THE JEPSON GLOBE a Newsletter from the Friends of the Jepson Herbarium

THE JEPSON GLOBE A Newsletter from the Friends of The Jepson Herbarium

VOLUME 15 NUMBER 1 June 2004

Director’s Column: for possible updates to TJM, conducting Graduate Student Research research to evaluate the reports, revis- by Brent D. Mishler ing distribution or other descriptive information in TJM accounts, updating the on-line accounts, and reviewing One of the most important roles photographs of California plants. In of a university-associated herbarium is addition to his important roles in the to foster the research of graduate stu- Jepson Herbarium, Jeff is also leading dents. The University and Jepson Her- the effort to accession the Baja Califor- baria have developed an increasingly nia collections made by Annetta Carter broad and interactive group of graduate into the University Herbarium. students over the last decade. They span an incredible array of topics and ap- Staci Markos proaches; we are very proud of them all. Project Manag. & Development Several Ph.D. students have graduated Coord. recently and are off to promising starts Jepson Flora Project Staff In 1995, Staci came to UC in their careers (including Raymund by Bruce G. Baldwin Berkeley as a graduate student working Chan, John McMurray, Staci Markos, Curator, Jepson Herbarium in the Herbarium and in the Department Terry O’Brien, Patricia Sanchez, Lisa Convening Editor, Jepson Flora Project of Integrative Biology. She studied Shultheis, and Dennis Wall). I’ll let the the evolutionary patterns in Lessingia current group of students tell you about As many of you know, the (Compositae). She completed her de- themselves in their own words: production of a Second Edition of The gree in December 2000 but not before Jepson Manual (TJM2) is underway. she had accepted the position of Coor- Kirsten Fisher Continued success of the project will dinator of Public Programs (February, Because they have little eco- depend in part on authors who will be 1999). Staci coordinated the program for nomic value to humans, and are there- contributing revised taxonomic treat- five years. Last fall, we hired a new Ed- fore unlikely subjects for anthropogenic ments, the Jepson Flora Project Editors, ucation Coordinator (Cynthia Perrine) dispersal, mosses provide an excellent and the hard-working and dedicated and Staci became more involved in the system for investigating natural dis- Jepson Flora Project staff. The staff is Jepson Flora Project (although you may persal patterns and diversification a great group of people and I hope this still see her on a workshop or two). processes across oceanic islands. My article will serve as a brief introduction dissertation research has focused on to each of them. Richard Moe the systematics and biogeography of a Database Administer & Webmaster small clade of mosses, Leucophanella, Jeff Greenhouse Dick first visited the Herbarium that occur throughout the Paleotropics. Collections Research Specialist in 1973 to examine the marine algae Until recently, this group of mosses was After years of volunteering in collections on his way to Antarctica, recognized as a single polymorphic spe- the Herbarium, Jeff officially joined where he spent 15 consecutive months. cies, Syrrhopodon involutus; however, the Herbarium staff in 2001. Jeff is After his big southern adventure, Dick phylogenetic analyses of morphological responsible for gathering initial reports returned to UC Berkeley to pursue

Above: Richard Moe, Margriet Wetherwax, Continued on page 2. Continued on page 7. Staci Markos, Jeff Greenhouse, Bruce Bald- win, and Tom Rosatti (left to right) Jepson Horticultural Database

The use of native plants in restoration efforts and home gardens is an important conservation strategy. Up to now, the Jepson Herbarium has contributed to this effort through the publication of horticultural information in The Jepson Manual. To increase our support of native plant horticulture, the Herbarium has developed a horticultural database that provides new and innovative ways to access the large amount of morphological and horticultural data that already exists for ~3000 taxa in The Jepson Manual.

Native plant enthusiasts are now able to make personalized queries and create customized lists of plants that may grow well and suit particular needs in their gardens. The database contains information on plant name, climatic zone, environmental conditions, life-form, flower color, and many more characters. Possible database queries include: What plants will grow well in a particular climate zone? What plants will grow well in a particular climate zone if they have moderate summer water? What plants will grow well in my shady back yard and sunny front yard? Do cultivars exist for a particular species?

