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NAYARIT, MÉXICO 1 COMMON OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit,UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria”, Nayarit

Photos by: Jesús Loc, Antonio Robles, Jonathan Vargas, David Molina. Acknowledgments. To Emmanuel Miramontes, Carlos Villar, Stefanny Villagómez and Héctor Franz for the support of several photos indicated in the main text and to Tatzyana Wachter for the improvements to the document. © Jesús Alberto Loc-Barragán [[email protected]], José Antonio Robles-Martinez [[email protected]], Jonathan Vargas-Vega [[email protected]] and David Molina [[email protected]] [fieldguides.fieldmuseum.org] [921] version 1 8/2017

Signs: (R) = residente/resident, ( MI) = winter migratory, (SR) = summer resident; (♂) = Macho/Male, (♀) = Hembra/Female, (J) = Juvenil/Juvenile. Status of concern () based on NOM-059-SEMARNAT-2010: PR: special protection; A: threatened; P: extinction risk; IUCN, LC: least concern; NT: near threatened; Endemism, E: endemic, CE: nearly endemic, SE: semiendemic, I; exotic, invasive. The numeric values are the Vulnerability index, which takes into account parameters like population size, geographic distribution, seasonal threats and population trend; index values vary from 4 until 20 and a higher value implies greater vulnerability (Panjabi et al., 2005; Berlanga et al. 2015).

Marismas Nacionales Biosphere Reserve and Birds

In northwest Mexico, Marismas Nacionales, an extensive estuarine system, it has been historically recognized for its importance for birds, especially waterfowl, shorebirds, herons and coastal birds like gulls and terns (Leopold, 1959; Morrison et al., 1994; Ortega-Solís, 2011).

The importance of Marismas Nacionales is recognized by numerous institutions, conservationist groups and government agencies. In 1992, the Western Hemispheric Shorebird Group decreed Marismas Nacionales as a site of International Importance for shorebirds, because it is used by at least 100,000 shorebirds during winter season (Morrison et al., 1992; Morrison et al., 1994). The Ramsar Convention (1995) designated it as a Ramsar site, for its value to human communities and wildlife. Subsequently, in 1998 it was included in the list of Important Areas (IBA) of Mexico (IBA C-65). In addition, the North America Bird Conservation Initiative (NABCI) recognizes Marismas Nacionales as one of the most important sites for migratory bird conservation at a continental level. In addition, the National Commission for the Knowledge and Use of Biodiversity considers it as a marine, terrestrial and hydrological priority region (CONABIO, 2001). Finally, the Federal Government decreed it as a Natural Protected Area, under the category of Biosphere Reserve (DOF, 2010), for its forests and its importance for wildlife; it is noteworthy to say that also for the constant human use and exploitation of its resources (CONANP, 2005).

However, the estimated bird richness in the area depends on the scale and the limits considered. Taking into consideration the Biosphere Reserve, richness reaches around 288 species (CONABIO, 2017), but Birdlife (2017) recognizes around 396 species, in a more extensive area. This represents 53-72% of the total bird richness in the state of Nayarit (Jacobo 2014). Nevertheless, increasing study and observation effort has yielded new species records that have gradually increased the local richness (e.g. White-winged Scoter, Hooded Merganser, Sandhill Crane and American Flamingo; eBird 2017).

From 2010 to the present year, we conducted several surveys in the Biosphere Reserve Marismas Nacionales and in other adjacent areas outside the reserve but connected. We documented with photographs 164 species of birds, which are relatively common in the reserve. We intend this photographic guide to serve as a tool for students, professionals, decision makers, stakeholders, birders and others who are interested in bird diversity and the conservation of the area.

