The malacologicalsocietymalacological society of Japan

Hre VEts'US(Jap. Jour. Matuc,) -4] VoL 51, Nos.1&2(1992)/11

H 7ts v > 7tf" ･ Se ?fiitiii]FkltwEl Y' i [rtEN(iva,Ell an 7 Y"v7 lf"i N) CaDv)(

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The Oenopotinae (: Turridae) Mollusks from the Southeastern Part of the Japan Sea

Igor P. BoGDANov and Katsuchiyo ITo

(Dept. of lnvertebrate Zoology, Biological Faculty, Moscow State University, 119899 Russia

and ex-Japan Sea Regional Fisheries Research Laboratory, Suido-cho, Niigata 951, Japan)

Abstract: Twenty-seven species of the Subfamily Oenopotinae of the Family Turridae were identified in the materiats that had been previously reported by Ito (1967, 1985, 1989) and Ito et al. (1986) from Japan Sea. Among them 21 species among which three are new to science are described and illustrated. The most of them are widely distributed subarctic

species, but ranges of a few species seem to be limited in Japan Sea coast of Honshu,

This study is an attempt to summarize the material, which was collected recently. The

material consists of one of the most interesting groups of the family Turridae, the subfami-

ly Oenopotinae, which is mainly distributed in the North Polar Basin. All the reviewed

material was collected by the junior author (Katsuchiyo Ito) in different areas of the Japan

Sea near the coast of Honshu, the region of Ishikawa and Niigata Prefeetures at depths

from 200 to 350 metres. The photos of the part of the Oenopotinae shells were early

pub]ished by Ito without descriptions or diagnoses and sometimes without the specific determination (Ito, 1967, 1985, 1989, Ito et al., 1986), The geographical ceordinates of

Tab]e 1. Collecting stations of specimens under report (I-series)

Station Date Coordinate Depth (m)

I-1I-2I-3I-4l-51-614.06,1980 37eOl,O N-136022.0 E 2S531431820S25025S

26.06.1980 37012,7 N-.137e12.1 E

25.06.1980 37014,6 N-137e12.9 E

13.06.1980 37D05.5 N-136025.0 E

13.06.1980 37004.4 N-136e22.1 E

14.06.1980 36057.0 N-136e21,9 E

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12 VENUS: Vol. 51, Nos.1&2(1992)

the I-series stations are given in Table 1. The locality accompanying S designated off

Sado Island where sampling was done during August 1981 till July 1982. All stations

are fallen within the range within: 38"08.1-08.6N, 138e44.0-44.6eE, 200-300 m deepl

38002.5-03.2N, 138e38.3-38.7E, 200-275 m deep, Summarizing up the material and literature data, it became clear that Oenopotinae fauna of the southeastern part of the Japan Sea is represented by five genera and 27 species.

Genus Curtitoma Bartsch, 1941

1. Curtitoma violacea (Mighe)s et Adams, 1842)

(Figs. Ia-e, 22)

Pteurotoma bicarinata Couthouy, 1838 (non W. Wood, 1828): 104, pl. 1, fig. 11. Pteurotoma violacea Mighels et Adarns, 1842: 51, pl. 4, fig. 21.

Dqf7ancia livida MO]ler, 1842: 14.

Bela violacea var. cylindracea MOrch, 1868: 215,

Beta bicarinata var. ruLfescens G.O. Sars, 1878: 238, tab. 16, fig. 11.

Bela bicarinata var. ge,ninotineata Friele, 1879: 277; 1886: 16, pl. 8, fig. 19.

Bela minuscularia LocaTd, 1897: 2S9, pr. 12, fig, 28-30.

Nbmatoma sp., Ito, 1985: 29, pl, 7, fig. 4. Propebela sp. No. 9, Ito, 1985: 30, pl. 8, fig. 3.

IVbmatoma sp, No. 3, Ito, 1986: 17, pl. 21, fig. 4.

Benthomangelia ? sp. Ito, 1989: 52, pl. 12, fig. 7.

The shell is fusiform, small, from 6 to 8 mm in height, consists of 4.5-5 whorls.

The spiral is represented sometimes by two well-defined ribs. But, as a rule,

there are two more spiral ribs. In some cases, spiral elements are deve!oped on the pro-

toconch, but absent on the definitiye shell whorls. It is worth to mention that this species

is characterized by significant variety of sculptures, The axial sculpture is developed to

some extent. It is represented by the folds evenly developed on the surface, straight

on the periphery of the whorl and sigmoidally curved on the shoulder. has a large number of microscopic granules, concentrated on the spiral ribs.

Remarks: The species is well separate from other Curtitoma species by fusiform shell and two or more well-defined spiral ribs. A great variability of spiral and axial sculptures and the shell polymorphism caused the previous investigators to create a lot of new species.

But, all of them are the infraspecific variety of Curtitoma violaeea because of the identical

morphology of the radular tooth.

Distribution: The boreal-arctic, widely spread species. It is distributed in all arctic

seas, and is found in the boreal regions of the Atlantic and Pacific Oceans. It was recorded

from the Canadian Arctic : Labrador Strait, British Columbia, Newfoundland, New England,

Massachusetts, western and eastern parts of Greenland, west Ireland and Iceland, near

the coast of Norway, Spitzbergen, in Barentz and White Seas, near the islands of Franz-

Josef Land, Novaja Zemlja, Severnaja Zemlja, in the Laptev Sea near Novosiberian Islands

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Bogdanov-Ito: Oenopotinae from Japan Sea 13

and in the Chukotski Sea near Wrangel Island, It is also found in the Bering Sea, off

the coast of eastern Kamchatka and in the Okhotsk Sea. In the Japan Sea it was recorded

in the Tartary Strait and near the Primorie coast. It was found near the Japanese coast

at Stations S-67, 59 and S-52.

2. Curtitoma decussata (Couthouy, 1839)

(Figs. 2a-b)

Pleurotoma decussata Ceuthouy, 1839: 183, pl. 4, fig. 8. Pleurotoma tenuicostata M. Sars, 1868: 259 (nomen nudum). Bela tenuicostata var. wiltei Friele, 1886: 13, pl. 8, fig. 16, 17, pl. 10, fig. 14 (radula).

Bela willei Kobelt, 1905: 254, pl, 84, fig, 6.

IVlematoma dissoluta Ito, 1989: 52, pl. 12, fig. 2-3.

