ZOOTAXA 77: 1-11 (2002) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA 77 Copyright © 2002 Magnolia Press ISSN 1175-5334 (online edition)

Nomenclatural and taxonomic notes on cyanescens (Ratze- burg, 1837), A. pratensis (Ratzeburg, 1837) and A. convexicollis Redtenbacher, 1849 (Coleoptera: : Agrilinae)

EDUARD JENDEK Institute of Zoology, Slovak Academy of Sciences, Dúbravská cesta 9, SK-842 06 Bratislava, Slovakia E-mail: [email protected]

Abstract

A nomenclatural analysis of names cited in synonymy of Agrilus cyanescens (Ratzeburg, 1837), A. pratensis (Ratzeburg, 1837) and A. convexicollis Redtenbacher, 1849 is presented. The availabi- lity and status of each name is examined, discussed and an updated list of subspecies, available and unavailable synonyms is given. Agrilus bidentulus Ganglbauer, 1890 is considered a valid species and excluded from the synonymy of A. cyanescens; A. pratensis praeclarus Krogerus, 1925 is revalidated as a subspecies with the new synonym fennicus Obenberger, 1936.

Key words: , nomenclature, synonymy, Coleoptera, Buprestidae, Agrilinae, Agrilus, Pa- laearctic Region

Introduction

This contribution belongs to the nomenclatural-taxonomic series of papers (Jendek 2002a, 2002b, 2002c) on Palaearctic species of the genus Agrilus Curtis, 1825 undertaken with the aim to clean up the synonymy from unavailable names. All names, whether available or unavailable, which were discovered in literature citations are treated herein. Although the primary effort was focused to treat the nomenclatural status of names, several taxo- nomic changes are performed in context with the changes in the nomenclatural status of particular names.

Material and methods

Abbreviations used: square brackets “[ ]” are used for my remarks and addenda; the mark “[sic!]” (= so, thus) denotes that a word, phrase, passage, etc., that may appear strange or incorrect has been written intentionally or has been quoted verbatim.

Accepted: 1 October 2002; published: 11 October 2002 1 ZOOTAXA Issues of nomenclature were resolved by consulting the fourth (ICZN, 1999) edition of 77 the International Code of Zoological Nomenclature. Articles of the Code cited herein are simply cited as, for example, “Article 74.5”. The analyzed names are divided into three groups. The names in each group are listed chronologically, then alphabetically. The names originally proposed for a species and subspecies or replacement names include the name, the author, date, and page number for the original description, followed by (in parentheses) the verbatim transcription from the original publication, when needed with my supplements in “[ ]”. The names originally proposed for an infrasubspecific entity are cited with the same attributes as the foregoing category, but in remarks, the availability of each name is treated and the corresponding Article of the Code is cited. The unavailable names origi- nally proposed for infrasubspecific entities are included in synonymy only in order to eliminate confusion or questions about their status. Nonetheless, they have no nomencla- tural status. The names that were never proposed as new (unpublished names, misidentification, incorrect subsequent spelling etc) are, nevertheless, considered names, although they were never proposed as a new name of taxon or a new replacement name proposed to replace junior homonym. Nevertheless, they have been cited in references as though they were synonyms. This practice is misleading and confusing. These names are cited herein only to resolve their status. Each name is listed in the form (name, author) in which it was firstly introduced, followed by (in parentheses) the verbatim transcription from the original pub- lication. In remarks, the author is cited, who is regarded the first introducing this name. Type localities are cited only for available names and they are quoted in form of the original citation. My complementary comments and updates are consistently given in “[ ]”. Localities have been updated according to the GEOnet Names Server providing access to the database of foreign geographic feature names of the National Imagery and Mapping Agency (NIMA).

Agrilus cyanescens (Ratzeburg, 1837)

cyanescens Ratzeburg, 1837: 54-55. (B[uprestis]. cyanescens Ill[iger]. [sic!])

Although Illiger is cited after cyanescens, Ratzeburg is the author of the taxon. Ratze- burg’s citation of Illiger’s name relates to his remark: “Cyanescens von Illiger in Berl. Mus. genannt”. Type locality: not given, but presumably Germany as indicated from the title: “Wäl- dern Preufsens”.