2 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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1 Dendrocygna autumnalis 2 Anas discors ♂ 3 Anas cyanoptera ♂ 4 Anas clypeata ♂ Black-bellied Whistling-Duck Blue-winged Teal Cinnamon Teal Northern Shoveler Pijije Ala Blanca Cerceta Ala Azul Cerceta Canela Pato Cucharón Norteño (R) (LC) (8) (MI) (LC) (8) (MI, R) (LC) (10) (MI) (LC) (7) Jesús Loc Antonio Robles Antonio Robles Jesús Loc ANATIDAE ANATIDAE ANATIDAE ANATIDAE

5 Anas acuta ♂ 6 Anas crecca ♂ 7 Aythya affinis ♂ ♀ 8 Ortalis wagleri Northern Pintail Green-winged Teal Lesser Scaup Rufous-bellied Chachalaca Pato Golondrino Cerceta Ala Verde Pato Boludo Menor Chachalaca Vientre Castaño (MI) (LC) (11) (MI) (LC) (6) (MI) (LC) (12) (R) (LC) (E) (15) Antonio Robles Emmanuel Miramontes Antonio Robles Antonio Robles ANATIDAE ANATIDAE ANATIDAE CRACIDAE

9 Callipepla douglasii ♂ ♀ 10 Tachybaptus dominicus 11 Podiceps nigricollis 12 Patagioenas flavirostris Elegant Quail Least Grebe Eared Grebe Red-billed Pigeon Codorniz Cresta Dorada Zambullidor Menor Zambullidor Orejudo Paloma Morada (R) (LC) (E) (14) (R) (PR) (8) (MI) (LC) (9) (R) (LC) (12) Jesús Loc Antonio Robles Antonio Robles Antonio Robles ODONTOPHORIDAE PODICIPEDIDAE PODICIPEDIDAE COLUMBIDAE

3 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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13 Streptopelia decaocto 14 Columbina inca 15 Leptotila verreauxi 16 Zenaida asiatica Eurasian Collared-Dove Inca Dove White-tipped Dove White-winged Dove Tórtola Turca Tórtola Cola Larga Paloma Arroyera Paloma Ala Blanca (R) (LC) (I) (5) (R) (LC) (8) (R) (LC) (8) (R, MI) (LC) (8) Antonio Robles Antonio Robles Antonio Robles Jesús Loc COLUMBIDAE COLUMBIDAE COLUMBIDAE COLUMBIDAE

17 Piaya cayana 18 Coccyzus minor 19 Crotophaga sulcirostris 20 Chordeiles acutipennis Squirrel Cuckoo Mangrove Cuckoo Groove-billed Ani Lesser Nighthawk Cuclillo Canela Cuclillo Manglero Garrapatero Pijuy Chotacabras menor (R) (LC) (9) (R) (LC) (14) (R) (LC) (7) (MI, R) (LC) (8) Antonio Robles Antonio Robles Antonio Robles Antonio Robles CUCULIDAE CUCULIDAE CUCULIDAE CAPRIMULGIDAE

21 Nyctidromus albicollis 22 Nyctibius jamaicensis 23 Cynanthus latirostris 24 Amazilia rutila Common Pauraque Northern Potoo Broad-billed Hummingbird Cinnamon Hummingbird Chotacabras Pauraque Bienparado Norteño Colibrí Pico Ancho Colibrí Canela (R) (LC) (7) (R) (LC) (14) (R) (LC) (SE) (10) (R) (LC) (10) Jesús Loc Antonio Robles Antonio Robles Antonio Robles CAPRIMULGIDAE NYCTIBIDAE TROCHILIDAE TROCHILIDAE

4 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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25 Amazilia violiceps 26 Rallus obsoletus 27 Porzana carolina 28 Porphyrio martinicus Violet-crowned Hummingbird Ridgway's Rail Sora Purple Gallinule Colibrí Corona Violeta Rascón Costero del Pacífico Polluela Sora Gallineta Morada (R) (LC) (SE) (10) (R) (NT) (14) (MI) (LC) (10) (R) (LC) (11) Antonio Robles Antonio Robles Antonio Robles Antonio Robles TROCHILIDAE RALLIDAE RALLIDAE RALLIDAE

29 Gallinula galeata 30 Fulica americana 31 Aramus guarauna 32 Himantopus mexicanus ♀ Common Gallinule American Coot Limpkin Black-necked Stilt Gallineta Frente Roja Gallareta Americana Carao Candelero Americano (R, MI) (LC) (8) (MI, R) (LC) (11) (R) (A) (11) (MI, R) (LC) (11) David Molina Antonio Robles Stefanny Villagòmez Jesús Loc RALLIDAE RALLIDAE ARAMIDAE RECURVIROSTRIDAE