The shell is oval or oval-fusiform, small, from 8 to 9 mm in height, consists of 4-5

whorls. (Dimensions of the holotype: 8.7 mm in height of the shell (H) and 3,7 mm

in body whorl diameter (D). The maximum sizes of the shell from Massachusetts: H= 12,5,

D=6,O mm). The spiral sculpture is represented by thread-like ribs, which are, usually

2-3 times wider than spaces between them. The ribs are equal in size on the whole

surface of the whorl and the base of the shell. The axial sculpture is represented by numer-

ous thin folds (from 14 to 34 in number) divided by spaces equal in width to the folds, The most developed folds are on the shoulder and on the upper part of the periphery.

They are obsolete on the base. The characteristic feature of the species is the flattened

shell base, transforming into the siphonal canal.

Remarks: This species is close to C. Iawrenciana by radular morphology and the shell

form. but can be distinguished therefrom by numerous thin axial folds.

Distribution : Boreal and arctic parts of the North Pacific, spread in the East Siberian,

Kara, Laptev and Chukehi Seas, in the Labrador, near the New Scotland coast, in the

Massachusetts Bay and in Canadian Arctic, in the Bering, Okhotsk and Japan Seas. It

was found near the Japanese coast at Stn. [-5.

3. Curtitoma becklemishevi Bogdanev, 1989

(Figs. 3a-g, 23)

iVematoma hokkaidoensis Okutani, 1964 (non Bartsch, 1941): 431, pl, 7, fig. 9; 1966: 24,

pl, 2, fig. 8, 9; Ito, 1985: 29, pl. 7, fig. 5: lto, 1989: 52, pl, 12, fig, 1. Curtitoma becklemishevi Bogdanov, 1989: 134, fig. 1, 1-2; 2, a-b. Propebela sp. No. 3, rto et al., 1986: 18, pl, 23, fig, 1.

The shell is oval-fusiform, light-gray, with thin walls. (Dimensions of the holotype:

H=7,7, D=3.9 rnm.) The spiral sculpture is represented by string-like ribs, which are

equally spaced and are of the same size on the whole surface of the shell. The axial sculp-

ture is represented by thin folds, which have a deep sigmoidal bend on the shoulder.

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14 VENUS: Vol. 51, Nos,1&2(1992)

There are from 14 to 18 folds on the body whorl and from 13 to 16 on the penultimate

one.

Remarks: This species is the closest to C, hebes but may be separated by its elongate

siphonal canal and more round shell base.

Distribution: The Asian boreal parts of the North Pacific. Okhotsk (51010'N-

154017'E), in the Tartary Strait and near the Primorie coast. It was found near the Japanese

coast at Stn. I-1, I-2, I-3, S-48, S-S8 and S-60.

4. Curtitoma trevelliana (Tllrton, 1834)

(Figs. 5a-c)

Pleurotoma reticulata Brown, 1827: pl. 48, fig. 29, 30 (only figures);1844: 8, pl.5, fig. 29-30.

Pteurotoma tneveUianum Turton, 1834: 351.

1886: 12, 8, 11-13. Beta decussata var. .finmarchia Friele, pl, fig.

The shell is fusiferm or turretiform, small, from 4.5 to 8.0 mm in height. The Iargest

±= specimen from Murmansk is H= 10.6, D=5.2, hb.;7.7 (height of body whorl), h, 5.4 (height of ); d=2.3 mm (width of aperture). The spiral sculpture is represented by fine striae on the shoulder and string-like ribs on the other surface of the whorl and

on the shell base. As a rule, the most developed rib lies on the shoulder and in some

specimens it becomes the keel. The axial elements are different in size and number from

16 to 24. The most developed folds are on the shoulder and on the upper part of the

periphery. They become obsolete on the base of the shell. The characteristic feature of the species is the callous growths, which are formed at the crossing of axial and spiral

sculptural elements.

Remarks: This species is close to C. violaeea and C. .fiora by the shell form, but

can be identified by dagger-shaped radula tooth. The usage of Pleurotoma trevelliana was

recommended by Bouchet and Waren (1980) as well-known and frequent in literature. The "reticutata" name PL reticulata is nomen oblitum, from 1827 to 1904 the name was used

only three times in 78 scientific works (Dautzenberg and Fischer, 1912), Distribution: The boreal-arctic species, Recorded in the Canadian Arctic and off the

arctic shore ef Alaska, near western and eastern Greenland, in Davis strait, Labrador,

near New England, Newfoundland and Nova Scotia, and southward up to Cape Cod in

Atlantic. In the eastern Atlantic it was found near Iceland, Scotland, in the Gulf of

Gascogne, near Norway and Great Britain, also in the Barentz, White, Kara and Laptev

Seas, common near the coast of Franz Josef Land, Sewernaja Semlja and Novosiberian

Islands. It was found near the Japanese coast at the Stn. I-3.

5. CZtrtitoma niigataensis n. sp.

(Fig. 4)

The shell is oval-fusiform, white, small: H=8,O; D=4.2; h.b=5.7; h,== 1.8 mm. The

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Begdanov.lto: Oenopotinae from Japan Sea 15

spiral sculpture is represented by the numerous fine string-like ribs, which are, as a rule, 3-4 times wider than the spaces between them. The ribs are equal in size on the whole surface of the shell. The spiral sculpture of the shoulder consists of 7-8 thread-like

ribs with spaces between them being 2-3 times wider. The axial sculpture is represented

by prominent but thin sigmoidally curved folds. The characteristic feature of the species is these sigmoidal folds, which occupy 314 of the whorl peripheral upper part. The folds

are divided by spaces equal in width to the folds.

Remarks: This species is similar to Curtitoma decussata, though it differs by a very prominent sheli base and a larger sigmoidal bend.

Distribution: Known only from Stn. S--49.

Genus Granotoma Bartsch, 1941

6. Granotoma albrechti (Krause, 1885) (Figs. 6a-d, 24a-b)

Bela albrechti Krause, 1885: 276, pl. 18, fig. 3, 11; Dalr, 1886: 300, pl 4, fig. 1. ?Vbmatoma dissoluta, lto, 1985: 30, pl. 7, fig, 6. Nbmatoma sp. No. 2, Ito et al., 1986: 17, pl, 21, fig. 3.

The shell is oval-fusiform, white, sometimes with the greenish hue. An average size ofthe shell with 4-5 whorls is from 7 to 8 mm. (The holotype: H=10.0, D=5.0 mm. The largest specimen from the Okhotsk Sea: H=10.2, D=5.5, hb.=7.2, h.=5･8;

d=2.3 rnm.) The spiral sculpture consists of numerous fine ribs, which are equal in size

on the whole surface of the whorl and, as a rule, 4-5 times wider than the spaces be-

tween them. The axial sculpture is represented by medium-sized and sharply angulated fold$, which are equally deyeloped on the shoulder, on the periphery and on the base of the shell, There is a small sigmoidal band on the upper part of the shoulder with intervals3-7 times narrower than the folds. There are 20 to 38 folds on the body

whorl, and 17 to 34 on the penultimate one. The equal sizes of the axial and spiral sculp- tures produce reticulate pattern on the shell surface, The first whorl of protoconch has no macrosculptural elements, but is evenly covered by minute granules (Fig. 24b). The spiral sculpture develops on the second embryonic whorl. All microscuipture granules are

concentrated on the thread-like spiral ribs and on the axial folds.

