20013-7012, USA Editor: Patrick S. Herendeen Premise of research. A large number of floating aquatic aroid fossils have been recovered from pond sediments in the Hell Creek Formation (Upper Cretaceous) of South Dakota, providing valuable new data about aquatic vegetation of the uppermost Cretaceous, that are used to describe a new species of the genus Cobbania,and to evaluate associated reproductive structures and phylogenetic relationships among floating aquatic monocots. Methodology. Fossils were uncovered as needed with fine needles to reveal surface features of the specimens. Images were captured with a digital scanning camera, and phylogenetic analyses were conducted with TNT implemented through WinClada. Pivotal results. The new species, Cobbania hickeyi Stockey, Rothwell & Johnson, extends the range of the genus to the uppermost Cretaceous, supports the taxonomic integrity of the genus Cobbania, and increases our understanding of structural variation and species richness within the genus. Associated reproductive structures include an aroid spadix, strengthening the assignment of Cobbania to the Araceae.
    [Show full text]
  • An Encyclopedia of Shade Perennials This Page Intentionally Left Blank an Encyclopedia of Shade Perennials
    An Encyclopedia of Shade Perennials This page intentionally left blank An Encyclopedia of Shade Perennials W. George Schmid Timber Press Portland • Cambridge All photographs are by the author unless otherwise noted. Copyright © 2002 by W. George Schmid. All rights reserved. Published in 2002 by Timber Press, Inc. Timber Press The Haseltine Building 2 Station Road 133 S.W. Second Avenue, Suite 450 Swavesey Portland, Oregon 97204, U.S.A. Cambridge CB4 5QJ, U.K. ISBN 0-88192-549-7 Printed in Hong Kong Library of Congress Cataloging-in-Publication Data Schmid, Wolfram George. An encyclopedia of shade perennials / W. George Schmid. p. cm. ISBN 0-88192-549-7 1. Perennials—Encyclopedias. 2. Shade-tolerant plants—Encyclopedias. I. Title. SB434 .S297 2002 635.9′32′03—dc21 2002020456 I dedicate this book to the greatest treasure in my life, my family: Hildegarde, my wife, friend, and supporter for over half a century, and my children, Michael, Henry, Hildegarde, Wilhelmina, and Siegfried, who with their mates have given us ten grandchildren whose eyes not only see but also appreciate nature’s riches. Their combined love and encouragement made this book possible. This page intentionally left blank Contents Foreword by Allan M. Armitage 9 Acknowledgments 10 Part 1. The Shady Garden 11 1. A Personal Outlook 13 2. Fated Shade 17 3. Practical Thoughts 27 4. Plants Assigned 45 Part 2. Perennials for the Shady Garden A–Z 55 Plant Sources 339 U.S. Department of Agriculture Hardiness Zone Map 342 Index of Plant Names 343 Color photographs follow page 176 7 This page intentionally left blank Foreword As I read George Schmid’s book, I am reminded that all gardeners are kindred in spirit and that— regardless of their roots or knowledge—the gardening they do and the gardens they create are always personal.
    [Show full text]
  • Woodard Bay Natural Resources Conservation Area Vascular Plant List
    Woodard Bay Natural Resources Conservation Area Vascular Plant List Courtesy of DNR staff and the Washington Native Plant Society. Nomenclature follows Flora of the Pacific Northwest 2nd Edition (2018). * - Introduced Genus/Species Common Name Plant Family Abies grandis Grand fir Pinaceae Acer circinatum Vine maple Sapindaceae Acer macrophyllum Bigleaf maple Sapindaceae Achillea millefolium Common yarrow Asteraceae Achlys triphylla Vanilla leaf Berberidaceae Actaea rubra Baneberry Ranunculacae Adenocaulon bicolor Path finder, trail plant Asteraceae Adiantum aleuticum (A. pedantum) Northern maidenhair fern Pteridaceae Agrostis exarata Spike bentgrass Poaceae Aira caryophyllea* Silver hairgrass Poaceae Amelanchier alnifolia Western serviceberry, saskatoon Rosaceae Anaphalis margaritacea Pearly everlasting Asteraceae Anthemis cotula* Stinking chamomile, mayweed chamomile Asteraceae Aquilegia formosa Red columbine Ranunculaceae Arbutus menziesii Pacific madrone Ericaceae Arctostaphylos uva-ursi Kinnikinnick Ericaceae Arctium minus* Common burdock Asteraceae Arum italicum* Italian arum Araceae Asarum caudatum Wild ginger Aristolochiaceae Athyrium filix-femina Common lady fern, northern lady fern Athyriaceae Atriplex patula* Saltbush, spearscale Amaranthaceae Bellis perennis* English lawn daisy Asteraceae Mahonia aquifolium Tall Oregon grape Berberidaceae Mahonia nervosa Dull Oregon-grape, Low Oregon-grape Berberidaceae Struthiopteris spicant Deer fern Blechnaceae Brassica nigra* Black mustard Brassicaceae Bromus spp** Brome grasses Poaceae