33 Recurvirostra americana ♂ 34 Haematopus palliatus 35 Pluvialis squatarola 36 Charadrius collaris ♂ American Avocet American Oystercatcher Black-bellied Plover Collared Plover Avoceta Americana Ostrero Americano Chorlo Gris Chorlo de Collar (MI) (LC) (12) (R) (P) (13) (MI) (LC) (12) (R) (LC) (12) Jesús Loc Antonio Robles Antonio Robles Antonio Robles RECURVIROSTRIDAE HAEMOTOPODIDAE CHARADRIIDAE CHARADRIIDAE

5 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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37 Charadrius nivosus 38 Charadrius wilsonia ♂ 39 Charadrius semipalmatus 40 Charadrius vociferus Snowy Plover Wilson's Plover Semipalmated Plover Killdeer Chorlo Nevado Chorlo Pico Grueso Chorlo Semipalmeado Chorlo Tildío (MI, R) (A) (14) (R, MI) (LC) (14) (MI) (LC) (9) (R, MI) (LC) (9) Jonathan Vargas Antonio Robles Stefanny Villagomez Antonio Robles CHARADRIIDAE CHARADRIIDAE CHARADRIIDAE CHARADRIIDAE

41 Jacana spinosa 42 Numenius phaeopus 43 Numenius americanus 44 Limosa fedoa Northern Jacana Whimbrel Long-billed Curlew Marbled Godwit Jacana Norteña Zarapito Trinador Zarapito Pico Largo Picopando Canelo (R) (LC) (11) (MI) (LC) (13) (MI) (LC) (15) (MI) (LC) (15) Jonathan Vargas Jesús Loc Jesús Loc Carlos Villar JACANIDAE SCOLOPACIDAE SCOLOPACIDAE SCOLOPACIDAE

45 Arenaria interpres 46 Calidris canutus 47 Calidris himantopus 48 Calidris alba Ruddy Turnstone Red Knot Stilt Sandpiper Sanderling Vuelvepiedras Rojizo Playero Canuto Playero Zancón Playero Blanco (MI) (LC) (11) (MI) (P) (15) (MI) (LC) (14) (MI) (LC) (14) Stefanny Villagomez Antonio Robles Antonio Robles Antonio Robles SCOLOPACIDAE SCOLOPACIDAE SCOLOPACIDAE SCOLOPACIDAE

6 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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49 Calidris minutilla 50 Calidris mauri 51 Limnodromus griseus 52 Actitis macularius Least Sandpiper Western Sandpiper Short-billed Dowitcher Spotted Sandpiper Playero Chichichuilote Playero Occidental Costurero Pico Corto Playero Alzacolita (MI) (LC) (11) (MI) (LC) (15) (MI) (LC) (13) (MI) (LC) (9) Antonio Robles Antonio Robles Antonio Robles Jesús Loc SCOLOPACIDAE SCOLOPACIDAE SCOLOPACIDAE SCOLOPACIDAE

53 Tringa solitaria 54 Tringa melanoleuca 55 Tringa semipalmata 56 Tringa flavipes Solitary Sandpiper Greater Yellowlegs Willet Lesser Yellowlegs Playero Solitario Patamarilla Mayor Playero Pihuiuí Patamarilla Menor (MI) (LC) (12) (MI) (LC) (10) (MI) (LC) (14) (MI) (LC) (12) Jesús Loc Antonio Robles Antonio Robles Antonio Robles SCOLOPACIDAE SCOLOPACIDAE SCOLOPACIDAE SCOLOPACIDAE

57 Phalaropus tricolor 58 Leucophaeus atricilla 59 Larus heermanni 60 Larus delawarensis Wilson's Phalarope Laughing Gull Heermann's Gull Ring-billed Gull Falaropo Pico Largo Gaviota Reidora Gaviota Ploma Gaviota Pico Anillado (MI) (LC) (11) (MI, R) (LC) (8) (R) (PR) (NT) (SE) (16) (MI) (LC) (6) Antonio Robles Antonio Robles Antonio Robles Antonio Robles SCOLOPACIDAE LARIDAE LARIDAE LARIDAE

