Notes on New and Poorly Known Palaearctic Species of the Genera
Total Page:16
File Type:pdf, Size:1020Kb
Bull. Br. arachnol. Soc. (2004) 13 (2), 33–40 33 Notes on new and poorly known Palaearctic Stockholm, Sweden (Dr T. Kronestedt); species of the genera Neon, Sitticus and Synageles YMTU=personal collection of Dr Yuri Marusik, (Araneae: Salticidae) temporarily kept in Zoological Museum, Turku University, Finland; ZMTU=Zoological Museum, Dmitri V. Logunov University of Turku, Turku, Finland (Dr S. Koponen); Manchester Museum, ZMUM=Zoological Museum, Moscow State University of Manchester, University, Moscow, Russia (Dr K. G. Mikhailov). Oxford Road, Manchester, M13 9PL Abbreviations used in the text: AME=anterior median eyes, ap=apical, d=dorsal, Fm=femur, Summary Mt=metatarsus, PLE=posterior lateral eyes, pr=prolateral, Pt=patella, rt=retrolateral, Tb=tibia, Two new species are diagnosed, figured and described: v=ventral. The sequence of leg segment measurements is Neon kovblyuki sp. n. (_\; Ukraine: the Crimea) and Synageles persianus sp. n. (_\; Azerbaijan and Iran). The as follows: femur+patella+tibia+metatarsus+tarsus. male of Sitticus rivalis Simon, 1937 is figured for the first For the leg spination the system adopted is that used by time; furthermore, this species is removed from synonymy Ono (1988). All measurements are in mm. with S. striatus Emerton, 1911. Neon pusio Simon, 1937 is synonymised with Neon convolutus Denis, 1937. Neon (Dicroneon) kovblyuki sp. n. (Figs. 1–6) Introduction Types: Holotype _ (ZMUM), Ukraine, the Crimea, Cape Martyan Reserve (44(30#N, 34(15#E), 1–70 m Although the Salticidae of northern and central a.s.l., 10 March 2002, Y. M. Marusik. Paratypes: 4\ Europe are relatively well-known, those from southern (ZMUM), together with holotype. Europe and especially the Mediterranean region remain Etymology: The species is dedicated to my colleague, poorly known. Almost any new collection from these Mr Nikolai M. Kovblyuk (Ukraine, Simferopol’), who regions still reveals new species (e.g. Logunov, 2001; has been successfully studying the Crimean spider fauna Logunov & Kronestedt, 2003); furthermore, known and who organised the trip during which this new species can present problems relating to the con- species was collected. specificity of specimens collected from different parts of Diagnosis: This new species belongs to the subgenus a species’ range (e.g. Kronestedt & Logunov, 2003). Dicroneon and is closest to Neon levis (Simon, 1871), Many of the old names by Lucas, Simon, Denis and from which it can be easily separated by the shorter others, especially from the genera Euophrys, Neon, embolus (cf. Fig. 2 and fig. 223 in Z~abka, 1997) and the Aelurillus, etc., of which descriptions were based on larger, round primary receptacles (Fig. 4) (long and single sexes, are still known only from the original tube-shaped in N. levis; see fig. 227 in Z~abka, 1997). descriptions, with little understanding of how they relate Distribution: The type locality only. to newly collected material. A thorough revision of the Description: Male (holotype): Carapace 0.77 long, Mediterranean salticid fauna is clearly required. The aim 0.59 wide, 0.40 high at PLE. Ocular field 0.54 long, of the present paper is to examine some of these taxo- 0.60 wide anteriorly, 0.64 wide posteriorly. Diameter of nomic problems related to southern European jumping AME 0.19. Chelicerae 0.14 long. Clypeus 0.03 high. spiders. Abdomen 0.63 long, 0.53 wide. Length of leg segments: I 0.43+0.19+0.29+0.17+0.18; II 0.36+0.17+0.21+ 0.16+0.16; III 0.36+0.16+0.21+0.20+0.16; IV 0.44+ Material and methods 0.19+0.33+0.30+0.20. Leg spination: I: Tb v 2-2-0, Mt Specimens for this study were borrowed from or are v 2-2ap. II: Tb v 1-0, Mt v 2-2ap. III and IV spineless. deposited in the following museums or personal collec- Coloration: carapace yellow in centre and yellow-brown tions: BMNH=Department of Entomology, Natural on sides and margins, with wide black areas around History Museum, London, UK (Ms J. Beccaloni); eyes. Clypeus narrow, brown, hairless. Sternum yellow. HNHM=Hungarian Natural History Museum, Maxillae and labium yellow-brown. Chelicerae yellow, Budapest, Hungary (Mr T. Szûts); HECO=Hope with brown anterior sides. Abdomen yellow, with 3 Entomological Collections, Oxford, UK (Mr J. Hogan); longitudinal brown stripes on dorsum and similar brown HUJI=Hebrew University of Jerusalem (Zoological stripe on each side; venter yellow. Book-lung covers Department), Israel (Dr G. Levy); MCZH=Museum of yellow, tinged with grey. Spinnerets yellow. Leg I dark- Comparative Zoology, Harvard University, Cambridge, est: femora yellow, with pro- and retrolateral