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Dinosaur Species List E to M
Dinosaur Species List E to M E F G • Echinodon becklesii • Fabrosaurus australis • Gallimimus bullatus • Edmarka rex • Frenguellisaurus • Garudimimus brevipes • Edmontonia longiceps ischigualastensis • Gasosaurus constructus • Edmontonia rugosidens • Fulengia youngi • Gasparinisaura • Edmontosaurus annectens • Fulgurotherium australe cincosaltensis • Edmontosaurus regalis • Genusaurus sisteronis • Edmontosaurus • Genyodectes serus saskatchewanensis • Geranosaurus atavus • Einiosaurus procurvicornis • Gigantosaurus africanus • Elaphrosaurus bambergi • Giganotosaurus carolinii • Elaphrosaurus gautieri • Gigantosaurus dixeyi • Elaphrosaurus iguidiensis • Gigantosaurus megalonyx • Elmisaurus elegans • Gigantosaurus robustus • Elmisaurus rarus • Gigantoscelus • Elopteryx nopcsai molengraaffi • Elosaurus parvus • Gilmoreosaurus • Emausaurus ernsti mongoliensis • Embasaurus minax • Giraffotitan altithorax • Enigmosaurus • Gongbusaurus shiyii mongoliensis • Gongbusaurus • Eoceratops canadensis wucaiwanensis • Eoraptor lunensis • Gorgosaurus lancensis • Epachthosaurus sciuttoi • Gorgosaurus lancinator • Epanterias amplexus • Gorgosaurus libratus • Erectopus sauvagei • "Gorgosaurus" novojilovi • Erectopus superbus • Gorgosaurus sternbergi • Erlikosaurus andrewsi • Goyocephale lattimorei • Eucamerotus foxi • Gravitholus albertae • Eucercosaurus • Gresslyosaurus ingens tanyspondylus • Gresslyosaurus robustus • Eucnemesaurus fortis • Gresslyosaurus torgeri • Euhelopus zdanskyi • Gryponyx africanus • Euoplocephalus tutus • Gryponyx taylori • Euronychodon -
New Data on Small Theropod Dinosaurs from the Upper Jurassic Morrison Formation of Como Bluff, Wyoming, USA
Volumina Jurassica, 2014, Xii (2): 181–196 Doi: 10.5604/17313708 .1130142 New data on small theropod dinosaurs from the Upper Jurassic Morrison Formation of Como Bluff, Wyoming, USA Sebastian G. DALMAN1 Key words: dinosaurs, Theropoda, Upper Jurassic, Morrison Formation, Como Bluff, Wyoming, western USA. Abstract. In 1879, Othniel C. Marsh and Arthur Lakes collected in the Upper Jurassic Morrison Formation Quarry 12 at Como Bluff, Wyoming, USA, several isolated axial and appendicular skeletal elements of small theropod dinosaurs. Since the discovery the specimens remained unnoticed for over a century. The skeletal remains of small theropods are rare at Como Bluff and throughout the Morrison Forma- tion. Their bones are delicately constructed, so they are not as well-preserved as the bones of large-bodied theropods. The bones of small theropods described here were found mixed with isolated crocodile teeth and turtle shells. Comparison of the skeletal materials with other known theropods from the Morrison Formation reveals that some of the bones belong to a very small juvenile Allosaurus fragilis and Tor vosaurus tanneri and also to a new ceratosaur taxon, here named Fosterovenator churei, whereas the other bones represent previously unidentified juvenile taxa of basal tetanuran and coelurid theropods. The discovery and description of these fossil materials is significant because they provide important information about the Upper Jurassic terrestrial fauna of Quarry 12, Como Bluff, Wyoming. The presence of previously unidentified theropod taxa in the Morrison Formation indicates that the diversity of basal tetanuran and coelurid theropods may have been much greater than previously expected. Although the fossil material here described is largely fragmentary, it is tenable that theropods of different clades co-existed in the same ecosystems at the same time and most likely competed for the same food sources. -
Strength Indicator Values of Theropod Long Bones, with Comments on Limb Proportions and Cursorial Potential
GAIA N' 15, LlSBONLISBON, DEZEMBROIDECEMBER 1998, pp. 241-255 (ISSN: 0871-5424) STRENGTH INDICATOR VALUES OF THEROPOD LONG BONES, WITH COMMENTS ON LIMB PROPORTIONS AND CURSORIAL POTENTIAL Per CHRISTIANSEN Zoological Museum, Department of Vertebrates. Universitetsparken 15, OK 2100 COPEN HAG EM 0. DENMARK E-mail: [email protected] ABSTRACT: Body mass and strength indicator values of the three hindlimb long bones have been calculated for a large number oftheropod dinosaurs and compared to extant mammals of varying size and locomotory capability_Small to medium sized theropods have strength indicator values comparable to fast-moving ungulates and carnivorans, whereas all large genera