You may access the horticultural database through the Online Interchange (http://ucjeps.berkeley.edu/interchange. html) or directly at: http://ucjeps.berkeley.edu/interchange/hort_form.html

The Jepson Herbarium gratefully acknowledges the Elvenia J. Slosson Endowment Fund for support of this project.

Graduate Students, continued from page 1. and molecular data indicate the pres- tern of reduced fertility and increased since the early 1800’s. The “memecy- ence of several morphologically and asexual reproduction is one component loids” have large seeds and also very geographically distinct lineages within of the paradigmatic “island syndrome” hard wood – derived characteristics S. involutus. I have prepared both a proposed by Sherwin Carlquist in 1974. conferring a specialization for shaded, traditional Linnean taxonomic treat- forest-understory habitats. Also unusu- ment and a rank-free treatment of the Doug Stone al are the anthers, each of which bears Leucophanella clade; both recognize a peculiar dorsal gland on the connec- five taxa:Involutus , Revolutus, Banksii, tive tissue. It has been suggested that Rufescens, and Borneensis. these are oil-producing glands, and The Leucophanella group most if this is true then the flowers should likely originated in Southeast Asia/In- be pollinated by oil-collecting bees donesia and subsequently dispersed to (a common syndrome in the tropics). increasingly remote islands in both the Given the wide geographic distribution Indian and South Pacific oceans. The and nearly 500 recognized species in occurrence of Leucophanella lineages the Memecyloideae, the fundamental on continents, near-continental islands, challenge for my study has been one of and remote oceanic islands makes it sampling. While other botanists have possible to investigate the evolution generously sent samples to me from Memecylon amshoffiae of reproductive traits in island mosses. tropical America and Indo-Malesia, I Like many other groups of plants, the have personally undertaken two suc- Leucophanella group displays reduced The focus of my dissertation cessful collecting expeditions – first in fertility (decreased production of research is the systematics and evo- the forests of Madagascar and eastern sporophytes and female gametangia) lution of the large and widespread Africa and more recently in western on remote oceanic islands. Addition- tropical family Melastomataceae. In Africa (Cameroon). A substantial ally, there appears to be an increased particular, I have been studying the portion of my research has involved reliance on asexual reproduction (an morphologically anomalous subfamily phylogenetic reconstruction using increase in the production of asexual Memecyloideae, a group of shrubs and evidence from morphology, anatomy, propagules known as gemmae) in the small trees of which the systematic and DNA sequence data. In addition, remote island moss lineages. This pat- position has been a matter of debate through my field work and herbarium 2 Continued on page 3. Graduate Students, continued from page 2. studies I have found many species that nesian archipelagoes. This may imply Elizabeth Zacharias are completely new to science. While that there was selection for sterile plants my interests will always lie more with that possessed different morphological studying plants than developing policy, traits. In conclusion, my research on ti I hope that my training and experience plants will help to better understand will also help me contribute to future events in Polynesian prehistory and the tropical conservation efforts. cultural role that ti may have played, particularly with its use as a source of Anya Hinkle food.