brown USA (Ms L. Leibensperger); MMUM=Manchester stripes; patellae, tibiae and metatarsi brown, with yellow Museum, University of Manchester, Manchester, UK dorsal stripes; tarsi yellow. Palps yellow-brown. Palpal (Dr D. V. Logunov); MNHN=Muséum national structure as in Figs. 1–3. d’Histoire naturelle, Paris, France (Dr C. Rollard); Female (paratype): Carapace 0.89 long, 0.64 wide, PCAM=personal collection of Dr Mark Alderweireldt 0.43 high at PLE. Ocular field 0.53 long, 0.66 wide (Gent, Belgium); PCKT=personal collection of Dr anteriorly, 0.67 wide posteriorly. Diameter of AME Konrad Thaler (Innsbruck, Austria); PCRS=personal 0.21. Chelicerae 0.17 long. Clypeus 0.04 high. Abdomen collection of Dr Tony Russell-Smith (Kent, 0.99 long, 0.73 wide. Length of leg segments: I 0.46+ UK); SMNH=Swedish Museum of Natural History, 0.21+0.31+0.21+0.19; II 0.41+0.21+0.21+0.19+0.17; 34 New and poorly known Palaearctic Salticidae III 0.41+0.19+0.24+0.22+0.19; IV 0.51+0.23+0.36+ Zoological Society of London for 1936, which was 0.36+0.24. Leg spination: I: Tb v 2-2-0, Mt v 2-2ap. II: published in January 1937 [apparently earlier than the Tb v 1-1, Mt v 2-2ap. III and IV spineless. Coloration as posthumous work by Simon (1937)]. for male; general appearance as in Fig. 6. Epigyne and Both N. convolutus and N. pusio could also be junior spermathecae as in Figs. 4–5. synonyms of the well-known Euro-Central Asian species Neon rayi (Simon, 1875), but the tiny differences in the structure of the tip of the tibial apophysis shown re- cently by some authors (e.g. Roberts, 1998: 207) and the Neon (Dicroneon) convolutus Denis, 1937 (Figs. 7–9) absence of adequate comparative material do not allow Neon convolutus Denis, 1937: 1055, pl. 5, figs. 7–9 (D_; _ holotype in me to confirm this assumption. Nevertheless, it should MNHN; examined). be noted that the _ illustrated by Roberts (1998) under Neon pusio Simon, 1937: 1183, 1254 (D_, _ syntypes in MNHN; the name N. pusio is slightly different from the type of examined). New synonymy. this species (cf. Fig. 9) and from that of N. convolutus For a complete set of taxonomic references see Platnick (2002). (cf. Figs. 7, 8) in having a larger distal coil of the Type: Holotype _ (MNHN), Algeria, Zouagha for- embolus and a larger distal embolic groove of the est. cymbium, which occupies nearly half of the cymbial Comments: Having re-examined the _ holotype of length (no more than a third in N. convolutus, as in true N. convolutus from Algeria (Figs. 7–8) and the two _ N. rayi), and hence it may belong to a different species syntypes of N. pusio from France (Fig. 9), I came to the (whereas N. convolutus itself may be a junior synonym of opinion that both species names should be synonymised. N. rayi). Being short of comparative material of N. rayi, The males of both species are characterised by an I cannot evaluate the variation of these characters and identical conformation of the copulatory organs (viz. the thus prove or reject the above suggestion. The problem long coiled embolus and the tibial apophysis with its tip should be addressed later when more material becomes slightly pointed and bent ventrad, cf. Figs. 8, 9). The available. name N. convolutus seems to have priority, as Denis’ Also, it is worth noting the rather surprising fact that paper appeared in part IV of the Proceedings of the Denis (1937: 1055) described N. convolutus from a single Figs. 1–6: Neon kovblyuki sp. n. (_ holotype, \ paratype). 1 Male palp, mesal view; 2 Ditto, ventral view; 3 Ditto, retrolateral view; 4 Spermathecae, dorsal view; 6 Female, general appearance. Scale lines=0.1 mm (1–5), 0.5 mm (6). D. V. Logunov 35 Figs. 7–9: 7–8 Neon convolutus Denis, 1937 (_ holotype). 7 Male palp, mesal view; 8 Ditto, retrolateral view. 9 Neon pusio Simon, 1937 (_ syntype), male palp, retrolateral view. Scale lines=0.1 mm. male collected together [i.e. from the same locality considered a junior synonym for a long time. This is (Djebel Daya), the same habitat (Quercus forest; op. cit.: hardly surprising, since the females of both species 1027) and on the same date] with two females identified have virtually identical epigynes and spermathecae by him as N. rayi, but did not pay any attention to this (see Figs. 17–19 and 22–24) and differ only in body fact. Both these females probably should be assigned to coloration (cf. Figs. 20 and 25), whereas the male of S. N. convolutus as well. rivalis has remained unknown until now, apart from the The record of N. convolutus from a single \ from the description without figures in Simon (1937). The males Azores (Wunderlich, 1992) should be disregarded, judg- of both species are similar, but that of S.