DOCSLIB.ORG

Small Ornithopod Dinosaur Tracks And

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Small Ornithopod Dinosaur Tracks And LJMU Research Online Milàn, J, Falkingham, PL and Mueller-Töwe, IJ Small ornithopod dinosaur tracks and crocodilian remains from the Middle Jurassic Bagå Formation, Bornholm, Denmark: Important additions to the rare Middle Jurassic vertebrate faunas of Northern Europe http://researchonline.ljmu.ac.uk/id/eprint/14014/ Article Citation (please note it is advisable to refer to the publisher’s version if you intend to cite from this work) Milàn, J, Falkingham, PL and Mueller-Töwe, IJ (2020) Small ornithopod dinosaur tracks and crocodilian remains from the Middle Jurassic Bagå Formation, Bornholm, Denmark: Important additions to the rare Middle Jurassic vertebrate faunas of Northern Europe. Bulletin of the Geological LJMU has developed LJMU Research Online for users to access the research output of the University more effectively. Copyright © and Moral Rights for the papers on this site are retained by the individual authors and/or other copyright owners. Users may download and/or print one copy of any article(s) in LJMU Research Online to facilitate their private study or for non-commercial research. You may not engage in further distribution of the material or use it for any profit-making activities or any commercial gain. The version presented here may differ from the published version or from the version of the record. Please see the repository URL above for details on accessing the published version and note that access may require a subscription. For more information please contact [email protected] http://researchonline.ljmu.ac.uk/ BULLETIN OF THE GEOLOGICAL SOCIETY OF DENMARK · VOL. 68 · 2020 Small ornithopod dinosaur tracks and crocodilian remains from the Middle Jurassic Bagå Formation, Bornholm, Denmark: Important additions to the rare Middle Jurassic vertebrate faunas of Northern Europe JESPER MILÀN, PETER L. FALKINGHAM & INKEN JULIANE MUELLER-TÖWE Milàn, J., Falkingham, P.L. & Mueller-Töwe, I. J. 2020. Small ornithopod dinosaur tracks and crocodilian remains from the Middle Jurassic Bagå Formation, Born- holm, Denmark: Important additions to the rare Middle Jurassic vertebrate faunas of Northern Europe. Bulletin of the Geological Society of Denmark, Vol. 68, pp. 245–253. ISSN 2245-7070. https://doi.org/10.37570/bgsd-2020-68-11 Geological Society of Denmark Two new small tridactyl dinosaur tracks are found in the Middle Jurassic Bagå https://2dgf.dk Formation of Bornholm and are interpreted as ornithopodian in origin. A skeletal fragment is identified as a crocodilian skull fragment. Previous finds of dinosaur Received 15 May 2020 tracks from the locality consist of two sizes of sauropods, a medium sized theropod Accepted in revised form and thyreophorans. The addition of tracks from ornithopod dinosaurs and skeletal 26 October 2020 Published online evidence of crocodilians now give a broader picture of a diverse Middle Jurassic 17 November 2020 vertebrate fauna. This is an important addition to the understanding of the terres- trial Mesozoic ecosystem of Denmark, and a valuable addition to the scarce Middle © 2020 the authors. Re-use of material is Jurassic vertebrate record of Europe. permitted, provided this work is cited. Creative Commons License CC BY: Keywords: Dinosaur, ornithopod, crocodile, ichnology, Middle Jurassic, Bathonian, https://creativecommons.org/licenses/by/4.0/ Bajocian. Jesper Milàn [[email protected]], Geomuseum Faxe, Østsjællands Museum, Østervej 2, DK-4640 Faxe, Denmark. Peter L. Falkingham [[email protected]], School of Biological and Environmental Sciences, Liverpool John Moores University, L3 3AF, UK. Inken Juliane Mueller-Töwe [[email protected]], Geological Survey of Denmark and Greenland, Øster Voldgade 10, DK-1350 Copenhagen K, Denmark. Mesozoic terrestrial vertebrates are rare in Denmark, An extensive vertebrate fauna including crocodiles, as terrestrial outcrops are limited to a few geographi- dinosaurs, birds, turtles, amphibians, sharks, other cally restricted