Remarks: This species is the closest to G. krausei but may be separated by triangular axial folds in section and cylindrical form of the radular tooth base. Distribution:The Pacific boreal species. Distributed mainly along the Asian coast, in the Chukotski Sea near the Wrangel lsland, in the Bering Sea from east Kamchatka to Bering Strait, in the northern and eastern parts of Primorie, near Japanese and Okhotsk coasts of Sakhalin, in the northern part of the Okhotsk Sea and near the middle Kurile Islands. It was found in the Japan Sea coast at Stn. S-52.

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7. 6ranotoma raduga (Bogdanoy, 1985)

(Fig. 7)

Oenopota raduga Bogdanov, 198S: 451, fig, 1, D-3, 2, 2.

The shelt is oval-fusiform, cream colored, with thin walls. An average size of the shell with 3-4 whorls is from 8 to 9 mm. (The holotype: H=10.3, D=5.9, hbw==7.6, h.=5.8, d=2.7 mm). The spiral sculpture consists of numerous, equal, flat, fine ribs on the periphery and base of the shell. Ribs are, as a rule, 2-2.5 times wider than the spaces between them. The spiral elements are feebly expressed on the shoulder, The axial sculpture is represented by practically straight, but oblique folds with sigmoidal bend on upper part of periphery. The axial folds are equal on the whole surface and divided by spaces equal in width. There are 16 to 22 folds on the body whorl, 14 to 18 on the penultimate one. Remarks: G. raduga is distinguished from others by its protoconch with microsculp- ture on the spirally sculptured surface, and thin spiral ribs of the definitive shell with

wide axial folds,

Distribution: The Pacific boreal species. Distributed mainly near the Asian coast.

Lives near eastern coast of Sakhalin in the Okhotsk Sea, and near eastern part of Kamchatka

in the Bering Sea. It was found near the Japanese coast at Stn. S-58.

Genus Oenopota MOrch, 1852

8. 0enopota cancellata (Mighels et Adams, 1840)

(Figs. 8a-h, 25)

Fletsus canceUatus Mighels et Adams, 1840: 50; 1842: 52, pl, 4, fig. 18. Oenopota dectivis Macpherson, 1971 (non Loven, 1846): 115, pl. 7, fig. 10, Propebela sp. No. S, No. 6, No. 8, Ito, 1985: 30, pl. 7, fig, 11, 12. Propebela sp. No. 1, Ito et al,, 1986: 18, pl. 22, fig. 5. Obesotoma hanazakiensis, lto et al., 1986: 19, pl. 24, fig. 4.

The shell is fusiform, white or sometimes yellowish brown, large in size. An average

size of the shell with 7-8 whorls is from 11 to 15 mm. (The largest dimensions were reported for the holotype: H=20.0, D=7.75 mm by Verrill (1882)). The spiral sculpture of adult shell consists of numerous, hair-like, fine ribs, which are equal on the whole surface of the whorl and on the upper part of shell base. The axial sculpture is represented by very prominent folds, They are equal on the whorl surface and divided by spaces equal in width. The elements of axial sculpture are practically straight, but show a considerable inclination on the back. These folds have a srnall sigmoidal bend only on the upper part of periphery. There ae 18 to 22 folds on the body whorl, 16 to 19 on the penultimate one.

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Bogdanov.Ito: Oenopotinae from Japan Sea 17

Remarks: This species is like to O. biconiea, but may be separated by more prominent

whorl form and less developed spiral sculpture.

Distribution: The amphiboreal, widely distributed species. Recorded in the boreal

waters of Pacific Ocean and in high-boreal western Atlantic. It was found near the New

England (type locality), in the Casco Bay, Massachusetts Bay, Labrador and near the coast

of Nova Scotia. In the Japan Sea it was found in the Gulf of Peter the Great and near

the Japanese coast at Stns. I-4, S-49, S-66, S-73 and S-69.

9. 0enqpota candlda (Yokoyama, 1926)

(Figs. 9a-d)

Beta candida Yekoyama, 1926: 261, pl. 34, fig. 1.

Propebela (Tleirritoma) candida, Yokoyama, 1958: pl. 46, fig. 1. Tberritoma candida, Ito, 1985: 30, pl. 8, fig. 6.

Obesotoma sp. Ito, 1967: 57, pl. 5, fig. 6. Prqpebeta sp, No.1 Ito, 1989: Sl, pl. 11, fig. 8. '

The shell is oval-fusiform, from gray to brownish color, consists of 4-5 whorls. An

average size of the shell is 12 mm. (The holotype: H= 13.6, D=6.0 m.) The spiral sculp-

ture is represented by numerous, equal, thread-like ribs with equal spaces between them.

The axial sculpture is represented by oblique, gently sloping, moderate folds. The mainly

developed folds are on the shoulder and on the upper part of the periphery, They faded

away on the base of the shell. They have a small sigmoidal band on the upper part of

the shoulder. There are 18 to 22 folds on the body wherl, 16 to 20 on the penultimate one. Remarks: This species can always be identified by its longitudinal siphonal canal and

the flexious axial folds.

Distribution: The Pacific low boreal species. Distributed only near the Asian coast.

It was found near the Japanese coast at Stn. S-48,

Genus Obasotoma Bartsch, 1941

10. 0besotoma robusta (Packard, 1867)

(Figs. 10a-f)

Bela robusta Packard, 1867: 232, pl. 7, fig, 12. Propebela sp, No.2 Ito, 1985: 30, pl. 7, fig. 8.

Propebela venusta, Ito, 1989: 51, pl. 11, fig. 6, Propebeta sp. No.4 Ito, 1989: Sl, pl. 11, fig. 11.

The shell is turretiform or fusiform, with various hues of brown color, an average size ef the shell with 5-6 whorls is from 9 to 11mm, (The holotype: H=4.5, D= 2.8 mm. The maximum specimen (15,2 mm) was from Japan Sea (De-Kastry)), The

keel is more developed among the different elements of spiral sculpture and it lies on the

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18 VENUS: Vol. 51, Nos,1&2(1992)

shoulder. Ten to twelve fine, hair-like ribs are placed on the sutural ramp. The spiral sculpture on the periphery of the whorl and on the shell base consists of numerous, equal, thread-like ribs. The axial sculpture is represented by angulated, straight and wide folds, which mainly developed on the sutural ramp but they faded away on the base of the shell. There are 16 to 18 folds on the body whorl, 14 to 16 on the penultimate one. The different coloration of sculptural elements is the characteristic to this species. The axial folds and spiral ribs are lighter than the intervals between them, and, as the result, the sculpture is like

a relief.