have considerably lower strength indicator values, roughly comparable to elephants and hippopotamuses. This suggests that their locomotory potential was reduced compared to the smaller forms. Limb bone ratios of a large number of extant mammals clearly differen tiate fast-moving forms, classified from their anatomy as subcursorial or cursorial, from forms capable of less rapid locomotion, classified as graviportal and mediportal. Limb bone ratios for theropods, however, somewhat contradict the above, as all theropods group among subcursorial mammals_ Calculations on estimated peak locomotory performance in dicates that even large theropods could have been fast moving without having to include a suspended phase in the stride, thus not subjecting their appendicular anatomy to large amounts of stress, due to their very long limbs. INTRODUCTION However, COOMBS (1978) analyzed a long list of anatomical characters in various tetrapods and Theropod dinosaurs were the only undoubtedly found that a number of these had probably de carnivorous terrestrial tetrapods for most of the veloped convergently and were found in all forms ca Mesozoic that, at least theoretically, were large and pable of fast locomotion. -
An Upper Jurassic Theropod Dinosaur from the Section 19 Mine, Morrison Formation, Grants Uranium District
New Mexico Geological Society Guidebook, 54th Field Conference, Geology of the Zuni Plateau, 2003, p. 309-314. 309 AN UPPER JURASSIC THEROPOD DINOSAUR FROM THE SECTION 19 MINE, MORRISON FORMATION, GRANTS URANIUM DISTRICT ANDREW B. HECKERT1, JUSTIN A. SPIELMANN2, SPENCER G. LUCAS1, RICHARD ALTENBERG1 AND DANIEL M. RUSSELL1 1New Mexico Museum of Natural History, 1801 Mountain Road NW, Albuquerque, NM 87104-1375; 2Dartmouth College, Hinman Box 4571, Hanover, NH 03755 ABSTRACT.—Known vertebrate fossil occurrences from Jurassic strata in the mines in the Grants uranium district are few. Here, we describe fragmentary teeth, skull, and jaw elements that probably pertain to the theropod dinosaur Allosaurus. These fossils were recovered from the Upper Jurassic Salt Wash (=Westwater Canyon) Member of the Morrison Formation in the underground workings of the Section 19 mine near Grants, New Mexico. If properly identified, these fossils are the stratigraphically lowest occurrence (LO) of Allosaurus in New Mexico and one of the oldest records of Allosaurus in the Morrison Formation. INTRODUCTION Miners in the Grants uranium district doubtless encountered dinosaur bones relatively frequently, but very few of these dis- coveries were ever documented (Chenoweth, 1953; Smith, 1961) and even fewer were reposited in museums (Hunt and Lucas, 1993; Lucas et al., 1996). In this paper, we document fragmen- tary tooth-bearing dinosaur bones and associated teeth from the Morrison Formation Section 19 mine and discuss their biostrati- graphic significance (Fig. 1). Throughout this paper, NMMNH refers to the New Mexico Museum of Natural History and Sci- ence, Albuquerque, and UMNH refers to the Utah Museum of Natural History, Salt Lake City. -
Caudipteryx As a Non-Avialan Theropod Rather Than a Flightless Bird
Caudipteryx as a non−avialan theropod rather than a flightless bird GARETH J. DYKE and MARK A. NORELL Dyke, G.J. and Norell, M.A. 2005. Caudipteryx as a non−avialan theropod rather than a flightless bird. Acta Palaeontolo− gica Polonica 50 (1): 101–116. Caudipteryx zoui is a small enigmatic theropod known from the Early Cretaceous Yixian Formation of the People’s Re− public of China. From the time of its initial description, this taxon has stimulated a great deal of ongoing debate regarding the phylogenetic relationship between non−avialan theropods and birds (Avialae) because it preserves structures that have been uncontroversially accepted as feathers (albeit aerodynamically unsuitable for flight). However, it has also been pro− posed that both the relative proportions of the hind limb bones (when compared with overall leg length), and the position of the center of mass in Caudipteryx are more similar to those seen in extant cusorial birds than they are to other non−avialan theropod dinosaurs. This conclusion has been used to imply that Caudipteryx may not have been correctly in− terpreted as a feathered non−avialan theropod, but instead that this taxon represents some kind of flightless bird. We re− view the evidence for this claim at the level of both the included fossil specimen data, and in terms of the validity of the re− sults presented. There is no reason—phylogenetic, morphometric or otherwise—to conclude that Caudipteryx is anything other than a small non−avialan theropod dinosaur. Key words: Dinosauria, Theropoda, Avialae, birds, feathers, Yixian Formation, Cretaceous, China. Gareth J. -