exposures on the west and southwest fish and a single mammal tooth has been retrieved coast of the Baltic island of Bornholm (Kear et al. 2016) by screen washing of samples from the lowermost (Fig. 1). The oldest evidence of terrestrial vertebrates Cretaceous Rabekke Formation in the coastal cliff consists of cross-sections through footprints in the east of Arnager (Lindgren et al. 2004, 2008; Rees et al. Lower Jurassic (Hettangian) Sose Bugt Member of 2005; Schwarz-Wings et al. 2009), and large dinosaur the Rønne Formation (Clemmensen et al. 2014), and tracks up to 70 cm in length associated with lungfish a diminutive theropod footprint known from the aestivation burrows are exposed in cross-section Pliensbachian Hasle Formation (Milàn & Surlyk 2015), (Surlyk et al. 2008). Inland quarries in the overly- together with a small fragment of a femur from a juve- ing Jydegaard Formation have produced remains nile sauropodomorph (Milàn & Cuny 2019). The Mid- of dinosaurs, turtles, crocodiles, fish scales, shark dle Jurassic (Bajocian – Bathonian) Bagå Formation has teeth and coprolites (Noe-Nygaard et al. 1987; Noe- hitherto yielded tracks from theropod, thyreophoran Nygaard & Surlyk 1988; Bonde & Christiansen 2003; and sauropod dinosaurs, preserved as natural casts of Christiansen & Bonde 2003; Bonde 2004, 2012; Milàn sandstone (Milàn & Bromley 2005; Milàn 2011). et al. 2012). After the initial report of dinosaur tracks Dinosaur tracks and crocodilian remains from the Bagå Formation, Bornholm · 245 from the Middle Jurassic Bagå Formation (Milàn & including ferns, cycads and conifers (Bartholin 1892; Bromley 2005), which included tracks attributed to Florin 1958; Gry 1969; Hoelstad 1985), and vertebrate large sauropods and thyreophorans, new tracks from trace fossils are known from tracks of sauropod, theropods, thyreophorans and small sauropods were theropod, and thyrophorean dinosaurs (Milan 2011; described by Milàn (2011). This paper describes new Milàn & Bromley 2005). tracks from ornithopod dinosaurs as well as the first skeletal evidence of a crocodilian skull fragment from the Bagå Formation. Material and Methods Two tracks are described here. The first specimen is a Geological setting small track preserved as track and naturally formed cast (Track 1) found in a sandstone boulder in the Bagå The Bagå Formation is located on the west coast of clay pit. (Fig. 2A). The boulder was split along a thin the Danish Island of Bornholm situated in the Baltic clay seam, revealing the impression and naturally Sea, and the type section is the Bagå Graven clay pit formed cast of a small tridactyl track. To enhance the of the now closed Hasle Klinkefabrik at the small features of the track, the infilling clay was mechani- town of Sorthat (Gravesen et al. 1982; Michelsen et al. cally removed in some parts of the track. The speci- 2003; Nielsen et al. 2010) (Fig. 1). Today quarrying has men was found by amateur geologist Christian Bach ceased, and the clay pit is water-filled and known as Hansen in the summer of 2018. The specimen was Pyritsøen (pyrite lake). The Bagå Formation comprises declared Danekræ (National treasure) (DK-1007) and thick grey clay, dark to black coaly clays with rootlets, stored in the collection of the Natural History Museum coal beds, and medium- to fine-grained, cross-bedded of Denmark (NHMD 625523). or laminated sandstone. In the upper part, poorly A second track (Track 2) is larger and was brought sorted, muddy and pebbly sandstone beds locally con- to our attention by Rene Vilsholm from the exhibition tain boulders of weathered granite, and deposition is centre NaturBornholm. The cast is preserved in coarse interpreted to have taken place in lakes and swamps, sandstone (Fig. 2B). The specimen is stored in the col- small crevasse channels, lacustrine deltas and fluvial lection of the Natural History Museum of Denmark channels (Gravesen et al. 1982; Koppelhus & Nielsen (NHMD 676864). 