Remarks: This species is similar to O. woodiana but can be separated by angulated, straight and wide folds that are devcloped on the sutural ramp, and small radular tooth

with rhomboid keel.

Distribution: The Pacific boreal-arctic species. Originally described from Labrador (type locality), but it is distributed mainly in the boreal regions of the Pacific, near the Asian coast. Recorded in the Chukotski Sea near the Wrangel Island, in the northern part of Bering Sea and near the Bering Island, near the coast of West and East Kamchatka,

near Okhotsk coast of Sakhalin, in the Gulf of Peter the Great and in northern part of

Japan Sea. It was found near the Japanese coast at the Stns, S-60, S-49, I-3 and I-4.

11. 0besotoma sotide (Dall, 1886) (Figs. 10g-h, 11a-e)

Lora soiida Dall, 1886: 301, pl. 3, fig. 4. Ttirritoma mitrata, Ito, 1985: 30, pl. 8, fig. 9. 7letrritema candidu, Ito et al., 1986: 18, pl. 21, fig. 5-1.

Obesotoma schantarica, Ito et al., 1986: 19, pl. 24, fig. 5. Obesotoma simpiex, Ito et al,, 1986: 19, pl. 24, fig, 6.

The shell is oval-fusiform, pinkish white or creamy colored. A moderate size of the shell with5-5.5 whorls is from 12 to 13 mm, (The largest shell from the type series: H=13,O, D=7.0, hb.=8,O mm; a large specimen from the Gulf of Peter the Great is H== 18 mm). The spiral sculpture consists of numerous, string-like ribs, which are equal

in size on the periphery and base of the shell and, as a rule, 2-3 times wider than the

spaces between them. The axial sculpture is represented by straight, wide folds, equal

in size on the whorl surface and divided by spaces equal in width. The most developed

folds are on the periphery of the whorl and eroded away on the upper 113 of shell base. There are 13 to 18 folds on the body whorl, 12 to 16 on the penultimate one.

Remarks: This species is morphologically close to O. robusta, but has stronger and wider axial sculpture, conical base and finer radular tooth. Distribution : The Pacific, wide boreal species, distributed mainly near the Asian coast, near the Alaska and western part of Aleutian Islands, in Japan Sea in the Gulf of Peter

the Great and Gulf De Kastry. It was found at the Stns. S-49, I-5 and I-･6.

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Bogdanov-Ito: Oenopotinae from Japan Sea 19

12. 0besotoma hokkaidoensis (Bartsch, 1941) (Figs. 12a-f, 26)

Nlematoma hokkaidoensis Bartsch, l941: 6, pl. 1, fig. 1. Asperdaphne sp., Ito, 1985: 30, pl. 8, fig, 11. Propebeta sp, No. 4, Ito et al., 1986: 18, pl. 23, fig, 2.

The shell is fusiform, white, brownish yellow in the upper part. A moderate size of the sheil with 4-S whorls is frorn 7 to 8mrn. (The holotype: H=9.3, D=5.3mrn. The largest specimen H == 11.6 mm). The spiral sculpture is represented by numerous, fine, thread-like ribs, which are equal in size on the whole surface of the shell. The axial sculp- ture is represented by straight, gently sloping folds, which are developed on the central part of the periphery, and have a sigmoidal band on the shoulder, The spaces are 1.5 times narrower than the folds. The number of folds is very variable, 16 to 24 on the body whorl, and 14 to 18 on the penultimate one.

Remarks: This species is close to O. candicla, but it may be separated by more ovate shell base and short siphonal canal.

Distribution: The Pacific low boreal species. Recorded only near the Asian coast: near southern part of Sakhalin, in the Tartary Strait and in the Gulf of Peter the Great, the eastern part of Japan Sea near south-eastern part of the Hokkaido. We found it from

Stns. S-48, I-2, I-4 and I-6.

13. 0besotoma pulchern'ma n. sp.

(Figs. 13a--b)

The shell is elongate fusiform, yellowish brown, 8.9 mm in height of the holotype. Other dimensions are D=3.9, hb.=5.8, h,=4.5, d=1,4 mm. Paratype H=8.2, D=3.5, hbw=S.4, ha=3.9, d=1,4 mm. The spiral sculpture is represented by two different ele- ments, viz. fine striae on the shoulder and string-like ribs on the other surface of the whorl and the shell base. The ribs are different in size and, as a rule, 2-4 times narrower than spaces between them. The axial sculpture is represented by prominent, straight and wide folds, with spaces 1.5-2 times wider than folds, which are oblique on the periphery 'sigmoidal with a small band on the upper part of the shoulder. There are about 12 folds on the body whorl and on the penultimate one. The most developed folds are on the shoulder and on the upper part of the periphery. They eroded away on the base of the shell. Remarks: This species is similar to Oenqpota pyramidatis, but distinguishes by a more prominent shell base and less numerous axial folds. It differs from Propebela atitakensis by less flexuous axial folds and narrower spiral sculpture elements. Distribution: Collected from Stn. I-4 (37e05,5'N-136025.0'E).

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･ 14. 0besotoma okutanii n. sp.

CFigs. 14a-d)

The shell is fusiform, bright brown, with 4-5 whorls. The holotype: H=8.0, D=4.0, mM･ hb. =6.3, h.=5.1, d=1.7 mm; paratype: H=7.8, D=3.7, hb.=5.8, h.=4,5, d=1,5

The spiral sculpture consists of flat, string-like ribs, which are 6-8 time wider than the

spaces between them. The more developed ribs lie on the shoulder and, as the result, they create the impression that the wide, flat keel is present on the shoulder. The axial

sculpture is represented by semicircular, straight and wide folds, which are most developed on the periphery of the whorl and eroded away on the shell base abapically. The spaces usually are 1.5-2 times narrower than the folds. The folds are oblique on the periphery

and show a faint sigmoidal curve on the sutural ramp, There are 13 to 15 folds on the

body whorl, 12 to 13 on the penultimate one. Remarks: This species can be separated by a slanting shell base and wide axial folds. Distribution: The Pacific low boreal species. Known only near the Asian coast. Type locality is off Ishikawa Prefecture, 36057.0' N-136e21.9' E, depth 255 m (Stn. I-6).