Ruth Kirkpatrick Elizabeth examining Atriplex I am investigating evolutionary relationships, historical biogeography, life-form evolution, and ecophysiological change in the North American lineages of Atriplex (Chenopodiaceae/ Amaranthaceae), with an emphasis on Anya in the greenhouse Californian diversity. I am conducting molecular and morphological phylo- The plant I am working on genetic analyses to determine whether is Cordyline fruticosa. Its common Atriplex, as traditionally recognized, name is “ti.” It is a woody monocot Pellaea truncata and the North American taxa are each and is closely related to species in monophyletic groups and whether the the family Agavaceae. It was intro- I am interested in evolutionary current circumscription of the genus duced throughout Polynesia and is an processes, history, and phylogenetic needs revision. Many of the annual important cultural plant in the Pacif- relationships among green plants, es- Californian taxa are found in extreme ic. Uses for the plant include using pecially ferns. For my dissertation I environments and are highly limited in the leaves for costumes, decoration, plan to apply molecular, cytogenetic, geographic and ecological distribution, wrapping food, and for medicines for morphological, and eco-physiological thus making them interesting from an fevers, rashes, swellings, sprains, etc. data to help resolve the phylogeny of ecophysiological standpoint. Many (basically topically for heat related the fern genus Pellaea. I plan to utilize taxa grow in isolated salt or alkaline disorders), also as a living fence, and this information to elucidate the adap- scalds, and several have been classified for religious purposes. Its rhizome was tive evolutionary history of several Pel- as rare or endangered. Atriplex plants used as a source of sugar, particularly laea taxa to their xeric environments. I are morphologically cryptic in addition in past centuries. My research uses this have more than 100 greenhouse main- to being able to tolerate very harsh con- plant as a proxy for understanding the tained pellaeas and close relatives that ditions, studying their ecophysiology colonization of this vast area of Earth, I collected from throughout California may determine adaptations that allow the Pacific Ocean. Multiple lines of and the southwestern U.S. I plan to them such tolerance. I am especially evidence suggest a general west-to-east perform physiological, morphologi- interested in the evolutionary response pattern of colonization, with Hawaii cal, and cytogenetic analyses on these to aridity and salinity. My research has and New Zealand being some of the plants. I have performed preliminary taken me all around the Central Valley, last places to have been inhabited by desiccation tolerance experiments and Owen’s Valley, and the Mojave Desert humans. I am using patterns of genetic phylogenetic analyses on many of these in California, as well as Utah, Nevada, variation in ti plants from Polynesia individuals. I am currently sequencing and Arizona to collect Atriplex. My to try to better understand the order of several genes (rps4, trnS-spacer, and research has significance for under- Polynesian colonization events. Addi- ITS) of Pellaea and outgroup taxa in standing the origin of arid systems in tionally, I am looking at reproductive order to generate a robust phylogenetic angiosperm diversification. With sali- and morphological differences in ti hypothesis. nization and drought increasing in many from different parts of Polynesia. Most areas of the world, studying questions Eastern Polynesian ti plants have sterile about salt- and drought-tolerance using pollen and appear not to set any seed, an evolutionary framework is very im- in contrast to ti plants in Western Poly- portant. Continued on page 4. 3 Graduate Students, continued from page 3 Vicente Garcia of research that has been called “eth- tree crop, Aleurites moluccana Willd. My project focuses on evo- nobryology.” My research is focused (Euphorbiaceae) to study the human lutionary and ethnobotanical aspects on the moss genus Plagiomnium, one colonization of the Pacific Islands. of two plant genera in the Piperaceae, of the few mosses that has specific Within the last 5,000 years, Aleurites Piper and Macropiper. The genus human uses. This genus is interesting moluccana (Candlenut tree, bancoulier, Piper, which Macropiper may be because it has reported medicinal uses kukui, tutui, tia’iri, kemiri) was spread nested within, has over 1,000 species by the Oweekeno and Bella Coola from Southeast Asia or Near Oceania distributed in tropical areas around the people of the central coast of British through the Pacific Islands by the first world. I’m interested in elucidating Columbia and by herbalists in China. human colonists. Because of the nature patterns of colonization and subsequent Also, some species of Plagiomnium are of this relationship in which humans diversification of the species within the cultivated in areas of southwest China have served as the dispersal agent for South Pacific. The 40+ species ofPiper to encourage the growth of gall-aphids the Candlenut tree, I have proposed and Macropiper in this region appear that over-winter in Plagiomnium. The that the geographic distribution patterns to form two groups with different his- aphids are critical in the production of of the genetic variants within the spe- tories. The Piper vines probably colo- economically valuable gallnuts on su- cies, as well as the cultural uses of the nized the South Pacific via Southeast mac trees (Rhus spp.). I am currently tree, will reflect the human settlement Asia where their closest relatives are working on reconstructing the evo- route through Oceania. This research found, whereas the shrubby Macropip- lutionary history of Plagiomnium in provides me with ample opportunities er assemblage could be more closely order to understand