1994; Michelsen et al. 2003; Nielsen et al. 2010). The Bagå Both tracks were digitized via photogrammetry, Formation is palynologically dated to Middle Jurassic, using photos taken with a 16mp FujiFilm FinePix Bajocian – Bathonian (Gry 1969; Koppelhus & Nielsen XP80, and processed using AliceVision Meshroom 1994). The formation contains abundant plant fossils 2019.1.0 (https://alicevision.github.io) (Moulon et al. NORWAY 15°E N SWEDEN 56°N DENMARK Hasle GERMANY 14°E Fig. 1. Map of Bornholm with Meso- Bornholm zoic and Palaeozoic outcrops indi- Kultippen Sorthat Pyritsøen cated. The type locality of the Bagå Formation is the water-filled clayp- it, Pyritsøen, between Hasle and Rønne Rønne towns, located at 55.150816°N, 14.704770°E. Kultippen is situat- Cretaceous Arnager ed further north along the coast Jurassic just south of Hasle at 55.170088°N, Palaeozoic 14.705822°E. Bagå Graven and Kul- Precambrian 0 10 km 55°N Fault tippen are indicated by red stars. Modified from Graversen (2009). 246 · Bulletin of the Geological Society of Denmark 2012; Jancosek & Pajdla, 2011). Models of track 1, its wide, however, part of one of the outer digits is miss- natural cast, and track 2 were constructed using 21, ing and we estimate that the total width would have 18, and 20 photos respectively. Digital models are pre- been around 18 cm. The cast is 2.9–4.1 cm deep. Each sented as textured and false colour images (Fig. 3), and digit cast shows the presence of one large, rounded all photos and models are made available at h t t p s :// pad encapsulating the whole digit (Fig. 2B). The two doi.org/10.6084/m9.figshare.7994216.v1 following the complete digit casts terminate in casts of short blunt protocol set out by Falkingham et al. (2018). claws. The casts of the digits are separated from the The hip height of the track makers was estimated cast of the metatarsal-phalangeal pad by a shallower using the relations between foot length and hip height. area (Fig. 3B). The metatarsal-phalangeal pad is broad Alexander (1976) proposed that hip-height could be with a wide proximal part. The divarication angle approximated as 4 × foot length, while Thulborn (1990) between the outer digits is estimated to be about 75°.
Load more

Recommended publications
  • New Theropod, Thyreophoran, and Small Sauropod Tracks from the Middle Jurassic Bagå Formation, Bornholm, Denmark
    New theropod, thyreophoran, and small sauropod tracks from the Middle Jurassic Bagå Formation, Bornholm, Denmark JESPER MILÀN Milàn, J. 2011. New theropod, thyreophoran, and small sauropod tracks from the Middle Jurassic Bagå Formation, Bornholm, Denmark © 2011 by Bulletin of the Geological Society of Denmark, Vol. 59, pp. 51–59. ISSN 0011–6297. (www.2dgf.dk/publikationer/bulletin) https://doi.org/10.37570/bgsd-2011-59-06 Three new dinosaur tracks are described from the Middle Jurassic Bagå Formation of Bornholm, Denmark. The tracks are all preserved as natural casts on the underside of fluvial sandstone blocks originating from the old Hasle Klinkefabrik’s clay pit, now called Pyritsøen. The new tracks are from a medium-sized theropod, a thyreophoran, and a small sauropod. Together with a thyreophoran track and large sauropod tracks described in 2005, the Middle Jurassic dinosaur fauna of Bornholm now comprises theropods, two sizes of sauropods and at least one type of thyreophoran dinosaur. This is important additional data for the very scarce Middle Jurassic dinosaurian skeletal record of Europe. Received 22 November 2010 Accepted in revised form Key words: Dinosaur fauna, trace fossils, Middle Jurassic, theropod, thyreophoran, sauropod. 21 September 2011 Published online Jesper Milàn [[email protected]], GeomuseumFaxe, Østsjællands Museum, Østervej 2, DK-4640 Faxe, 30 September 2011 Denmark. Also Department of Geography and Geology, University of Copenhagen, Øster Voldgade 10, DK-1350 Copenhagen K, Denmark. Remains of Mesozoic terrestrial vertebrates are scarce Dinosaur remains are more commonly encountered in Denmark and have so far only been found in the in the southern part of Sweden, where numerous di- few Mesozoic outcrops along the west and southwest nosaur tracks and trackways of theropod dinosaurs, a coast of the Baltic island of Bornholm (Fig.