New Information on Segisaurus Halli, a Small Theropod Dinosaur from the Early Jurassic of Arizona
Journal of Vertebrate Paleontology 25(4):835–849, December 2005 © 2005 by the Society of Vertebrate Paleontology NEW INFORMATION ON SEGISAURUS HALLI, A SMALL THEROPOD DINOSAUR FROM THE EARLY JURASSIC OF ARIZONA MATTHEW T. CARRANO1*, JOHN R. HUTCHINSON2, and SCOTT D. SAMPSON3 1Department of Paleobiology, Smithsonian Institution, P.O. Box 37012, MRC 121, Washington, DC 20013-7012, U.S.A., [email protected]; 2Structure and Motion Laboratory, The Royal Veterinary College, University of London, North Mymms, Hatfield, Hertfordshire, AL9 7TA, United Kingdom, [email protected]; 3Utah Museum of Natural History and Department of Geology and Geophysics, 1390 East Presidents Circle, University of Utah, Salt Lake City, UT 84112-0050, U.S.A., [email protected] ABSTRACT—Here we redescribe the holotype and only specimen of Segisaurus halli, a small Early Jurassic dinosaur and the only theropod known from the Navajo Sandstone. Our study highlights several important and newly recognized features that clarify the relationships of this taxon. Segisaurus is clearly a primitive theropod, although it does possess a tetanuran-like elongate scapular blade. Nonetheless, it appears to be a coelophysoid, based on the presence of a pubic fenestra, a long and ventrally curved pubis, and some pelvic (and possibly tarsal) fusion. Segisaurus does possess a furcula, as has now been observed in other coelophysoids, thus strengthening the early appearance of this ‘avian’ feature. The absence of an external fundamental system in bone histology sections and the presence of sutural contact lines in the caudal vertebrae, scapulocoracoid, and (possibly) between the pubis and ischium support the inference that this specimen is a subadult, neither a true juvenile nor at full skeletal maturity. -
The Locomotor and Predatory Habits of Unenlagiines (Theropoda, Paraves)
bioRxiv preprint doi: https://doi.org/10.1101/553891; this version posted February 18, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY 4.0 International license. 1 The locomotor and predatory habits of unenlagiines (Theropoda, Paraves): 2 inferences based on morphometric studies and comparisons with Laurasian 3 dromaeosaurids 4 5 6Federico A. Gianechini1*, Marcos D. Ercoli2 and Ignacio Díaz-Martínez3 7 8 9 101Instituto Multidisciplinario de Investigaciones Biológicas (IMIBIO), CONICET-Universidad 11Nacional de San Luis, Ciudad de San Luis, San Luis, Argentina. 122Instituto de Ecorregiones Andinas (INECOA), Universidad Nacional de Jujuy-CONICET, 13IdGyM, San Salvador de Jujuy, Jujuy, Argentina. 143Instituto de Investigación en Paleobiologia y Geología (IIPG), CONICET-Universidad 15Nacional de Río Negro, General Roca, Río Negro, Argentina. 16 17 18 19 20*Corresponding author 21E-mail: [email protected] 22 23 24 25 26 1 1 2 bioRxiv preprint doi: https://doi.org/10.1101/553891; this version posted February 18, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY 4.0 International license. 27Abstract 28 29Unenlagiinae is mostly recognized as a subclade of dromaeosaurids. They have the modified 30pedal digit II that characterize all dromeosaurids, which is typically related to predation. 31However, derived Laurasian dromaeosaurids (eudromaeosaurs) differ from unenlagiines in 32having a shorter metatarsus and pedal phalanx II-1, and more ginglymoid articular surfaces in 33metatarsals and pedal phalanges. -
The Phylogeny of Ceratosauria (Dinosauria: Theropoda)
Journal of Systematic Palaeontology 6 (2): 183–236 Issued 23 May 2008 doi:10.1017/S1477201907002246 Printed in the United Kingdom C The Natural History Museum The Phylogeny of Ceratosauria (Dinosauria: Theropoda) Matthew T. Carrano∗ Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560 USA Scott D. Sampson Department of Geology & Geophysics and Utah Museum of Natural History, University of Utah, Salt Lake City, UT 84112 USA SYNOPSIS Recent discoveries and analyses have drawn increased attention to Ceratosauria, a taxo- nomically and morphologically diverse group of basal theropods. By the time of its first appearance in the Late Jurassic, the group was probably globally distributed. This