Genus Propebela Iredale, 1918 1941) (=TZtrritoma Bartsch, 1941; Titrritomella Bartsch,

15. RrQpebela (Propebela) nobilis (Moller, 1842) (Figs. 6a-h)

Def}'ancia nobilis Moller, 1842: 12. Pteurotoma turricuta var. nobilis, Morch, 1877: 439. Bela nobilis var, brevispira Loyning, 1931; 15, pl. 2, fig, 26. Rhodopetoma erosa, Ito, 1967: S7, pl, S, fig. 3. 7leirritoma venusta, lto, 1985: 30, pl. 8, fig. 8. 71trritoma candido, Ito et al., 1986: 18, pl, 21, fig. 5-1.

The shell is turretiform, ashy gray, sometimes bright brown, large. An average size of the shell with 6--7 whorls is from 12 to 14 mm. (The holotype: H=7.5 mm. The

largest specimen from the Barents Sea: H;29.0, D=13.9, hb.=19.3, ha=14.1, d==5.0 mm.) The spiral sculpture consists of numerous, string-like ribs, which are equal

in size on the whole shell suTface and, as a rule, l.5-2 times wider than the spaces between them. The keel lies on the shoulder and more developed than the other elements of spiral sculpture. The axial sculpture is represented by prominent, high folds, with 2-3

times wider spaces between them, The folds are equal in size on the whorl surface and

they eroded away on the upper part of the shell base. There are 16 to 19 folds on the

body whorl, 13 to 16 on the penultimate one, Remarks: This species is the closest to R rugulata but can be distinguished by 5-7

thin spiral ribs which lie on the keel, and the presence of an elongate keel on radular tooth. Distribution: The boreal-arctic species. Recorded off the Arctic shore of Alaska, near

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Bogdanov.Ito: Oenopotinae from Japan Sea 21

Greenland, in Labrador and in the Canadian Arctic. In the eastern Atlantic it was found

near Norway and Great Britain, spread in the southeast part of Barentz Sea and near

Spitzbergen, in the White Sea, near the eastern coast of Novaja-Zemlja, near Novosiberian

Islands and in the western part of Laptev Sea and in the Chukotski Sea, near Alaska

and Aleutian, Kemandor, Kurile Islands and Sakhalin. In Japan Sea it is noted in the

Gulf of Peter the Great and in the Tartary straight, and from Stns. S-50, S-60, I-1,

I-2 and I-5.

16. ltqpebela (Ilropebela) assimilis (G.O. Sars, 1878) (Figs. 5a-b)

Bela assimitis G,O. Sars, 1878: 231, tab, 23, fig, 8.

Bela turricuia Tryon, 1884 (non Montagu, 1803): 219, pl, 28, fig. 44.

Bela rugutata var, assimilis Friele, 1886: 5, pl. 9, fig, 9, 10,

Bela rugulata var, schneideri Yokoyama, 1922: 44, pl. 1, fig, 37, Beta candida var, angulata Yokoyama, 1926: 262, pl. 34, fig. 2.

Propebela yokayamai Oyama, 1973: 53, pl, 16, fig. 7. OenQpota scalaris Bouchet et Waren 1980 (non Moller, 1842): 73, fig. 149, 150, Tletrritoma angulata, Ito, 1985: 30, pl. 8, fig. 7; Ito, 1989: 52, pl. 11, fig. 15,

The shell is turretiform, pinkish white or ashy gray, large. A moderate size of the

shell with 5-6 whorls is from 9 to 11 mm. (The holotype H=15 mm. The largest

specimen: H=17.5, D=8.9, hb.=12.2, h,!=8,7, d=4.0 mm.) The keel is the most de-

veloped elements among spiral sculptures and the shoulder is marked. From 5 to 10 equivalent

thread-like ribs are placed on the sutural ramp. The spiral sculpture on the periphery

of the whorl and on the shell base consists of numerous, equal in size, fine ribs, with

equal distances between them. The axial sculpture is represented by round, straight and

middle-sized folds. The mostly developed folds are on the periphery and they eroded away on the shell base. There are 14 to 17 folds on the body whorl, 13 to 14 on the penultimate one.

Remarks: This species can be separated from R scaiaris by stronger axial folds on

the periphery and flattened base of radular tooth.

Distribution: The boreal-arctic species. It is found in the boreal regions of the North

Atlantic and in the Bering, Okhotsk and Japan Seas. It was found at Stns. S-50 and S-60.

17. Propebela

(Figs. 17a-d, 27)

Propebeta venusta Okutani, 1964: 429, pl, 7, fig, 8,

The shell is small, subfusiform, dirty white. A moderate size of the shell is 9-10 mm.

(The holotype: H=10.4, D=5.1 mm.) The spiral sculpture is represented by two type

ofelements, viz. 3-5 thin, hair-like ribs on the sutural ramp, and numerous, equal-sized

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22 VENUS: Vol. 51, Nos,1&2(1992)

thread-like ribs, which are, as a rule, 1,5-2 times wider than the spaces between them.

The axial sculpture is represented by the gently sloping, fine folds, they are obsolete on

the upper part of base. They are equal in size and with spaces between them. There are 18 to 22 folds on the body whorl, 16 to 21 on the penultimate one.

Remarks: This species can be distinguished from Propebela exquisita by a gently slop-

ing axial folds.

Distribution: The Pacific lowboreal species. Recorded only from near the Asian coast.

The type locality is Sagami Bay, 35001.0' N--139"19.6' E. It was found at Stns, I-2,

I-3 and S-48.

18. PrQpebela (Rropebela) exquisita Bartsch, 1941

(Fig. 18)

Propebela (Titrritotna) exquisita Bartsch, 1941: 12, pl. 1, fig. 4. Propebela (7iirritoma) exquisita, Ito, 1967: 58, pl, 5, fig, 9,

The shell is turretiform, cream-colored, with muddy-brown hue on the shoulder, mid-

dle size. (The holotype: H=10.0, D=4.9 mm) The spiral sculpture consists of middle- sized, equivalent thread-like ribs on the periphery and base. The keel is marked the shoul-

der, 8-10 thin, hair-like ribs lie on the sutural ramp. The axial sculpture is represented

by the gently sloping, middle-sized, straight folds, which most develope on the periphery

of the whorl and obsolete on the shell base. They are equal in size to spaces between

them. The folds are oblique on the periphery and have a slight sigmoidal band on the

sutural ramp. There ar ¢ 13 to 16 folds on the body whorl, 12 to 15 on the penultimate one.

Remarks: This species is similar to P. ruguiata but can be separated by higher spire

and numerous ribs on the sutural ramp,

Distribution: The Pacific lowboreal species. The type locality is Albatross Station

3738. It was collected from Stn. r-2.

19. Rropebela (Cbnetoma) viridula (Moller, 1842) (Figs. 21a-k)

Def)'ancia viriduta Moller, 1842 (non 7)"itonium viridutum Fabricius, 1760): 14. Beia arctica Adams, 1855: 121.