    [Show full text]
  • Dinosaur Tracks from the Early Cretaceous (Albian) of Parede (Cascais, Portugal): New Contributions for the Sauropod Palaeobiology of the Iberian Peninsula
    Journal of Iberian Geology 41 (1) 2015: 155-166 http://dx.doi.org/10.5209/rev_JIGE.2015.v41.n1.48662 www.ucm.es /info/estratig/journal.htm ISSN (print): 1698-6180. ISSN (online): 1886-7995 Dinosaur tracks from the Early Cretaceous (Albian) of Parede (Cascais, Portugal): new contributions for the sauropod palaeobiology of the Iberian Peninsula V.F. Santos1,2*, P.M. Callapez2,3, D. Castanera4, F. Barroso-Barcenilla5,6, N.P.C. Rodrigues1, C.A. Cupeto7 1Museu Nacional de História Natural e da Ciência, Rua da Escola Politécnica 58, 1250-102 Lisboa, Portugal 2Centro de Investigação da Terra e do Espaço da UC – CITEUC, Avenida Dr. Dias da Silva, 3000-134 Coimbra, Portugal 3Departamento de Ciências da Terra da Universidade de Coimbra, Largo Marquês de Pombal, 3001-401 Coimbra, Portugal 4Grupo Aragosaurus-IUCA, Paleontología, Facultad de Ciencias, Universidad de Zaragoza, Calle Pedro Cerbuna 12, 50009 Zaragoza, Spain 5Departamento de Paleontología, Facultad de Ciencias Geológicas, Universidad Complutense de Madrid, 28040 Madrid, Spain 6Grupo de Investigación IberCreta, Universidad de Alcalá de Henares, 28871 Alcalá de Henares, Spain 7Departamento de Geociências da Universidade de Évora, Rua Romão Ramalho 59, 7000-671 Évora, Portugal e-mail addresses: [email protected] (V.F.S., *corresponding author); [email protected] (P.M.C.); [email protected] (D.C.); [email protected] (F.B.B.); [email protected] (N.P.C.R.); [email protected] (C.A.C.) Received: 17 January 2014 / Accepted: 18 December 2014 / Available online: 25 March 2015 Abstract A recently discovered Early Cretaceous (early late Albian) dinosaur tracksite at Parede beach (Cascais, Portugal) reveals evidence of dinoturbation and at least two sauropod trackways.
    [Show full text]
  • Research Paper Inventory and Management of Dinosaurs Traces
    International Journal of Global Science Research ISSN: 2348-8344 (Online) Vol. 5, Issue. 2, October 2018, pp. 793-814 DOI: 10.26540/ijgsr.v5.i2.2018.106 Available Online at www.ijgsr.com © Copyright 2014 | ijgsr.com | All Rights Reserved Research Paper Inventory and Management of Dinosaurs Traces: A case Study of Azilal Region, Morocco Fayçal AMINE, M’Hamed BERRAHMA and Abdellatif AARAB Mohamed V University, ENS, Department of Earth Sciences, Geosciences, Natural Resources and Development Laboratory, Avenue Mohamed Bel Hassan El Ouazzani, P.O. Box 5118 Takaddoum, Rabat, Morocco. *Corresponding author E-mail: [email protected] Received: 30/09/2018 Revised: 13/10/2018 Accepted: 18/10/2018 Abstract: The preservation of geosites in under considerations has been the scope of the heritage framework is necessary at a numerous geological studies since the 20th time when urbanization, the exploitation of Century and particularly in the past 30 years. fossils and minerals are real threats to this Recent advances have revealed that irreversible heritage. This work is a geotourism is an emerging and still contribution to the preservation and underdeveloped face of tourism, mainly in development of dinosaur trace deposits by Morocco, and mapping the geoheritage of geological and geomorphological Azilal region remains lacunars. The description of the study area, then by importance of study of geoheritage of the mapping using Geographic Information Azilal Region is related the dinosaurs traces, Systems (GIS) and finally by the proposal of worked by many paleontologists and the models of development and preservation geologists (Dresnay, 1971, 1975; Couvreur, of these geosites. 1977; Dutuit & Ouazzou, 1980; Jenny et al., Keywords: Azilal region, Morocco, 1981a,b; Laville, 1981; Monbaron, 1978, Geoheritage, Geo-Routes, GIS, Geotouristic 1980, 1981, 1982a, b, 1983; Biron, 1982; Interess Local, Dinosaurs.