2 © 2002 Magnolia Press JENDEK available synonyms ZOOTAXA - caeruleus Rossi, 1792 77 - amabilis Gory & Laporte, 1837 - sulcaticeps Abeille de Perrin, 1869 - acuticornis Abeille de Perrin, 1897 - teriolensis Obenberger, 1916 unavailable synonyms - virens Schilsky, 1888 - combinator Obenberger, 1925 - subcarpathicus Roubal, 1929 - cuprescens Fleischer, 1930 - pseudroberti Fleischer, 1930 subspecies - ssp. johanidesi Niehuis, 1999

Available names proposed for species and subspecies or replacement names caeruleus Rossi, 1792: 62. (B[uprestis]. caerulea [sic!]) This name is the junior primary homonym of Buprestis caerulea Thunberg, 1789 (now in Meliboeus) and Buprestis caerulea Olivier, 1790 (now in Psiloptera Dejean, 1833); and senior subjective synonym of Agrilus cyanescens (Ratzeburg, 1837). Type locality: “in planta silvae Pisanae [= Pisa province, Tuscany, CW Italy]”. amabilis Gory & Laporte, 1837: 52-53. (Agrilus Amabilis) The first synonymy of this name as the junior subjective synonym of A. caeruleus was by Kiesenwetter (1857: 141, 142). The lectotype designation by Jendek (1998: 317). Type locality: “Saxe [= Saxonia, E Germany]”. sulcaticeps Abeille de Perrin, 1869: 79-80. (Agrilus sulcaticeps) The first synonymy of this name as the junior subjective synonym of A. caeruleus was by Abeille de Perrin (1897: 9). Type locality: “Briançon [44°54' N, 6°37' E] (Hautes-Alpes) SE France]”. bidentulus Ganglbauer, 1890: 22, 31. species revalidated This species was described from China, Gansu and was synonymized by Alexeev (1959: 26) as the junior subjective synonym of A. caeruleus. This taxon belongs to the cyanescens species group but it is not conspecific with A. cyanescens and it shall be removed from this synonymy.

AGRILUS © 2002 Magnolia Press 3 ZOOTAXA acuticornis Abeille de Perrin, 1897: 3, 10. ([Agrilus] acuticornis n. sp.) 77 The first time synonym of this name as the junior subjective synonym of A. cyane- scens was by Niehuis & Tezcan (1993: 1, 22). Type locality: “Caucase arménien [Armenia]”.

teriolensis Obenberger, 1916: 275. (Agrilus artemisiae subsp. teriolensis m. n. sp. [sic!]) The name teriolensis was proposed as a subspecies of A. artemisiae Brisout de Barneville, 1863. Curletti (1994: 167) studied the type and synonymized it as the junior subjective synonym of A. cyanescens. Type locality: “Tirol [SW Austria]”.

johanidesi Niehuis, 1999: 35-41. (Agrilus cyanescens johanidesi n. ssp.) Proposed as the subspecies of A. cyanescens. Type locality: “Syria, Quatanã –Burqush env. [33°27' N, 36°02' E]”.

Names originally proposed for an infrasubspecific entities

virens Schilsky, 1888: 187. (Agrilus coerulus [sic!] v. virens) The name virens was proposed as a variety of A. caeruleus. The content of the work unambiguously revealed that names were proposed as an infrasubspecific entity, since the taxon was very briefly diagnosed only by differences in color: “Oberseite grün”. This name was never adopted as a valid name of a species or a subspecies. Following Article 45.6.4, such name should be deemed infrasubspecific and unavailable from the original publication.

combinator Obenberger, 1925: 102. (Agrilus fissifrons var. combinator m. n. var.) The type specimen was examined and is conspecific with A. cyanescens. The content of the work unambiguously revealed that names were proposed as an infrasubspecific entity, since the taxon was very briefly diagnosed only by differences in color: “A typo thorace ac capite aeneobrunneis, elytris clare viridibus differt” and Obenberger, in subse- quent references, treated it as a mere aberration. This name was never adopted as a valid name of a species or a subspecies. Following Article 45.6.4, such name should be deemed infrasubspecific and unavailable from the original publication.

subcarpathicus Roubal, 1929: 192. (Agrilus Kyselyi formam hanc a. subcarpathica n. puto) The type specimen was examined and is conspecific with A. cyanescens. It was origi- nally proposed as an aberration and thus an unavailable name (Article 45.6.2).

4 © 2002 Magnolia Press JENDEK cuprescens Fleischer, 1930: 207. ([Agrilus coeruleus] [sic!] a. cuprescens Fleisch.) ZOOTAXA Obenberger (1936: 1004) cited it in combination with A. kyselyi Obenberger, 1924. It 77 was originally proposed as an aberration and thus is an unavailable name (Article 45.6.2). pseudroberti Fleischer, 1930: 207. ([Agrilus coeruleus] [sic!] a. pseudroberti [sic!] Fleisch.) It was originally proposed as an aberration and thus is an unavailable name (Article 45.6.2).