Bela kobeiti Verkruzen, 1875: 238.

Propebeta sp. No, 1, Ito, 1985: 30, pL 7, fig. 7; No. 2, Ito et al., 1986: 18, pl. 22, fig. 6; No. 3, No. 6, Ito, 1989: 51, pL 11, fig, 10, 13.

The shell is oval-fusiform, from ashy grey to bright chestnut in color, middle size. . (The holotype H=4.3 mm. Thelargest specimen from the Barents Sea; H=14.3, D;7.0, hb.= 10.3, h,= 7.7, d=3.0 mm.) The spiral sculpture on the whole shell surface consists

of numerous, equal, string-like ribs, with equal spaces between them. The axial sculpture

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Bogdanov.Ito: Oenopotinae from Japan Sea 23

consists of gently sloping, wide folds. There is a small bend at anterior part of the sutural ramp. There are 18 to 25 folds on the body whorl, 16 to 20 on the penultimate one. Remarks: This species is close to P. variabilis, but can be separated by higher body

whorl and numerous axial folds.

Distribution: The borear-arctic species, Recorded in the Canadian Arctic, near western

Greenland, in the North Atlantic in the Arctic Basin, in south part of Barentz Sea, in the White Sea and near southern coast of Novaja-Zemlja, in the Bering, Okhotsk and Japan Seas. It was found at Stns. I-1, I-2, I--5, I-6, S-50, S-58, S-60 and S-66.

20. Propebela (thnetoma) concinnula (Verrill, 1882)

(Figs. 19a-b)

Bela exarata Verrill, 1880 (non Moller, 1842): 336.

Bela concinnula Verrill, 1882: 468, pl. 43, fig. IS, pl. 57, fig, 11.

Bela concinnula var. acuta Verrill, 1882: 470, pl. S7, fig. 10.

Propebela sp, No,7, Ito, 1985: 30, pl. 8, fig. 1.

P. komakahide, Ito, 1985: 30, pl. 8, fig. 5.

The shell is fusiform, of ashy color, an average size of the shell is from 9 to 10 mm. (The largest shell in the type series: H=11.5, D=5.3, h.==8,O, d=2.0 mm; the middle

size one: H=10.0, D=4.5, h.=5.5 mm. The shell with height 12.5 mm was found near the coast of West Kamchatka.) The spiral sculpture consists of different elements: 4-6

thread-like ribs on the sutural ramp, and numerous, equal in size, string-like ribs on the

periphery and base of the shell. The axial sculpture is represented by numerous, round,

medium-sized folds, which are equivalent on the whorl surface and divided by the equal

in width. The folds have a small bend at anterior part on the sutural ramp. The thicken-

ing is formed at the point of crossing of axial and spiral elements. There are 20 to 28

folds on the body whorl, 14 to 17 on the penultimate one. Remarks: This species is nearest to P. mitrata, but can be distinguished by high spire,

axial and spiral elements forming thick knots on crossing points,

Distribution: The amphiboreal widely spread species. It is known from all boreal

waters of the Pacific Ocean and high-boreal ones of the western Atlantic. It was found

from New England to Nova Scotia, near the Newport and Cape Cod, in the Casco Bay

and Massachusetts Bay. In the Pacific Ocean it is known from near the Bering Island

and near the coast of West Kamchatka. It was collected near the Japanese coast at the

Stn. S-66.

21. Propebela (Canetoma) mitrata (Dall, 1919) (Figs. 20a-b)

Lora mitrata Dall, 1919: 43, pl. 16, fig. 1.

The shell is turretiform, grey, large. (The holotype: H=24.0, D=11.0, hb.=16.0 mm.)

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24 VENUS: Vol.Sl, Nos.1&2(1992)

The spiral sculpture is represented by numerous, string-iike ribs, which are equal in size on

the whole shell surface, and, asarule, 1.5-1.7 times wider than the space$ between them,

The axial sculpture is represented by prominent, fine folds, with 1.5-2 times narrower spaces

between them. The folds are equivalent on the whorl surface and they become obsolete only

on the lower part of the base. There are 24 to 26 folds on the body whorl, 16 to 18 on the

penultimate one. The elements of spiral and axial sculptures are equal in size, and, as the result of crossing, they produce characteristic reticulate pattern on the shell surface. Remarks: This species is morphologically close to P. exarata, but can be separated by

fine prominent axial folds, characteristic reticulate pattern on the shell surface and the typical triangular flag at apical point of the radular tooth.

Pistribution: The Pacific, boreal, widely spread species from near American coast, Bering

Strait to Schumagin Island, near the Kurile Islands and Okhotsk coast of Sakhalin. It was

found at Stn. I-3.

To review Oenopotinae living in the southeastern part of Japan Sea, a list of species reported

from Ishikawa and Niigata Prefectures are given here based on the photographed specimens by Ito (1967, 1985) and Ito et al. (1986).

22. Curtitoma hebes

Propebeta sp. No. 4, Ito, 1985: 30, pl. 7, fig. 10.

Propebela sp. No. 5, Ito et al,, 1986: 18, pl. 23, fig. 3, 4.

23. 0enopota impressa (Beck in Morch, 1868)

Propebeta sp. No. 6, Ito et al., 1986: 18, pL 23, fig. 5.

24. 0besotomaJ'aponica Bartsch, 1941

Obesotoma hanazakiensis, Ito, 1967: 57, pl. 5, fig. 4.

25. 0besotoma simplax (Middendorff, 1849)

Obesotoma sp., Ito, 1967: 57, pl. 5, fig. 5.

26. thqpebela (thnetoma) tet:sa (Bartsch, 1941)

Propebeta sp. No, 3, Ito, 1985: 30, pl. 7, fig. 9.

7)trritomayokayamai, Ito et al., 1986: 18, pl. 22, fig. 3, 4.

27. Rrqpebela (Canetoma) tayaensis (Nomura et Hatai, 1938)

Propebela sp. Ito, 1967: 57, pt. 5, fig, 8.