    [Show full text]
  • Tennant Et Al AAM.Pdf
    Zoological Journal of the Linnean Society Evolutionary relations hips and systematics of Atoposauridae (Crocodylomorpha: Neosuchia): implications for the rise of Eusuchia Journal:For Zoological Review Journal of the Linnean Only Society Manuscript ID ZOJ-08-2015-2274.R1 Manuscript Type: Original Article Bayesian, Crocodiles, Crocodyliformes < Taxa, Implied Weighting, Laurasia Keywords: < Palaeontology, Mesozoic < Palaeontology, phylogeny < Phylogenetics Note: The following files were submitted by the author for peer review, but cannot be converted to PDF. You must view these files (e.g. movies) online. S1 Atoposaurid character matrix.nex Page 1 of 167 Zoological Journal of the Linnean Society 1 2 3 1 Abstract 4 5 2 Atoposaurids are a group of small-bodied, extinct crocodyliforms, regarded as an important 6 3 component of Jurassic and Cretaceous Laurasian semi-aquatic ecosystems. Despite the group being 7 8 4 known for over 150 years, the taxonomic composition of Atoposauridae and its position within 9 5 Crocodyliformes are unresolved. Uncertainty revolves around their placement within Neosuchia, in 10 11 6 which they have been found to occupy a range of positions from the most basal neosuchian clade to 12 13 7 more crownward eusuchians. This problem stems from a lack of adequate taxonomic treatment of 14 8 specimens assigned to Atoposauridae, and key taxa such as Theriosuchus have become taxonomic 15 16 9 ‘waste baskets’. Here, we incorporate all putative atoposaurid species into a new phylogenetic data 17 10 matrix comprising 24 taxa scored for 329 characters. Many of our characters are heavily revised or 18 For Review Only 19 11 novel to this study, and several ingroup taxa have never previously been included in a phylogenetic 20 21 12 analysis.
    [Show full text]
  • A New Narrow-Gauge Sauropod Trackway from the Cenomanian Candeleros Formation, Northern Patagonia, Argentina
    Accepted Manuscript A new narrow-gauge sauropod trackway from the Cenomanian Candeleros Formation, northern Patagonia, Argentina Arturo Miguel Heredia, Pablo José Pazos, Diana Elizabeth Fernández, Ignacio Díaz Martínez, Marcos Comerio PII: S0195-6671(18)30249-0 DOI: https://doi.org/10.1016/j.cretres.2018.11.016 Reference: YCRES 4019 To appear in: Cretaceous Research Received Date: 14 June 2018 Revised Date: 8 October 2018 Accepted Date: 20 November 2018 Please cite this article as: Heredia, A.M., Pazos, P.J., Fernández, D.E., Martínez, I.D., Comerio, M., A new narrow-gauge sauropod trackway from the Cenomanian Candeleros Formation, northern Patagonia, Argentina, Cretaceous Research, https://doi.org/10.1016/j.cretres.2018.11.016. This is a PDF file of an unedited manuscript that has been accepted for publication. As a service to our customers we are providing this early version of the manuscript. The manuscript will undergo copyediting, typesetting, and review of the resulting proof before it is published in its final form. Please note that during the production process errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain. ACCEPTED MANUSCRIPT MANUSCRIPT ACCEPTED A new narrow-gauge sauropod trackway from the Cenomanian Candeleros Formation, northern Patagonia, Argentina Arturo Miguel Heredia1, 2, Pablo José Pazos1, 2, Diana Elizabeth Fernández1, 2, Ignacio Díaz Martínez3, Marcos Comerio4 1- Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales. Departamento de Ciencias Geológicas. Buenos Aires, Argentina. 2- CONICET - Universidad de Buenos Aires. Instituto de Estudios Andinos Don Pablo Groeber (IDEAN). Buenos Aires, Argentina.
    [Show full text]
  • New Sauropod Trackways from the Middle Jurassic of Portugal
    New sauropod trackways from the Middle Jurassic of Portugal Vanda F. Santos, José J. Moratalla, and Rafael Royo-Torres Acta Palaeontologica Polonica 54 (3), 2009: 409-422 doi: http://dx.doi.org/10.4202/app.2008.0049 The Galinha tracksite reveals a sequence of Bajocian–Bathonian limestones belonging to the Serra de Aire Formation (West−Central Portugal) and is one of the few sites in the world where Middle Jurassic sauropod dinosaur tracks can be found. This tracksite is characterised by the presence of long, wide gauge sauropod trackways, the Middle Jurassic age of which suggests these dinosaurs were more widely distributed over time than previously thought. Two trackways contain unique pes and manus prints with morphologies that allow a new sauropod ichnotaxon to be described: Polyonyx gomesi igen. et isp. nov. On the basis of different manus/pes prints and trackway features, the proposal is made to subdivide Sauropodomorpha ichno−morphotypes into five groups: Tetrasauropus−like, Otozoum −like, Breviparopus/Parabrontopodus−like; Brontopodus−like, and Polyonyx−like. Polyonyx gomesi igen. et isp. nov. is thought to represent a nonneosauropod eusauropod, with a well developed manus digit I. The posterior orientation of this digit print suggests they were made by a eusauropod dinosaur with a posteriorly rotated pollex. The manus print morphologies observed in two trackways suggest a stage of manus structure intermediate between the primitive non−tubular sauropod manus and the tubular metacarpal arrangement characteristic of more derived sauropods. The low heteropody (manus:pes area ratio 1:2) of the trackway renders it possible they could have been made by eusauropods such as Turiasaurus riodevensis, which has a similar manus:pes area ratio.