7 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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61 Larus californicus 62 Sternula antillarum 63 Gelochelidon nilotica 64 Hydroprogne caspia California Gull Least Tern Gull-billed Tern Caspian Tern Gaviota Californiana Charrán Mínimo Charrán Pico Grueso Charrán Caspia (MI) (LC) (12) (SR) (PR) (16) (R, MI) (LC) (14) (MI) (LC) (10) Antonio Robles Antonio Robles Antonio Robles Stefanny Villagómez LARIDAE LARIDAE LARIDAE LARIDAE

65 Chlidonias niger 66 Sterna forsteri 67 Sterna hirundo 68 Thalasseus maximus Black Tern Forster's Tern Common Tern Royal Tern Charrán Negro Charrán de Forster Charrán común Charrán Real (MI) (LC) (12) (MI) (LC) (12) (MI) (LC) (12) (MI, R) (PR) (12) Antonio Robles Antonio Robles Antonio Robles Jesús Loc LARIDAE LARIDAE LARIDAE LARIDAE

69 Rynchops niger 70 Mycteria americana 71 Fregata magnificens ♂ ♀ 72 Sula nebouxii Black Skimmer Wood Stork Magnificent Frigatebird Blue-footed Booby Rayador Americano Cigüeña Americana Fragata Magnìfica Bobo Pata Azul (MI, R) (LC) (14) (R) (PR) (13) (R) (LC) (16) (R) (PR) (16) Antonio Robles Jesús Loc Jonathan Vargas Héctor Franz LARIDAE CICONIIDAE FREGATIDAE SULIDAE

8 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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73 Phalacrocorax brasilianus 74 Anhinga anhinga ♀ 75 Pelecanus erythrorhynchos 76 Pelecanus occidentalis Neotropic Cormorant Anhinga American White Pelican Brown Pelican Cormorán Oliváceo Anhinga Americana Pelícano Blanco Pelícano Pardo (R) (LC) (16) (R) (LC) (10) (MI) (LC) (12) (R) (LC) (11) Jesús Loc Jesús Loc Jesús Loc Jesús Loc PHALACROCORACIDAE ANHINGIDAE PELECANIDAE PELECANIDAE

77 Tigrisoma mexicanum 78 Ardea herodias 79 Ardea alba 80 Egretta thula Bare-throated Tiger-Heron Great Blue Heron Great Egret Snowy Egret Garza Tigre Mexicana Garza Morena Garza Blanca Garceta Pie Dorado (R) (PR) (13) (MI, R) (LC) (7) (MI, R) (LC) (7) (MI, R) (LC) (8) Antonio Robles Jesús Loc Jesús Loc Jesús Loc ARDEIDAE ARDEIDAE ARDEIDAE ARDEIDAE

81 Egretta caerulea 82 Egretta tricolor 83 Egretta rufescens 84 Bubulcus ibis Little Blue Heron Tricolored Heron Reddish Egret Cattle Egret Garceta Azul Garceta Tricolor Garceta Rojiza Garza Ganadera (MI, R) (LC) (13) (MI, R) (LC) (12) (MI, R) (PR) (18) (R, MI) (LC) (I) (6) Jesús Loc Antonio Robles Jesús Loc Antonio Robles ARDEIDAE ARDEIDAE ARDEIDAE ARDEIDAE

9 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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85 Butorides virescens 86 Nycticorax nycticorax 87 Nyctanassa violacea 88 Cochlearius cochlearius Green Heron Black-crowned Night-Heron Yellow-crowned Night-Heron Boat-billed Heron Garceta Verde Pedrete Corona Negra Pedrete Corona Clara Garza Cucharón (R, MI) (LC) (11) (R, MI) (LC) (10) (R, MI) (LC) (12) (R) (LC) (10) Jesús Loc Jesús Loc Jesús Loc Antonio Robles ARDEIDAE ARDEIDAE ARDEIDAE ARDEIDAE

89 Eudocimus albus 90 Plegadis chihi 91 Platalea ajaja 92 Coragyps atratus White Ibis White-faced Ibis Roseate Spoonbill Black Vulture Ibis Blanco Ibis Cara Blanca Espátula Rosada Zopilote Común (R, MI) (LC) (12) (R, MI) (LC) (8) (R, MI) (LC) (9) (R) (LC) (5) Jesús Loc Jonathan Vargas Antonio Robles Jesús Loc THRESKIORNITHIDAE THRESKIORNITHIDAE THRESKIORNITHIDAE CATHARTIDAE