Those reported by Ito (1985, 1989) and Ito et al. (1986) as subfamily Oenopotinae are

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25 Bogdanov ・lto ： Oenopotinae from Japan Sea

excluded from this subfamily ：

： 8 fig．4 ． Propebeta sp 。No 、且0 ，Ito，1985 30，pL ， Venustoma tacunosa ， Ito et al ．， 1986 ： 51 ， pL 10， fig． 12，

No 2 Ito et aL 1986 ： 11 fig．9 ． ProPebela sp ． ， ， ， 51，p1． ，

1989 ： 17 22 ．1 ． Venustoma tacunosa，lto， ， pL ，fig ，2

要 約

主 と し て 著 者 の 1 人伊 藤 （1967，1985，1989） 及 び 伊 藤他 （1986） が 日 本 海 か ら報告 し た フ タ マ の ． る こ と が た が そ の ン ジ ガ イ亜 科 Oenopotinae の 諸種 を再 検 討 し た 結呆 27種 種 名 を確定す 出 来 ，

う ち 次 の 21種 に つ き記載 と 図示 を行 っ た 。 1． Cttrtitoma violacea （Mighels ＆ Adams ，1842）

Vemaiom α sp P α opehela sp ． ，8 fig．3 ； 同 1986 ）で ？ 伊 藤 （1985）が AJematoma ．（pL7 ，fig．4）， （pl ， ） （ ？ sp ． ．12 fig．7）と し た 種 。 sp ． no ．3 （pl．21 ，fig．4）； 同 （1989） Benthomangelia （p1 ， 2 ． C ．decussata（CouthoLLy，1939） ． 貸 ． と た 種 伊 藤 （1989） が Nenzatoma dissotuta〔pl 12 ， g 23 ） し 。 3 ． （；− becktenTishevi Bogdanov ，1989 Paopebela sp ．No ．3 ．23 伊藤 （1985， 1989） が Nematoma hokkaidoensis ，伊藤 他 （1986） が （pl ， fig，1）と し た 種 。 4． C ．trevettiana （Turton ，1834） ． ’ 伊 藤 （1985 ） が Pro ρehela sp ． N o ，1 （pl．7 ， fig．7）と した種 。 5 ． C ．niigataensis n ． sp ．（碧俸巨） No ． ． ． た 種 伊藤 （1985）が Propebeta sp ． 6 （pl 7， fi g 12）と し 。

6 ． Granotoma albre ‘rhti （Krause ．1885） 伊 藤 （1985） が Nematoma dissolutaに ，伊 藤 他 （1986 ） が rVematoma sp ．〔pl．21 ，11g．3）と し た 種 。 7． （｝．raduga （Bogdanov ，1985）

o と た 種 伊 藤他 （1986） が Proρebeta sp ． N ．4 （p1．23 ，fig．4 ） し 。 8 ． Oenopota caneellata （Mighels ＆ Adams ，1840） ． fig が Propebeta 伊 藤 （1985）が Propebeta sp ． Nos ．5 ，6 ，8 （pl．7 ， ．11，12 ，p1．8，fig．2），伊藤 他 （1986） sp ．（pl．22 ， fig．5）， Obestoma hanazakiensisとし た 種 。 9 ． 0 ．candida （Yekoyarna ，1926 ）コ ウ シ フ タ マ ン ジ ガ イ Pro ehela sp ． 、11 fig，8 と され 伊藤 （1967） で は obestoma sp ．（pl．5， fig．6），同 〔1989） で は ρ （pl ， ） た 。 10． Obestoma robusta （Packard，1867） fig sp Propehela sp ． No ．4 ， ．11） 伊 藤 （1985） で は Propebela ．（pl．7 ，fig．8）， 同 （1989） で は （pL11

と ，P ． venust α と 同定 さ れ た 種 。 11 ． 0 ，solida （DalL 1886＞

T c α ndida sehantarica O ． 伊 藤 （1985） が Turritoma mitrata ，伊 藤 他 〔1986） が ． ， Obestoma ，

simplex と し た 種 。 ． 12 ． 0 ，hokkaidoensis（Bartsch．1941）ミ ドリ フ タ マ ン ジ ガ イ ． Pro eheta sp ． ．23 fig．2）と 伊 藤 （1985） が ASperdaphne sp ．（pi．8，ng ，11），伊 藤他 （1986） が ρ （pi ， し た 種 。

13． O ． pulcherrima n ． sp ．（新 種 ） 伊藤他 （1986 ） が Propebela sp ． No ．1 （pL22 ， fig．5）と した種 。

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26 VENUS: Vel. 51, Nos.1&2(1992)

14. 0. okutanii n. sp. (M$fi) IJ+ee (1985) ht Turritoma angulata (pl.8, fig.7) t Lt:pt.. 15. Propebela (Propebela) nohifis (Moller, 1842) 'r'li TC'IS f#ma (1967) abfRhodopetomaerosa, ft (1985) Turritoma venusta, I#mafill (1986) Z candida t " tLkfiIL.

16. PL (P.) assin7itis (G. O. Sars, 1878) '(T' f#ec (1985, 1989) Turritoma anguLata t"nrt:. +("IS:1'IL< i'.='li 17. P, (PL) verustaOkutani,1964 pawh (1985) [lfi"ttv(L'aftiabL IJFwh (1989) TZtrritoma candida (pl.11, fig.l4) 2[ StLk.

18. P. (PL) exquisita BaTtsch, 1941 e,ee (1987) T'SiEL<1dl5E'3ft(vik. 19. P. (Canetoma) viridula (Moller. 1842) '("eS 't'Sraec ilfwh (1985) Propebela sp. No.1 (pl.7, fig.7), IYma (1986) (pl.22, fig,6) t5 "C'l)IFofct tl., f'Yes{te (1989) (pl.li, fig,10,13) 2 3iLkle'..

20. P. (Cl) eoncinnula (Verrill. 1882) JOCi ilfma (1985) Prqpebela sp. (pl.8, fig.l) Jts2L"' I'. komakahida 2 "2Lh$rft. 21. P. (C) mitrata (Dall, 1919> -C'Ci fP'ma C1985) Tberritoma mitrata t StLkElll.

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Bost, J, 7Vdt. Hist.,4 (1): 37-54. Moller, H.P.C. 1842. Ihdex M/oUuscorum Groenlandiae. Hafniae. 24 pp. Mbrch, O.A.L. 1852. Catatogus conchyliorum quae retiquit D. Alphonso Dtt4guirra Gaclea comes

de Yoldii. Hafniae, 170 pp. M6rch, O.A.L. 1868, Catalogue des mollusques du Spitzberg, requeillis par le Dr. H. Kroyer pendant le voyage de la corvette la Recherche en juni 1838. Mem. Soc. matacol, Beiguique. 4: 7-32, M6rch, O.A.L. 1877. The Greenland Fbuna IV. R, Brown (ed.) London. Appendix: 435-442.

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fap. Jburn, GeoL Geogr. 16 (1): 39-61. Okutani, T. 1965. Report on the archibenthal and abyssal gastropod mollusca mainly collected from Sagami Bay and adjacent waters by the R.V. Soyo-Maru during the years 195S-1963. JOur. of Ric. Sei.. Univ. 7bk),e, sect. II, 15: 171-447.