    [Show full text]
  • Microvertebrates of the Lourinhã Formation (Late Jurassic, Portugal)
    Alexandre Renaud Daniel Guillaume Licenciatura em Biologia celular Mestrado em Sistemática, Evolução, e Paleobiodiversidade Microvertebrates of the Lourinhã Formation (Late Jurassic, Portugal) Dissertação para obtenção do Grau de Mestre em Paleontologia Orientador: Miguel Moreno-Azanza, Faculdade de Ciências e Tecnologia da Universidade Nova de Lisboa Co-orientador: Octávio Mateus, Faculdade de Ciências e Tecnologia da Universidade Nova de Lisboa Júri: Presidente: Prof. Doutor Paulo Alexandre Rodrigues Roque Legoinha (FCT-UNL) Arguente: Doutor Hughes-Alexandres Blain (IPHES) Vogal: Doutor Miguel Moreno-Azanza (FCT-UNL) Júri: Dezembro 2018 MICROVERTEBRATES OF THE LOURINHÃ FORMATION (LATE JURASSIC, PORTUGAL) © Alexandre Renaud Daniel Guillaume, FCT/UNL e UNL A Faculdade de Ciências e Tecnologia e a Universidade Nova de Lisboa tem o direito, perpétuo e sem limites geográficos, de arquivar e publicar esta dissertação através de exemplares impressos reproduzidos em papel ou de forma digital, ou por qualquer outro meio conhecido ou que venha a ser inventado, e de a divulgar através de repositórios científicos e de admitir a sua cópia e distribuição com objetivos educacionais ou de investigação, não comerciais, desde que seja dado crédito ao autor e editor. ACKNOWLEDGMENTS First of all, I would like to dedicate this thesis to my late grandfather “Papi Joël”, who wanted to tie me to a tree when I first start my journey to paleontology six years ago, in Paris. And yet, he never failed to support me at any cost, even if he did not always understand what I was doing and why I was doing it. He is always in my mind. Merci papi ! This master thesis has been one-year long project during which one there were highs and lows.
    [Show full text]
  • Titanosaur Trackways from the Late Cretaceous El Molino Formation of Bolivia (Cal Orck’O, Sucre)
    Annales Societatis Geologorum Poloniae (2018), vol. 88: 223 – 241. doi: https://doi.org/10.14241/asgp.2018.014 TITANOSAUR TRACKWAYS FROM THE LATE CRETACEOUS EL MOLINO FORMATION OF BOLIVIA (CAL ORCK’O, SUCRE) Christian A. MEYER1, Daniel MARTY2 & Matteo BELVEDERE3 1 Department of Environmental Sciences, University of Basel, Bernoullistrasse 32, CH-4056 Basel, Switzerland; e-mail: [email protected] 2 Museum of Natural History Basel, Augustinergasse 2, CH- 4000 Basel, Switzerland; e-mail: [email protected] 3 Office de la culture, Paléontologie A16, Hôtel des Halles, P.O. Box 64, CH-2900 Porrentruy 2, Switzerland; e-mail: [email protected] Meyer, C. A., Marty, D. & Belvedere, M., 2018. Titanosaur trackways from the Late Cretaceous El Mo- lino Formation of Bolivia (Cal Orck’o, Sucre). Annales Societatis Geologorum Poloniae, 88: 223 – 241. Abstract: The Cal Orck’o tracksite is exposed in a quarry wall, approximately 4.4 km NW of Sucre (Department Chuquisaca, Bolivia) in the Altiplano/Cordillera Oriental, in the El Molino Formation (Middle Maastrichtian). Fossiliferous oolitic limestones, associated with large, freshwater stromatolites and nine levels of dinosaur tracks in the El Molino Formation document an open lacustrine environment. The main track-bearing level is almost vertical with a surface area of ~ 65,000 m2. The high-resolution mapping of the site from 1998 to 2015 revealed a total of 12,092 individual dinosaur tracks in 465 trackways. Nine different morphotypes of dinosaur tracks have been documented. Amongst them are several trackways of theropods, orni- thopods, ankylosaurs and sauropods, with the latter group accounting for 26% of the trackways.