93 Cathartes aura 94 Pandion haliaetus 95 Elanus leucurus 96 Rostrhamus sociabilis ♂ Turkey Vulture Osprey White-tailed Kite Snail Kite Zopilote Aura Gavilán Pescador Milano Cola Blanca Gavilán Caracolero (R) (LC) (5) (MI, R) (LC) (7) (R) (LC) (8) (R) (PR) (9) Jesús Loc Antonio Robles Antonio Robles Antonio Robles CATHARTIDAE PANDIONIDAE ACCIPITRIDAE ACCIPITRIDAE

10 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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97 Accipiter cooperii (J) 98 Geranospiza caerulescens 99 Buteogallus anthracinus 100 Buteogallus urubitinga Cooper's Hawk Crane Hawk Common Black Hawk Great Black Hawk Gavilán de Cooper Gavilán Zancón Aguililla Negra Menor Aguililla Negra Mayor (R, MI) (PR) (8) (R) (A) (11) (R) (PR) (11) (R) (PR) (12) Antonio Robles Antonio Robles Jesús Loc Antonio Robles ACCIPITRIDAE ACCIPITRIDAE ACCIPITRIDAE ACCIPITRIDAE

101 Parabuteo unicinctus 102 Buteo plagiatus 103 Buteo brachyurus 104 Buteo swainsoni Harris Hawk Gray Hawk Short-tailed Hawk Swainson's Hawk Aguililla Rojinegra Aguililla Gris Aguililla Cola Corta Aguililla de Swainson (R) (PR) (11) (R) (LC) (8) (R) (LC) (10) (MI) (PR) (12) Jesús Loc Jesús Loc Antonio Robles Antonio Robles ACCIPITRIDAE ACCIPITRIDAE ACCIPITRIDAE ACCIPITRIDAE

105 Buteo albonotatus 106 Buteo jamaicensis 107 Tyto alba 108 Glaucidium brasilianum Zone-tailed Hawk Red-tailed Hawk Barn Owl Ferruginous Pygmy-Owl Aguililla Aura Aguililla Cola Roja Lechuza de Campanario Tecolote Bajeño (MI, R) (PR) (10) (R, MI) (LC) (6) (R) (M) (LC) (9) (R) (LC) (8) Antonio Robles Antonio Robles Antonio Robles Antonio Robles ACCIPITRIDAE ACCIPITRIDAE TYTONIDAE STRIGIDAE

11 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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109 Athene cunicularia 110 citreolus 111 Chloroceryle americana ♀ 112 Melanerpes chrysogenys ♂ Burrowing Owl Citreoline Trogon Green Kingfisher Golden-cheeked Woodpecker Tecolote Llanero Trogon citrino Martín Pescador Verde Carpintero Enmascarado (R, MI) (LC) (12) (R) (LC) (E) (16) (R) (LC) (10) (R) (LC) (E) (12) Jonathan Vargas Antonio Robles Jonathan Vargas Antonio Robles STRIGIDAE TROGONIDAE ALCEDINIDAE PICIDAE

113 Melanerpes uropygialis ♂ 114 Picoides scalaris ♂ 115 Dryocopus lineatus 116 Campephilus guatemalensis ♂ Gila Woodpecker Ladder-backed Woodpecker Lineated Woodpecker Pale-billed Woodpecker Carpintero del Desierto Carpintero Mexicano Carpintero Lineado Carpintero Pico Plata (R) (LC) (10) (R) (LC) (9) (R) (LC) (9) (R) (PR) (14) Antonio Robles Antonio Robles Antonio Robles Jesús Loc PICIDAE PICIDAE PICIDAE PICIDAE

117 Herpetotheres cachinnans 118 Caracara cheriway 119 Falco sparverius ♀ 120 Falco columbarius Laughing Falcon Crested Caracara American Kestrel Merlin Halcón Guaco Caracara Quebrantahuesos Cernícalo Americano Halcón Esmerejón (R) (PR) (10) (R) (LC) (8) (MI, R) (LC) (11) (MI) (PR) (7) Antonio Robles Jesús Loc Jesús Loc Jonathan Vargas FALCONIDAE FALCONIDAE FALCONIDAE FALCONIDAE

