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Ent18 2 097 105 (Johnson).Pmd Russian Entomol. J. 18(2): 97105 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2009 Revised list of Old World types and names of Corticariinae (Coleoptera: Latridiidae) described by Mannerheim and Motschulsky, 18431867 Èñïðàâëåííûé ñïèñîê òèïîâ è íàçâàíèÿ Corticariinae (Coleoptera: Latridiidae), îïèñàííûõ Ìàííåðãåéìîì è Ìî÷óëüñêèì â 18431867 èç Ñòàðîãî Ñâåòà Colin Johnson Êîëèí Äæîíñîí 17 Peaknaze Close, Glossop, Derbys., SK13 6UN, England. E-mail: [email protected] KEY WORDS. Revised list, Latridiidae, Corticariinae, old world types and names, Mannerheim, Motschulsky, 18431867, lectotype designations, new status, new synonyms. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ. Ñïèñîê, òèïû è íàçâàíèÿ, Latridiidae, Corticariinae, Ñòàðûé Ñâåò, Ìàííåðãåéì, Ìî÷óëüñêèé, 18431867, îáîçíà÷åíèÿ ëåêòîòèïîâ, íîâûé ñòàòóñ, íîâûé ñèíîíèì. ABSTRACT. Types and names of all Mannerheim ÐÅÇÞÌÅ. Ïðèâåä¸í ñïèñîê òèïîâ è íàçâàíèé and Motschulsky old world (Eurasia, Africa, India and âèäîâîé ãðóïïû Corticariinae (Latridiidae), îïèñàí- Sri Lanka) species of Corticariinae (Latridiidae) from íûõ Ìàííåðãåéìîì è Ìî÷óëüñêèì â 18431867 ãã 1843 to 1867, are revised and listed, together with new èç Ñòàðîãî Ñâåòà (Åâðàçèÿ, Àôðèêà, Èíäèÿ è Øðè Lectotype designations and changes in nomenclature. Ëàíêà). Îáîçíà÷åíû ëåêòîòèïû Coricaria fusco- Lectotype of Coricaria fuscotestacea Motschulsky, testacea Motschulsky, 1861, C. axillaris Mannerheim, 1861, C. axillaris Mannerheim, 1844, C. badia Man- 1844, C. badia Mannerheim, 1844, C. baicalica nerheim, 1844, C. baicalica Mannerheim, 1844, C. Mannerheim, 1844, C. boreophila Motschulsky, 1867, boreophila Motschulsky, 1867, C. convexipennis Mots- C. convexipennis Motschulsky, 1861, C. crenicollis chulsky, 1861, C. crenicollis Mannerheim, 1844, C. Mannerheim, 1844, C. cylindrica Mannerheim, 1844, cylindrica Mannerheim, 1844, C. flavifrons Motschul- C. flavifrons Motschulsky, 1867, C. gracilis sky, 1867, C. gracilis Mannerheim, 1844, C. laticollis Mannerheim, 1844, C. laticollis Mannerheim, 1844, C. Mannerheim, 1844, C. melanophthalma Mannerheim, melanophthalma Mannerheim, 1844, C. parvula 1844, C. parvula Mannerheim, 1844, C. pharaonis Mannerheim, 1844, C. pharaonis Motschulsky, 1867, Motschulsky, 1867, C. piligera Mannerheim, 1844, C. C. piligera Mannerheim, 1844, C. psammeticha psammeticha Motschulsky, 1867, C. punctatissima Motschulsky, 1867, C. punctatissima Motschulsky, Motschulsky, 1867, C. rubripes Mannerheim, 1844, C. 1867, C. rubripes Mannerheim, 1844, C. setosa setosa Motschulsky, 1867, C. subrugosa Motschulsky, Motschulsky, 1867, C. subrugosa Motschulsky, 1867, 1867, C. subtilis Mannerheim, 1844, C. transversicollis C. subtilis Mannerheim, 1844, C. transversicollis Motschulsky, 1867, C. tincta Mannerheim, 1844, C. Motschulsky, 1867, C. tincta Mannerheim, 1844, C. truncatella Mannerheim, 1844, C. unicarinulata Mots- truncatella Mannerheim, 1844, C. unicarinulata chulsky, 1867, C. validipes Motschulsky, 1867, Latrid- Motschulsky, 1867, C. validipes Motschulsky, 1867, ius formicetorum Mannerheim, 1843, Melanophthalma Latridius formicetorum Mannerheim, 1843, angulicollis Motschulsky, 1866, M. cylindricollis Mots- Melanophthalma angulicollis Motschulsky, 1866, M. chulsky, 1866, M. gibbula Motschulsky, 1866, M. ret- cylindricollis Motschulsky, 1866, M. gibbula roculis Motschulsky, 1866 and Mineauxia villigera Motschulsky, 1866, M. retroculis Motschulsky, 1866 è Motschulsky, 1867 designated. Cortinicara corpulenta Mineauxia villigera Motschulsky, 1867. Cortinicara (Motschulsky, 1866), stat.n. from Sri Lanka and Mel- corpulenta (Motschulsky, 1866), stat.n èç Øðè Ëàíêè è anophthalma (Cortilena) picina Motschulsky, 1866, Melanophthalma (Cortilena) picina Motschulsky, 1866, stat.n. from South Africa., given as valid species; Mel- stat.n. èç Þæíîé Àôðèêè, ïðèâîäÿòñÿ êàê âàëèäíûå anophthalma capicola Belon, 1898 is syn.n. of M. (C.) âèäû; Melanophthalma capicola Belon, 1898, syn.n. picina Motschulsky, 1866. êàê ñèíîíèì M. (C.) picina Motschulsky, 1866. 98 Colin Johnson Introduction List of Species and Types Count Carl Gustaf von Mannerheim (17971854) Species are listed in alphabetical order for ease of was the first coleopterist to attempt a monograph of the reference and given a chronological number [in square small beetles of the genera Corticaria Marsham, 1802 brackets] to demonstrate priority of names for nomen- and Lathridius Illiger, 1801 (now Latridius Herbst, clatural purposes. Type localities are given in full in 1793) which together made up the family Latridiidae their original published form, which is in Latin for [Mannerheim, 1844], an interest which seems to have Mannerheim, French for Motschulsky, together with been awakened by his first discovery of a beetle in this their modern