pattern eventually gave way to a primarily Gondwanan distribution by the Late Cretaceous. Ceratosaurs are one of several focal groups for studies of Cretaceous palaeobiogeography and their often bizarre morphological develop- ments highlight their distinctiveness. Unfortunately, lack of phylogenetic resolution, shifting views of which taxa fall within Ceratosauria and minimal overlap in coverage between systematic studies, have made it difficult to explicate any of these important evolutionary patterns. Although many taxa are fragmentary, an increase in new, more complete forms has clarified much of ceratosaur anatomy, allowed the identification of additional materials and increased our ability to compare specimens and taxa. We studied nearly 40 ceratosaurs from the Late Jurassic–Late Cretaceous of North and South America, Europe, Africa, India and Madagascar, ultimately selecting 18 for a new cladistic analysis. The results suggest that Elaphrosaurus and its relatives are the most basal ceratosaurs, followed by Ceratosaurus and Noasauridae + Abelisauridae (= Abelisauroidea). -
'Coelurosaurs' (Dinosauria, Theropoda) from the Late Jurassic of Tanzania
Geol. Mag. 142 (1), 2005, pp. 97–107. c 2005 Cambridge University Press 97 DOI: 10.1017/S0016756804000330 Printed in the United Kingdom Post-cranial remains of ‘coelurosaurs’ (Dinosauria, Theropoda) from the Late Jurassic of Tanzania OLIVER W. M. RAUHUT* Institut fur¨ Palaontologie,¨ Museum fur¨ Naturkunde, Humboldt-Universitat,¨ Invalidenstraße 43, D-10115 Berlin, Germany (Received 19 February 2004; accepted 7 September 2004) Abstract – Small theropod post-cranial material from Tendaguru, Tanzania, the only known Late Jurassic theropod locality in the Southern Hemisphere, is reviewed. Material originally described as ‘coelurosaurs’ includes at least one taxon of basal tetanuran and one taxon of small abelisauroid. Together with the abelisauroid Elaphrosaurus and the presence of a larger ceratosaur in Tendaguru, this material indicates that ceratosaurs were an important faunal element of Late Jurassic East African theropod faunas. One bone furthermore shares derived characters with the holotype of the poorly known Middle Jurassic Australian theropod Ozraptor and allows the identification of the latter as the oldest known abelisauroid, thus indicating an early divergence of ceratosaurids and abelisauroids within ceratosaurs. Abelisauroids might have originated in Gondwana and represent important faunal elements of Cretaceous Gondwanan theropod faunas in general. Keywords: Jurassic, Gondwana, Theropoda, diversity, biogeography. 1. Introduction Whereas the expeditions resulted in the discovery of thousands of specimens of sauropods, stegosaurs and Our knowledge of theropod dinosaur faunas from ornithopods, theropods are rather poorly represented Gondwana has rapidly improved in the past twenty (Janensch, 1914; Zils et al. 1995). Apart from the years. The description of numerous new taxa and partial skeleton of Elaphrosaurus (Janensch, 1920, identification of new groups have greatly changed 1925, 1929), only isolated elements have been found, our understanding of Cretaceous theropod evolution and theropod diversity and faunal composition in the and biogeography (e.g. -
A New Phylogeny of the Carnivorous Dinosaurs
GAIA Nº 15, LISBOA/LISBON, DEZEMBRO/DECEMBER 1998, pp. 5-61 (ISSN: 0871-5424) A NEW PHYLOGENY OF THE CARNIVOROUS DINOSAURS Thomas R. HOLTZ, Jr. Department of Geology, University of Maryland. College Park, MARYLAND 20735. USA E-Mail: [email protected] ABSTRACT: The last several years have seen the discovery of many new theropod dinosaur taxa. Data obtained from these and from fragmentary forms not previously utilized in cladis- tic analyses are examined. An analysis of forty one primary ingroup taxa and 386 characters yielded a set of most parsimonious cladograms which preserves many previously discov- ered relationships (e.g., a basal split between Ceratosauria and Tetanurae; a carnosaur- coelurosaur clade Avetheropoda outside of more primitive "megalosaur" - grade teta- nurines; Dromaeosauridae as the sister taxon to birds, and so forth). The Middle Jurassic English Proceratosaurus was discovered to be a basal coelurosaur, as was (on less secure evidence) the Middle Jurassic Chinese Gasosaurus: these are among the oldest coeluro- saurs yet described. Several characters previously considered to be restricted to birds and other advanced coelurosaurs (e.g., furcula, semilunate carpal block) were found to be more broadly distributed among tetanurines. Other characters, once considered synapomor- phies for Avetheropoda (e.g., loss of metacarpal IV, possession of a pubic obturator notch) were found to be convergent between advanced carnosaurs and advanced coelurosaurs, lacking in the basal members of both clades. At least three (and possibly four) separate ori- gins for the arctometatarsalian pes were supported in this study. The mosaic of derived character state distributions for troodontids relative to the dromaeosaurid-bird clade, the tyrannosaurid-ornithomimosaur clade, and the therizinosauroid-oviraptorosaur clade sug- gests that relationships alternative to the most parsimonious found here may be supported in future studies. -
Cretaceous, Maastrichtian) of India
CONTRIBUTIONS FROM THE MUSEUM OF PALEONTOLOGY THE UNIVERSITY OF MICHIGAN VOL. 3 1, NO. I, PP. 1-42 August 15,2003 A NEW ABELISAURID (DINOSAURIA, THEROPODA) FROM THE LAMETA FORMATION (CRETACEOUS, MAASTRICHTIAN) OF INDIA JEFFREY A. WILSON, PAUL C. SERENO, SURESH SRIVASTAVA, DEVENDRA K. BHATT, ASHU KHOSLA AND ASHOK SAHNI MUSEUM OF PALEONTOLOGY THE UNIVERSITY OF MICHIGAN ANN ARBOR CONTRIBUTIONS FROM THE MUSEUM OF PALEONTOLOGY Philip D. Gingerich, Director This series of contributions from the Museum of Paleontology is a medium for publication of papers based chiefly on collections in the Museum. When the number of pages issued is sufficient to make a volume, a title page plus a table of contents will be sent to libraries on the Museum's mailing list. This will be sent to individuals on request. A list of the separate issues may also be obtained by request. Correspondence should be directed to the Publications Secretary, Museum of Paleontology, The University of Michigan, 1109 Geddes Road, Ann Arbor, Michigan 48 109-1079 ([email protected]). VOLS. 2-3 1: Parts of volumes may be obtained if available. Price lists are available upon inquiry. Text and illustrations 02003 by the Museum of Paleontology, University of Michigan A NEW ABELISAURID (DINOSAURIA, THEROPODA) FROM THE LAMETA FORMATION (CRETACEOUS, MAASTRICHTIAN) OF INDIA JEFFREY A. WILSON', PAUL C. SERE NO^, SURESH SRIVASTAVA3, DEVENDRA K. BHATT4, ASHU KHOSLA5 AND ASHOK SAHN15 Abstract - Many isolated dinosaur bones and teeth have been recovered from Cretaceous rocks in India, but associated remains are exceedingly rare. We report on the discovery of associated cranial and postcranial remains of a new abelisaurid theropod from latest Cretaceous rocks in western India. -
An Upper Jurassic Theropod Dinosaur from the Section 19 Mine, Morrison Formation, Grants Uranium District Andrew B
New Mexico Geological Society Downloaded from: http://nmgs.nmt.edu/publications/guidebooks/54 An Upper Jurassic theropod dinosaur from the Section 19 mine, Morrison Formation, Grants Uranium district Andrew B. Heckert, Justin A. Spielmann, Spencer G. Lucas, Richard Altenberg, and Daniel M. Russell, 2003, pp. 309-314 in: Geology of the Zuni Plateau, Lucas, Spencer G.; Semken, Steven C.; Berglof, William; Ulmer-Scholle, Dana; [eds.], New Mexico Geological Society 54th Annual Fall Field Conference Guidebook, 425 p. This is one of many related papers that were included in the 2003 NMGS Fall Field Conference Guidebook. Annual NMGS Fall Field Conference Guidebooks Every fall since 1950, the New Mexico Geological Society (NMGS) has held an annual Fall Field Conference that explores some region of New Mexico (or surrounding states). Always well attended, these conferences provide a guidebook to participants. Besides detailed road logs, the guidebooks contain many well written, edited, and peer-reviewed geoscience papers. These books have set the national standard for geologic guidebooks and are an essential geologic reference for anyone working in or around New Mexico. Free Downloads NMGS has decided to make peer-reviewed papers from our Fall Field Conference guidebooks available for free download. Non-members will have access to guidebook papers two years after publication. Members have access to all papers. This is in keeping with our mission of promoting interest, research, and cooperation regarding geology in New Mexico. However, guidebook sales represent a significant proportion of our operating budget. Therefore, only research papers are available for download. Road logs, mini-papers, maps, stratigraphic charts, and other selected content are available only in the printed guidebooks.