12 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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121 Falco peregrinus 122 Eupsittula canicularis 123 Amazona albifrons 124 Xiphorhynchus flavigaster Peregrine Falcon Orange-fronted Parakeet White-fronted Parrot Ivory-billed Woodcreeper Halcón Peregrino Perico Frente Naranja Loro Frente Blanca Trepatroncos Bigotudo (MI, R) (PR) (9) (R) (PR) (14) (R) (PR) (12) (R) (LC) (13) Jonathan Vargas Jonathan Vargas Antonio Roble Antonio Robles FALCONIDAE PSITTACIDAE PSITTACIDAE FURNARIIDAE

125 Empidonax traillii 126 Empidonax minimus 127 Empidonax difficilis 128 Sayornis nigricans Willow Flycatcher Least Flycatcher Pacific-slope Flycatcher Black Phoebe Mosquero Saucero Mosquero Mínimo Mosquero Californiano Papamoscas Negro (MI) (LC) (10) (MI) (LC) (10) (MI, R) (LC) (SE) (12) (R, MI) (LC) (9) Jonathan Vargas Jonathan Vargas Antonio Robles Jesús Loc TYRANNIDAE TYRANNIDAE TYRANNIDAE TYRANNIDAE

129 Pyrocephalus rubinus 130 Myiarchus tuberculifer 131 Myiarchus cinerascens 132 Pitangus sulphuratus Vermilion Flycatcher Dusky-capped Flycatcher Ash-throated flycatcher Great Kiskadee Mosquero Cardenalito Papamoscas Triste Papamoscas Cenizo Luis Bienteveo (R, MI) (LC) (5) (R) (LC) (9) (MI, R) (LC) (9) (R) (LC) (5) Antonio Robles Antonio Robles Jonathan Vargas Antonio Robles TYRANNIDAE TYRANNIDAE TYRANNIDAE TYRANNIDAE

13 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

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133 Myiozetetes similis 134 Tyrannus melancholicus 135 Tyrannus crassirostris 136 Pachyramphus aglaiae ♂ Social Flycatcher Tropical Kingbird Thick-billed Kingbird Rose-throated Becard Luis Gregario Tirano Tropical Tirano Pico Grueso Mosquero Cabezón Degollado (R) (LC) (5) (R) (LC) (4) (R, MI) (LC) (SE) (12) (R) (LC) (11) Jesús Loc Jesús Loc Jesús Loc Antonio Robles TYRANNIDAE TYRANNIDAE TYRANNIDAE TITYRIDAE

137 Lanius ludovicianus 138 Calocitta colliei 139 beecheii (J) 140 sinaloae Loggerhead Shrike Black-throated - Purplish-backed Jay Crow Alcaudón Verdugo Urraca-hermosa Cara Negra Chara Sinaloense Cuervo Sinaloense (R, MI) (LC) (11) (R) (LC) (E) (13) (R) (P) (E) (17) (R) (LC) (E) (11) Antonio Robles Jesús Loc Jonathan Vargas Jesús Loc LANIIDAE CORVIDAE CORVIDAE

141 Tachycineta albilinea 142 sinaloa 143 Polioptila caerulea 144 Turdus rufopalliatus Mangrove Swallow Sinaloa Blue-gray Gnatcatcher Rufous-backed Robin Golondrina Manglera Chivirín Sinaloense Perlita Azulgris Mirlo Dorso Canela (R) (LC) (11) (R) (LC) (E) (13) (MI) (LC) (7) (R) (LC) (E) (10) Antonio Robles Antonio Robles Antonio Robles David Molina HIRUNDINIDAE TROGLODYTIDAE POLIOPTILIDAE TURDIDAE

14 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

[fieldguides.fieldmuseum.org] [921] version 1 8/2017

145 Mimus polyglottos 146 Oreothlypis celata 147 Oreothlypis ruficapilla 148 Setophaga pitiayumi Northern Mockingbird Orange-crowned Warbler Nashville Warbler Tropical Parula Centzontle Norteño Chipe Corona Anaranjada Chipe de Coronilla Parula Tropical (R, MI) (LC) (8) (MI) (LC) (9) (MI) (LC) (9) (R) (LC) (10) Jesús Loc Jonathan Vargas Antonio Robles Jonathan Vargas MIMIDAE PARULIDAE PARULIDAE PARULIDAE