Unpublished names and misidentification amethystinus Schönherr ([Buprestis] Amethystina Oliv[ier]. [sic!]) Schönherr (1817: 258) did not describe this species as new, as he attributed it to Oli- vier. Schönherr erroneously considered Buprestis amethystina Olivier, 1790 (now in Meli- boeus) to be conspecific with Buprestis caerulea Rossi, 1792 (now junior subjective syn- onym of A. cyanescens). It was first introduced in the synonymy of A. caeruleus by Gemminger & Harold (1869: 1438). Misidentification. cyaneus Lacordaire (A[grilus]. cyaneus Ol[ivier]. [sic!]) Lacordaire in Boisduval & Lacordaire (1835: 612-613) did not describe this species as new; he attributed it to Olivier1. This name was introduced by Obenberger (1936: 975) in the synonymy of A. caeruleus. Misidentification. cyaneus Gory & Laporte (Agrilus cyaneus, Olivier [sic!]) Gory & Laporte (1837: 52) did not describe this species as new; he attributed it to Olivier1. It was first introduced in the synonymy of A. caeruleus by Kiesenwetter (1857: 140, 141). Misidentification.

Agrilus pratensis (Ratzeburg, 1837) pratensis Ratzeburg, 1837: 56-57. (B[uprestis]. pratensis)

Type locality: not given, but presumably Germany. This species was described from speci- men provided on loan by the German entomologist E. G. Germar: “Indessen gehört das von H. Germar mir als B. linearis geliehene Stück zu einer ganz anderen Art B. pratensis”.

1. Olivier (1790: 91) also did not describe this species as new; he attributed it to Fabricius. Actually, Olivier only cited Buprestis cyanea Fabricius, 1775 (now in Phaenops Dejean, 1833).

AGRILUS © 2002 Magnolia Press 5 ZOOTAXA available synonyms 77 - robertii Chevrolat, 1838 - pseudocoeruleus Obenberger, 1930

unavailable synonyms - strigicollis Rey, 1891 - confusus Schaefer, 1936 - paracyaneus Schaefer, 1937 - aeneicolor Schaefer, 1946 - confusinella Schaefer, 1949

subspecies - ssp. praeclarus Krogerus, 1925 revalidated name available synonym - fennicus Obenberger, 1936 syn. nov.

-ssp. djukini Obenberger, 1935 available synonym - chankae Obenberger, 1935

- ssp. meridionalis Cobos, 1986

Available names proposed for species and subspecies or replacement names

robertii Chevrolat, 1838: 89-90. (Agrilus Robertii [sic!]) A junior subjective synonym of A. pratensis. It was first synonymized by Kiesenwet- ter (1857: 146). Type locality: “Belgia [= Belgium] ... des environs de Liège [50°38' N, 5°34' E]”.

pseudocoeruleus Obenberger, 1930: 112. (Agrilus pseudocoeruleus m. n. sp.) A junior subjective synonym of A. pratensis according to Alexeev & Volkovitsh (1989: 319). The lectotype designation was by Jendek (1994: 10-11). Type locality: “Asia centralis: Sefid-rud [in Iran ?]”.

praeclarus Krogerus, 1925: 73, 75-76. ([Agrilus] Robertii var. praeclarus n. var.) revali- dated name The name praeclarus was proposed as a variety of A. robertii. This taxon was briefly diagnosed only by differences in color: “Heller gefärbt als die Nominalform. Halsschild goldgrün, Flügeldecken blau”, but it is not considered unambiguously proposed for an infrasubspecific entity, because it was described from northernmost distribution area of A. pratensis and this taxon was deemed to be a subspecies of A. pratensis under the name fen- nicus Obenberger 1936. See also comments under fennicus Obenberger. Type locality: “im südlichen Finnland, St. Ylöjärvi [61°33' N, 23°35' E]”.

6 © 2002 Magnolia Press JENDEK fennicus Obenberger, 1936: 1029. ([Agrilus Roberti] [sic!] ab. [sic!] fennicus nom. nov.) ZOOTAXA syn. nov. 77 The name praeclarus Krogerus is the junior primary homonym of praeclarus Rey, 1891 proposed as a variety of A. viridis (Linné, 1758). The name praeclarus Krogerus was substituted by Obenberger in 1936 by the replacement name fennicus. This taxonomic act was mistakenly repeated by Obenberger also in 1937. Schaefer (1949: 392) erected fenni- cus to the subspecies level. I consider praeclarus Rey to be an unavailable name (this name will be treated in a future contribution on A. viridis) and thus praeclarus Krogerus is deemed a valid subspecies of A. pratensis and fennicus Obenberger is junior objective syn- onym of praeclarus Krogerus. djukini Obenberger, 1935: 169. (Agrilus Roberti [sic!] subsp. Djukini n. ssp.) Described as a subspecies of A. robertii. The lectotype designation was by Jendek (1994: 11). Type locality: “s. Trojckoe [= Troitskoye, 44°51' N, 132°02' E], oz. Chanka [Khanka lake], Primors. [Primorskiy Kray, Russia]”. The type locality precised according to the lec- totype locality data. chankae Obenberger, 1935: 169. (Agrilus betuleti subsp. Chankae n. ssp.) Described as a subspecies of A. betuleti (Ratzeburg, 1837). The proposal of a junior subjective synonym of A. pratensis djukini and the lectotype designation was given by Jendek (1994: 11). Type locality: “Chanka [Khanka lake] Ussuri Kamen-Rybolov [44°45' N, 132°04' E]”. The type locality precised according to the lectotype locality data. meridionalis Cobos, 1986: 241-243, 316. ([Agrilus (Anambus)] pratensis meridionalis ssp. nov.) Proposed as a subspecies of A. pratensis. Type locality: “Balenyá [= 41°49' N, 2°13' E, Catalonia, Spain]”.