how a group of for fieldwork in physically and spiritu- related to plants in Central America and potentially medicinally important ally challenging localities like Hawaii, South America. More work needs to be chemicals, flavonoids, have evolved Tahiti, Samoa, Indonesia, and many conducted to refine these biogeographic in this group. In addition, I will other blue-skied, beach-ridden islands hypotheses. Kava, Piper methysticum, soon be conducting ethnobotanical in between. holds an important cultural role in the fieldwork in Yunnan, China, to in- South Pacific. It is used ethnomedically vestigate the uses of Plagiomnium Danica Harbaugh to treat many ailments, it’s imbibed so- and other mosses in the area. cially to create and preserve communal bounds, and it’s an important drink in Anna Larsen many sacred ceremonies. This plant surely accompanied the Polynesians during their colonization of the Pacific Islands. By clarifying phylogenetic relationships among kava cultivars, and conducting interviews to document the Santalum freycinetianum ethnobotanical uses of related species, I can test the patterns of Polynesian I am studying the systematics movement across the Pacific. and biogeography of sandalwoods (Santalum spp.), a genus of plants that Eric Harris includes the sandalwoods. This is one of the most heavily exploited groups of Anna in the field plants across its range from Indomalay- sia to Hawaii because it has a diverse Through my graduate stud- array of uses and is commonly known ies, I am dabbling in diverse areas for its use as a scent in perfume and of study including: the processes of incense. The demand for its valuable plant evolution, the ethnobotany of oil has lead to drastic over-harvesting the Pacific Islands and Indonesia, the by people, in almost all regions where co-movement of plants and people, and it grows. It is an interesting group to the genetic and socio-cultural meth- study because all species of Santalum Plagiomnium insigne sporophytes ods used to tackle questions in these are hemi-parasitic on the roots of oth- fields. In my dissertation research, er plants. Parasitic plants represent My dissertation research is concerned I am using genetic and cultural data interesting models to study molecular with the ethnobotany of moss, an area from an ethnobotanically-important Continued on page 6. 4 Plant Identification Services The Jepson Herbarium Projects & Resources Provided by The Jepson Herbarium The Jepson Flora Project The Jepson Herbarium now offers a plant identification service. Second Edition of The Jepson Manual Online Interchange for Advances in Plant Identifications include determination of plant name, statement about California Florisitcs Jepson Desert Manual toxicity and weediness, geographic distribution, and if needed, library re- Online Horticultural Database search. Electronic Publication of Jepson’s A Flora of California Fees are determined on an hourly basis, with a 1-hour minimum charge of $75. After the first hour, the fee will be applied in 20-minute intervals. Publications & Research Projects Constancea: University of California electronic publications in botany Send specimens by mail: Tarweeds & Silverswords: Evolution of the Madiinae 1) Wrap the specimen in newspaper and place it between two pieces of DeCew’s Guide to the Seaweeds cardboard. Flora of Mount Diablo Unravelling the dynamics of mating-system 2) Include information on where the specimen was collected including habi- evolution in tribe Collinsieae Building the Tree of Life -- A National tat information, state, county, nearest town, and approximate elevation. Resource for Phyloinformatics and Computational Phylogenetics 3) Include the following contact information: name, address, telephone Deep Green Plant Phylogenetics: Novel number, and email address. Analytical Methods for Scaling Data from Genomics to Morphology 4) Please indicate if an official letter should be provided with determinations Beyond “Deep Green”: Towards an Integra- tion of Plant Phylogenetics and Plant Genomics 5) Mail the specimen to: Margriet Wetherwax, Jepson Herbarium, 1001 Demography and Germination Ecology of VLSB #2465, Berkeley, CA 94720-2465. the Endangered Santa Cruz Tarplant Sierra Nevada Plants Project Upon arrival at the Jepson Herbarium, dried specimens will be frozen for seven days before they are examined. Educational Services & Resources Botanical Workshops & Courses Plant Identification Once the plant specimens have been identified, you will be contacted.The 1,920,000+ Worldwide Plant Specimens specimens, a list of determinations, and an invoice will be mailed to you. Photographic Slide Collection Payment will be due within 30 days of receipt of the invoice. Map Collection & Locality File Botanical Library (non-circulating) Use our convenient drop-in service: Administration Trustees: Vice Chancellor Emeritus Rod- 1) Please call and schedule an appointment with Margriet Wetherwax (510) eric Park, Chairman; Vice Chancellor Beth 643-7008 or send an email to [email protected]. Depending Burnside (on leave); UC Botanical Garden on the number of specimens, you may be able to wait for identifications. Director, Paul Licht; Professors John Taylor and Brent Mishler (ex officio) Director: Professor Brent Mishler 2) When you come to the Herbarium, please bring the specimens and all of Curator: Assoc. Professor Bruce Baldwin the information requested above. Database Admin. & Webmaster: Richard Moe Scientific Editor: Tom Rosatti Managing Editor: Margriet Wetherwax Collection Research Sp.: Jeff Greenhouse Research Associate: Bridget Wessa Administrative Curator: Barbara Ertter For more information, please contact Margriet Wetherwax (510) 643- Senior Museum Scientist: Fosiee Tahbaz 7008 or [email protected] Senior Museum Preparator: Ana Penny Assistant Museum Scientist: Kim Kersh Public Programs & Development: Staci Markos Cynthia Perrine