149 Setophaga petechia 150 Setophaga coronata 151 Icteria virens 152 Volatinia jacarina Yellow Warbler Yellow-rumped Warbler Yellow-breasted Chat Blue-black Grassquit Chipe Amarillo Chipe Coronado Buscabreña Semillero Brincador (MI, R) (LC) (6) (MI) (LC) (6) (MI, R) (LC) (9) (R) (LC) (4) Jesús Loc Antonio Robles Antonio Robles Jonathan Vargas PARULIDAE PARULIDAE ICTERIIDAE THRAUPIDAE

153 Rhodinocichla rosea 154 Melozone kieneri 155 Peucaea ruficauda 156 Chondestes grammacus Rosy Thrush-Tanager Rusty-crowned Ground-Sparrow Stripe-headed Sparrow Lark Sparrow Tangara Cuitlacoche Rascador Nuca Rufa Zacatonero Corona Rayada Gorrión Arlequín (R) (LC) (17) (R) (LC) (E) (13) (R) (LC) (11) (MI) (LC) (10) Antonio Robles Antonio Robles Antonio Robles Jesús Loc RHODINOCICHLIDAE PASSERELLIDAE PASSERELLIDAE PASSERELLIDAE

15 NAYARIT, MÉXICO

COMMON BIRDS OF THE MARISMAS NACIONALES BIOSPHERE RESERVE

Jesús Alberto Loc-Barragán1, José Antonio Robles-Martínez2, Jonathan Vargas-Vega3 and David Molina4

1Fotógrafos de Naturaleza A.C., 2Universidad Autónoma de Nayarit, UAT, 3Terra Peninsular A.C. and 4Estación Ornitológica “Sierra de San Juan-La Noria, Nayarit

[fieldguides.fieldmuseum.org] [921] version 1 8/2017

157 Passerculus sandwichensis 158 Melospiza lincolnii 159 Quiscalus mexicanus ♀ 160 Molothrus aeneus Savannah Sparrow Lincoln's Sparrow Great-tailed Grackle Bronzed Cowbird Gorrión Sabanero Gorrión de Lincoln Zanate Mexicano Tordo Ojo Rojo (MI) (LC) (8) (MI) (LC) (7) (R) (LC) (5) (R) (LC) (6) Antonio Robles Antonio Robles Jonathan Vargas Jesús Loc PASSERELLIDAE ICTERIDAE ICTERIDAE ICTERIDAE

161 Molothrus ater ♂ 162 Icterus pustulatus ♀ 163 Icterus wagleri ♂ 164 Cassiculus melanicterus ♂ Brown-headed Blackbird Streak-backed Oriole Black-vented Oriole Yellow-winged Cacique Tordo Cabeza Café Bolsero Dorso Rayado Bolsero de Wagler Cacique Mexicano (MI) (LC) (7) (R) (LC) (10) (R) (LC) (12) (R) (CE) (LC) (10) Jonathan Vargas Jonathan Vargas Jonathan Vargas Antonio Robles ICTERIDAE ICTERIDAE ICTERIDAE ICTERIDAE

153 JESÚS LOC 154 ANTONIO ROBLES 155 JONATHAN VARGAS 156 DAVID MOLINA Biologist. Researcher for Biodiversa Student of Bachelor’s Degree in Biologist. Obtained his Master’s of Marine Biologist. He has worked in A.C., Nature Photographer for Tourism. Nature photographer and Science at Universidad Autónoma de various research and conservation Fotógrafos de Naturaleza A.C and specialized guide for bird watching. BCS. Nature photographer and bird projects at Marismas Environmental Educator of Grupo He currently collaborates in several specialized birding guide, certificate Nacionales. Currently coordinates Ecologista A.C. monitoring bird projects in Nayarit. of Nature and Culture Interpreter by environmental and conservation Author/co-author of 25 peer- the National Interpretation wildlife programs. Author and co- reviewed papers and has participated Association. Author/co-author of author of some peer-reviewed in national/international conferences. some peer-reviewed articles. He articles. He currently collaborates in several coordinates environmental wildlife projects in Nayarit. conservation and wildlife programs.