Names originally proposed for an infrasubspecific entities strigicollis Rey, 1891: 19. (Agrilus pratensis ... variété strigicollis R[ey].) The name strigicollis was proposed as a variety of A. pratensis. The content of the work unambiguously revealed that name was proposed for an infrasubspecific entity, since the taxon was briefly diagnosed only by a smaller size and denser structure of pronotum: “est moindre, avec les rides du prothorax paraissant un peu moins serrées” without the citation of type locality. This name was never adopted as a valid name of a species or a subspecies. Following Article 45.6.4, such a name should be deemed infrasubspecific and unavailable from the original publication.

AGRILUS © 2002 Magnolia Press 7 ZOOTAXA confusus Schaefer, 1936: 117. (Agrilus Roberti ab. confusus nov.) 77 Proposed as an aberration of A. roberti and thus an unavailable name (Article 45.6.2). See also confusinella Schaefer, 1949.

paracyaneus Schaefer, 1937: 84. ([Agrilus pratensis] ab. paracyaneus nov.) Proposed as an aberration of A. pratensis and thus an unavailable name (Article 45.6.2).

aeneicolor Schaefer, 1946: 73. (Agrilus pratensis ab. aeneicolor nov.) Proposed as an aberration of A. pratensis and thus an unavailable name (Article 45.6.2).

confusinella Schaefer, 1949: 391, 392, 490. ([Agrilus pratensis] ab. confusinella nom. nov.) Proposed as a replacement name for an unavailable name A. pratensis a. confusus Schaefer, 1936 not A. confusus Waterhouse, 1889 and thus an unavailable name.

Unpublished names and misidentification

auricollis Sturm ([Agrilus] auricollis. St[urm]) Sturm (1843: 63) cited the name “A. auricollis Sturm” in the catalogue list of species. Sturm never described this taxon as new. Gemminger & Harold (1869: 1445) listed “auri- collis Sturm” in synonymy of A. robertii. Unpublished name.

linearis Gory & Laporte (Agrilus linearis. Fabr[icius]. [sic!]) Gory & Laporte (1837: 55-56) misidentified A. pratensis under the name linearis according to Kiesenwetter (1857: 146, 147), who attributed linearis to Gory & Laporte and listed it in the synonymy of A. pratensis. Gory & Laporte did not describe the species as new; they attributed it to Fabricius. Misidentification.

linearis Paykull (Bupr[estis]. Linearis ... Fabr[icius]) Paykull (1799: 226-227) did not propose linearis as new, he attributed it to Fabricius. Schönherr (1817: 245) listed linearis Paykull in the synonymy of “Buprestis olivacea Gyllenhal”; Gory & Laporte (1837: 53) in the synonymy of “Agrilus pavidus Fabr[icius]” and Kiesenwetter (1857: 146, 147) in the synonymy of A. pratensis. Misidentification.

Agrilus convexicollis Redtenbacher, 1849

convexicollis Redtenbacher, 1849: 285. ([Agrilus] (Redt.) convexicollis)

8 © 2002 Magnolia Press JENDEK Type locality: Not given, but presumably Austria as indicated by title of the work “Fauna ZOOTAXA Austriaca”. 77 available synonym - mancinii Obenberger, 1927

Available names proposed for species and subspecies mancinii Obenberger, 1927: 84-85. (Agrilus convexicollis ssp. Mancinii m. n. ssp.) Proposed as a subspecies of A. convexicollis and as the junior subjective synonym of A convexicollis by Curletti (1987: 151). Type locality: “Italia: Poggio Cavallo [42°45' N, 11°09' E], Dint. Grossetto”.

Unpublished name brevitarsis Erichson, “[Agrilus] brevitarsis Erichs. i. litt.” An unavailable name first introduced by Gemminger & Harold (1869: 1438) in the synonymy of A. convexicollis. Unpublished name.

Acknowledgements

My sincere thanks are due to Charles L. Bellamy (Sacramento, California, USA) for lin- guistic revision of the manuscript. This work was partially supported by Grant No. 2/7176/ 20 from the Slovak Grant Agency for Science.

References

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