equivalents. International borders changed family [Mannerheim, 1843]. He published additional after World War 2, especially those between Finland papers on Alaskan beetles and also collaborated with and Russia, so it is essential to note that Villnäs (= other coleopterists of his day, especially Colonel Vic- Villnas), Mannerheims ancestral home, still lies in tor Ivanovitsch de Motschulsky (18101871) who was Finland about 25 km NW of Turku / Åbo. Vyborg (= also developing an interest in similar groups of beetles Viborg, Viipuri) is now the name of a north-west from his own studies and travels in both the old and Russian city and province, Vozrozhdenie (= Kavan- new world [Motschulsky, 1849, 1861, 1866, 1867]. tholm) being Mannerheims manor from later years Consequently, many species of Latridiidae were de- and 20 km NNE from Vyborg, both in the Karelian scribed by these two great workers, so that their collec- Isthmus. I have assumed that when there is only a tions, rich in type specimens, in the University Zoolog- single specimen in a collection, it represents the holo- ical Museums of Helsinki (Finland) and Moscow (Rus- type, in cases where the original descriptions lack sia) respectively, are of great importance in enabling variation in measurements and localities. Multiple us to interpret the many species which they described type specimens are therefore syntypes from which I within this difficult family. The advent of new work have selected Lectotypes, some of which have already which pioneered use of the male genitalia of Latridi- been validated in earlier papers [Johnson 1972, 1986], idae as an important and hitherto overlooked character but the bulk of them (15 Mannerheim names, 17 of [Strand, 1937], laid the basis for modern studies where- Motschulsky) are designated below. Label limits are by such characters provide the all important features indicated by diagonal lines / /. The location of speci- that enable us to identify species with much greater mens is indicated by the symbols (ZMH) for Helsinki accuracy and confidence than our predecessors. Since and (ZMM) for Moscow. most of the Mannerheim and Motschulsky names and Names lacking types have some additional data sum- types had never been subjected to modern treatment, I marized under Other material. Only three species are resolved, nearly forty years ago, to start a revision of without type material: Corticaria illaesa Mannerheim, all the Old World Corticariinae names published by 1844, C. subacuminata Mannerheim, 1844 and C. umbi- these two great coleopterists. The Mannerheim collec- licifera Mannerheim 1844. In the interests of nomencla- tion in Helsinki was much easier and quicker for me to tural stability, interpretation of these names follows tradi- study with postal loans from the museum than Mots- tional usage or synonymy, which is based on recent re- chulskys in Moscow, because of differences in geo- descriptions [Rücker, 1983], or correctly identified Oth- graphical position and political governance. I attempt- er material in the describers collections. ed to study all such species in both collections, and to Three new changes in nomenclature are: Cortinica- carefully clean (many Motschulsky specimens were ra corpulenta (Motschulsky, 1866) from Sri Lanka, originally hidden by mould), dissect, remount and stat.n.; Melanophthalma (Cortilena) picina Motschul- unambiguously label (although some specific names sky, 1866 from South Africa, stat.n.; M. capicola Be- are now slightly outdated) where appropriate, all type lon, 1898 is syn.n. of M. (C.) picina Motschulsky, 1866. material. As a result of my studies, many erroneous abietorum Motschulsky, 1867: 71 interpretations of catalogued species [Hetschko, 1926; (Corticaria) [71] Winkler, 1926] were found together with many new synonyms, all of which are given in the new Palaearc- TYPE LOCALITY: St. Pétersbourg = St. Peters- tic Catalogue [Johnson, 2007]. Recent revisional work burg, Russia. on the difficult Melanophthalma taurica species-group TYPE MATERIAL: Holotype #: /type/, /15 Aug.59 fichtengr/, /Corticaria abietorum Motsch. Petropol/ (ZMM). [Rücker & Johnson, 2007] made use of some of these IDENTITY: synonym of Corticaria longicornis (Herbst, new interpretations. Mannerheim described 39 species 1784) [Johnson, 2007]. from Europe, Asia Minor, Caucasus, Middle Asia and Siberia. The bulk of Motschulskys 41 species were adustipennis Motschulsky: 1867: 57 described from Eurasia and North Africa (27), with (Corticaria) [62] others from South Africa (5), India and Sri Lanka (9). TYPE LOCALITY: Cap de bonne Espérance = Cape of He also described two genera, the well-known and Good Hope, South Africa. ubiquitous Melanophthalma [Motschulsky, 1866] and TYPE MATERIAL: Holotype $: /Type/, /Corticaria Cortilena [Motschulsky, 1867], now considered a
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