    [Show full text]
  • Interpretation Manual of European Union Habitats - EUR27 Is a Scientific Reference Document
    INTERPRETATION MANUAL OF EUROPEAN UNION HABITATS EUR 27 July 2007 EUROPEAN COMMISSION DG ENVIRONMENT Nature and biodiversity The Interpretation Manual of European Union Habitats - EUR27 is a scientific reference document. It is based on the version for EUR15, which was adopted by the Habitats Committee on 4. October 1999 and consolidated with the new and amended habitat types for the 10 accession countries as adopted by the Habitats Committee on 14 March 2002 with additional changes for the accession of Bulgaria and Romania as adopted by the Habitats Committee on 13 April 2007 and for marine habitats to follow the descriptions given in “Guidelines for the establishment of the Natura 2000 network in the marine environment. Application of the Habitats and Birds Directives” published in May 2007 by the Commission services. A small amendment to Habitat type 91D0 was adopted by the Habitats Committee in its meeting on 14th October 2003. TABLE OF CONTENTS WHY THIS MANUAL? 3 HISTORICAL REVIEW 3 THE MANUAL 4 THE EUR15 VERSION 5 THE EUR25 VERSION 5 THE EUR27 VERSION 6 EXPLANATORY NOTES 7 COASTAL AND HALOPHYTIC HABITATS 8 OPEN SEA AND TIDAL AREAS 8 SEA CLIFFS AND SHINGLE OR STONY BEACHES 17 ATLANTIC AND CONTINENTAL SALT MARSHES AND SALT MEADOWS 20 MEDITERRANEAN AND THERMO-ATLANTIC SALTMARSHES AND SALT MEADOWS 22 SALT AND GYPSUM INLAND STEPPES 24 BOREAL BALTIC ARCHIPELAGO, COASTAL AND LANDUPHEAVAL AREAS 26 COASTAL SAND DUNES AND INLAND DUNES 29 SEA DUNES OF THE ATLANTIC, NORTH SEA AND BALTIC COASTS 29 SEA DUNES OF THE MEDITERRANEAN COAST 35 INLAND
    [Show full text]
  • Review of Recent Plant Naturalisations in South Australia and Initial Screening for Weed Risk
    Review of recent plant naturalisations in South Australia and initial screening for weed risk Technical Report 2012/02 www.environment.sa.gov.auwww.environment.sa.gov.au Review of recent plant naturalisations in South Australia and initial screening for weed risk Chris Brodie, State Herbarium of SA, Science Resource Centre, Department for Environment and Natural Resources and Tim Reynolds, NRM Biosecurity Unit, Biosecurity SA June 2012 DENR Technical Report 2012/02 This publication may be cited as: Brodie, C.J. & Reynolds, T.M. (2012), Review of recent plant naturalisations in South Australia and initial screening for weed risk, DENR Technical Report 2012/02, South Australian Department of Environment and Natural Resources, Adelaide For further information please contact: Department of Environment and Natural Resources GPO Box 1047 Adelaide SA 5001 http://www.environment.sa.gov.au © State of South Australia through the Department of Environment and Natural Resources. Apart from fair dealings and other uses permitted by the Copyright Act 1968 (Cth), no part of this publication may be reproduced, published, communicated, transmitted, modified or commercialised without the prior written permission of the Department of Environment and Natural Resources. Disclaimer While reasonable efforts have been made to ensure the contents of this publication are factually correct, the Department of Environment and Natural Resources makes no representations and accepts no responsibility for the accuracy, completeness or fitness for any particular purpose of the contents, and shall not be liable for any loss or damage that may be occasioned directly or indirectly through the use of or reliance on the contents of this publication.
    [Show full text]