    [Show full text]
  • Weathering Profiles and Clay Mineralogical Developments, Bornholm, Denmark
    Weathering profiles and clay mineralogical developments, Bornholm, Denmark Pingchuan Tan1, 2, Nikolas Oberhardt1, Henning Dypvik1,2, Lars Riber1, Ray E. Ferrell Jr3. 1 Department of Geoscience, University of Oslo, P.O.BOX 1047, Blindern, NO-0316 Oslo, Norway 2 Centre for Earth Evolution and Dynamics, University of Oslo, P.O.BOX 1028, Blindern, NO-0315 Oslo, Norway 3 Department of Geology & Geophysics, Louisiana State University, Baton Rouge, LA, 70803, USA Corresponding author – Pingchuan Tan: [email protected] ABSTRACT Saprock-saprolite associations were studied by field and laboratory methods (optical microscopy, X-ray powder diffraction, scanning electron microscopy, electron microprobe) in order to describe regolith development in the crystalline rocks of the Nygård kaolin pit (Bornholm, Denmark). The clay sequences and stages of porosity development are similar to those observed for reservoir rocks from the Utsira High (Riber, L., Dypvik, H., Sørlie, R. and Ferrell, R. (2016) Clay minerals in deeply buried paleoregolith profiles, Norwegian North Sea. Clays and Clay Minerals, in press). The weathering of the parent granite began before the end of the Mesozoic. Two stages of syn- /pre-burial alteration, followed by diagenesis during burial, and then post-uplift weathering have been recognized. In stage Ι, plagioclase and some biotite (biotite-vermiculite-kaolinite) reacted to form elongate booklets of highly-ordered kaolinite or smaller, blocky pseudohexagonal crystals. Stage II represented more extreme weathering developed along local fracture systems. The higher potential for fluid flow in the fractures caused highly-ordered kaolinite to alter to halloysitic, poorly- ordered kaolinite. Plagioclase, biotite, and K-feldspar continued to interact with formation water and formed additional quantities of secondary clay minerals.
    [Show full text]
  • Sauropod and Small Theropod Tracks from the Lower Jurassic Ziliujing Formation of Zigong City, Sichuan, China, with an Overview
    Ichnos, 21:119–130, 2014 Copyright Ó Taylor & Francis Group, LLC ISSN: 1042-0940 print / 1563-5236 online DOI: 10.1080/10420940.2014.909352 Sauropod and Small Theropod Tracks from the Lower Jurassic Ziliujing Formation of Zigong City, Sichuan, China, with an Overview of Triassic–Jurassic Dinosaur Fossils and Footprints of the Sichuan Basin Lida Xing,1 Guangzhao Peng,2 Yong Ye,2 Martin G. Lockley,3 Hendrik Klein,4 W. Scott Persons IV,5 Jianping Zhang,1 Chunkang Shu,2 and Baoqiao Hao2 1School of the Earth Sciences and Resources, China University of Geosciences, Beijing, China 2Zigong Dinosaur Museum, Zigong, Sichuan, China 3Dinosaur Trackers Research Group, University of Colorado, Denver, Colorado, USA 4Saurierwelt Palaontologisches€ Museum, Neumarkt, Germany 5Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada Keywords Grallator, Parabrontopodus, Hejie tracksite, Ma’anshan A dinosaur footprint assemblage from the Lower Jurassic Member, Zigong Ziliujing Formation of Zigong City, Sichuan, China, comprises about 300 tracks of small tridactyl theropods and large sauropods preserved as concave epireliefs (natural molds). The INTRODUCTION theropod footprints show similarities with both the ichnogenera Zigong is famous for the Middle Jurassic Shunosaurus Grallator and Jialingpus. Three different morphotypes are fauna and the Late Jurassic Mamenchisaurus fauna. However, present, probably related to different substrate conditions and extramorphological variation. A peculiar preservational feature fossils from the Early Jurassic and the Late Triassic are com- in a morphotype that reflects a gracile trackmaker with paratively rare. Previously reported Early Jurassic fossils extremely slender digits, is the presence of a convex epirelief that include fragmentary prosauropod and sauropod skeletal occurs at the bottom of the concave digit impressions.