Okutani, T. 1966. Archibenthal and abyssal mollusca collected by the R.V. Soyo-Maru from Japanese waters during 1964. BuU. Tbkai Reg. }ilsh. Res, Lab. 46: 1-27. Oyama, K. 1973. Revision ef Matajuro Yokoyama's Type Mollusca from the Tertiary and Quater- nary of the Kanto area. Paleonto. Soc. lapan. sp. paper, 17: 1-148. Packard, A.S. 1867. 0bservations on the glacial phenemena of Labrador and Maine, with a yiew ofthe Recent invertebrate fauna ofLabrador. Mem, Bost, Soc. Ndt, Hist., n.s. 1 (2): 210-303. Sars, G.O. 1878. Mollusca regionis arcticae Nbrvegiae. Bedrag tit Kunctskaben om Not:ges Arktiske

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Sars, M. 1868. Fbrtsatte bemerkninger over det clyriske iivs udbredningihavets tlybder. Forhandl,

Vidensk Kristiania. 246-275,

Strom, H. 1788. Beskrivelse over norske insecter, Kong, Dan. IXidensk. Selsk. Skrijtr. 3: 296-297,

Troschel, F.H. 1866, Das Gebiss der Schnecken. Berlin, Nicolaische Verlagsbuchhandlung, 2: 1-48,

Tryon, G.W. 1884. Manuat of conchology: , Pleurotomidae. Vol. 6: 1-413, 7-tirton, PVL i834. Description ofsome nondescript and rare British species ofshells, Londonls Mag.

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Verkruzen, T.A, 1875, Bericht uber einen Schale-Ausflug im Somrner 1874. Jahrb. Deutsche

MalakozooL Gex 2: 229-240.

Verrill, A.E. 1880. Notice of recent additions to the marine invertebrata of the northeastern coast

of America, with descriptions of new genera and species and critical remarks en others. Proc.

U.S. ?VllL Mus. 3: 3S6--405,

Verrill, A,E. 1882. Catalogue of rnarine mollusca. Added to the fauna of New England during

the past ten years. 7)'ans. Con. Acad. 5: 447-587.

Yokoyama, M. 1922. Fossils from the Upper Musashino of Kazusa and Shimosa, L ColL Sci. U)Tiv.

71)k;po, 44 (1): 1-200,

Yokoyama, M.1926. Fossil shells from Sado. Jour. fuc. Sci., Imp. Uhiv. Tbk],o,2(1): 249-312,

Yokoyarna, M. 1958. Tertiary fossils from various localities in Japan. Part II, Paiaeon. Soc. Jdpan.

sp. papers. 4: 2S-S7. [Received: OctobeT 1, 1991]

Figs. Ia-e. Curtitoma violacea (Mighels & Adams, 1842)

a. x7, S67-59; b. x7, S67-S3; c,d. x6.5, S67-59;

e. x7, S67-59.

Figs. 2a,b. Curtitoma decussata (Couthoy, 1839)

a. x6, I-5; b. x7, I-S.

Figs. 3a-g. Curtitoma becktemishevi Bogdanov, 1989.

a, x7, I-3; b, x8, I-3; c,d. x7. I-2; e. x7.5. I-2;

f.g. x7, I-2.

Fig. 4. Curtitoma niigataensis n,sp, x6.5, S-49.

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Figs. 5a-c. Curtitoma trevelliana (Turton, 1834)

a, x8. I-3; b,c. x7.S, I-3.

Figs. 6a-d. Granotoma albrechti (Krause, 1885)

a. x6, S-52; b, x7.5, S-S2; c, d. x8, S-S2.

x7, ,Fig. 7.Figs. Granotoma raduga (Bogdanov, 1985) S-58.

8a-h. Oenopota caneeltata (Mighels & Adams, 1840)

a. x5.5, S-69; b. x4,5, S-69; c,d. x6, S-69; e. x5, I-.4;

f. x5, I-4; g. x5.5, S-69; h. x5.5, S-69.

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Figs. 9a-d. Oenopota candida (Yokoyama, 1926)

a. x4,5, S-48; b. x4, S-48; c,d. x4.5, S-48,

Figs. 10a-f. Obestoma robusta (Packard, 1867)

a, b, x4.5, S--60; c,d. xS, I-3; e,f. x5, S-49.

Figs. 10g-h, 11a-c. Obestoma solida (Dall, 1886)

10-g,h. x4.5, I-4; 11-a, x5, I-6; b,c. x5, S-49;d,e. x5.5,I-5.

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Figs. 12g-f, Obestoma hokkaidoensis (Bartsch, 1941)

a. x6, I--4; b. x7, S-45; c. x6, S-48; d. x7, S-48;

e. x7, S-48; f. x6, S-48.

Figs. 13a,b. Obestoma puicherrima n,sp.

a, x6, I-4; b, x7, I-4.

Figs. 14a-d. Obestoma okutanii n.sp.

a, b. x4, I-6; c. x4,, I-6; d. x4, I-6

Figs.15a,b. Propebela (Propebela) assimitis (G,O. Sars, 1878)

a. xS, I-60; b. xs.s, S-6o.

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Figs. 16a-h. Rropebela (Propebela) nobitis (Moller, 1842)

a. x4, S-S6; b. x4.5, S-50; c,d. x3.5, S-50; e. x4,ST50;

f. × 4, I-5; g,h. x4, I-5.

Figs. 17a-d. PrQpebela (Propebela) venusta Okutani, 1964

a. x5, I-4; b, x5, I-3; c, xS, I-2; d. x6, I-2.

Fig. 18. Ilropebela (Propebela) exquisita Bartsch, 1941

x5.5, I-2.

Figs. 19a,b. PrQpebeta (Canetoma) concinnuta (Verrilr, 1882)

x5, S-66,

Figs. 20a,b. Propebela (Canetoma) mitrata (Dall,, 1919)

x4, I-3.

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Figs. 21aLj. Propebeia (Canetoma) viridula (Moller, 1842)

a,b. x5, I-2; c, x5.5, I-2; d. x4.5, S-58; e. x5, 1-2;

f. x5.5. S-58; g,h, x5, 1-2; i,j. x55, I-2.

Fig. 22. Curtitema violacea. Sculpture enlarged, (Scale=100 ptm).

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Figs.23-27. Protoconchs (Scale=100 "m) [except Fig. 24a]

23. Curtitoma becklemishevi

24b. Granostema atbrechti

25. Oenopota cancellata

26. Obestoma hokkaidoensis

27.24a.Propebeta venusta

Fig. Shell of Granostoma albrechti (Scale=1 mm).

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