    [Show full text]
  • Dinosaur Tracks in Lower Jurassic Coastal Plain Sediments (Sose Bugt Member, Rønne Formation) on Bornholm, Denmark
    Dinosaur tracks in Lower Jurassic coastal plain sediments (Sose Bugt Member, Rønne Formation) on Bornholm, Denmark LARS B. CLEMMENSEN, JESPER MILAN, GUNVER K. PEDERSEN, ANNE B. JOHANNESEN AND CONNIE LARSEN Clemmensen, L.B., Milan, J., Pedersen, G.K., Johannesen, A.B. & Larsen, C. 2014: Dinosaur tracks in Lower Jurassic coastal plain sediments (Sose Bugt Member, Rønne Formation) on Bornholm, Denmark. Lethaia,Vol. 47, pp. 485–493. Fluvial palaeochannels of coastal plain sediments of the Lower Jurassic Sose Bugt Member of the Rønne Formation exposed in the coastal cliffs at Sose Bugt, Bornholm, contain abundant dinosaur or other large vertebrate tracks in the form of deformation structures exposed in vertical section. The tracks are represented by steep-walled, flat- to-concave-bottomed depressions, with a raised ridge at each side. The tracks are filled with laminated sediments, draping the contours of the bottom of the depression. Un- derprints, stacked concave deformations beneath the prints, are present beneath each track. Contemporary Upper Triassic – Lower Jurassic strata from southern Sweden and Poland contain a diverse track fauna, supporting our interpretation. This is the earliest evidence of dinosaur activity in Denmark. □ Bornholm, coastal plain, dinosaur tracks, lake sediments, Lower Jurassic. Lars B. Clemmensen [[email protected]], Jesper Milan [[email protected]], Anne B. Joh- annesen [[email protected]], and Connie Larsen [[email protected]], Depart- ment for Geosciences and Natural Resource Managements, University of Copenhagen, Øster Voldgade 10, DK-1350 Copenhagen K, Denmark; Jesper Milan [[email protected]], Geomuseum Faxe/Østsjællands Museum, Østervej 2, DK-4640 Faxe, Denmark; Gunver K. Pedersen [[email protected]], GEUS Geological Survey of Denmark and Greenland, Øster Voldgade 10, DK-1350 Copenhagen K, Denmark; manuscript received on 12/04/2013; manuscript accepted on 04/12/2013.
    [Show full text]
  • Oj Va Le.Njoa.Ndl,Si Casa N Ods 5Ne4r
    New Sauropod Trackways from the Middle Jurassic of Portugal Author(s): Vanda F. Santos, José J. Moratalla, and Rafael Royo-Torres Source: Acta Palaeontologica Polonica, 54(3):409-422. 2009. Published By: Institute of Paleobiology, Polish Academy of Sciences DOI: http://dx.doi.org/10.4202/app.2008.0049 URL: http://www.bioone.org/doi/full/10.4202/app.2008.0049 BioOne (www.bioone.org) is a nonprofit, online aggregation of core research in the biological, ecological, and environmental sciences. BioOne provides a sustainable online platform for over 170 journals and books published by nonprofit societies, associations, museums, institutions, and presses. Your use of this PDF, the BioOne Web site, and all posted and associated content indicates your acceptance of BioOne#s Terms of Use, available at www.bioone.org/page/terms_of_use. Usage of BioOne content is strictly limited to personal, educational, and non-commercial use. Commercial inquiries or rights and permissions requests should be directed to the individual publisher as copyright holder. BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research. New sauropod trackways from the Middle Jurassic of Portugal VANDA F. SANTOS, JOSÉ J. MORATALLA, and RAFAEL ROYO$TORRES ATS(Wachetnaets oGPts$aCa,ll Viaennhe.Fota.ra,nlt MtroPa lcooorkgratsuiitctagaella Pa lr,e)oJ va le.nJoa.ndl,si icasa an ods 5ne4R eqo u( o3yefo)n :t$ch T4eeo0 r o9ffre%we B4s,2as R 2jiot.. escD2 ia0iO nn0I9% t: hB.1eN a0tw e.h4wo2onr0s lid2aa/un awrpl hopiepm.r2oede0s M0t t8roia.nd0cekd0sl4web 9a Je.yulsoranf rsgsoimincgs t aht ouert oM hpeiodS ddeld erriJ naoud sraeasuA sriict rreo afcF P kosormrc tauangtiab olen.
    [Show full text]