5 Graduate Students, continued from page 4. evolution because their evolutionary in the Marattiaceae has been termed a “liv- the Specimen Management System history may be linked to that of the host ing fossil” lineage in that the morphology for California Herbaria (SMASCH). species, or genes may be transferred has changed insignificantly over extensive The database now contains over from the host plant to the parasite. Cur- periods of time in the fossil record and the 365,000 specimen records and rently, I am building a phylogeny of the rate of molecular evolution of the genes serves countless numbers of people entire genus of about 25 taxa, using the thus far examined has been shown to be outside of the Herbaria who access nuclear rDNA gene ITS, and am pur- significantly slower in the Marattiaceae SMASCH over the web. In 2000, suing the use of additional nuclear and than in any closely related lineage of Tom began developing the Index to chloroplast genes. I have already done plants (with the possible exception of the California Plant Names, the core of extensive collecting in Hawaii and plan tree ferns of the Cyatheaceae and Dick- the Online Interchange for California to collect in Australia, the South Pacif- soniaceae). I hope, in the course of my Floristics. As Scientific Editor for ic, and India. With my resulting phylog- research, to reveal more about the nature TJM2, Tom’s roles include review- eny, I hope to address several questions, of living fossils and to determine whether ing and editing taxonomic treatments such as where and when the genus the diversity in the group as we see it today for scientific content, preparing diverged, the biogeographic patterns is a result of relatively recent speciation treatments for groups not assigned across the Pacific, host-specificity, and (i.e., going against the standard dogma to external authors, and assisting au- evolution of polyploidy, among others. surrounding living fossils) or a result or thors in the preparation of electronic In addition, I will use the phylogeny to more ancient divergence. I also hope to keys. make any appropriate changes to the use the phylogeny of the Marattiaceae to classification of the group, and identify shed light on questions concerning func- Margriet Wetherwax populations for conservation. tional arguments surrounding some novel Managing Editor morphological features found in the family Margriet officially joined Andy Murdock (incorporating evidence from both fossil the Herbarium staff in 1990 and was and living material), and to investigate the immediately involved in The Jepson historical biogeography of the family. Manual Project. She wrote 14 Scro- phulariaceae treatments and worked with the production team. From JFP Staff continued from page 1. 1994–1998, she managed correc- tions to TJM, oversaw the Jepson Li- graduate studies in phycology, including brary, and assisted me with growing the description of 4 new genera and 6 tarweeds for my research. Margriet’s new species of Antarctic seaweed. After attention then turned to The Jepson completing his degree, he stayed on as a Desert Manual. As Managing Editor, researcher and worked with Paul Silva on she edited treatments, corresponded the Catalogue of the Benthic Marine Algae with authors and the Editorial Board of the Indian Ocean. Dick’s career took Angiopteris frond uncurling about taxonomic treatments, worked a different turn when he started learning with illustrators, and interacted with more about electronic cataloguing; he is the production staff at UC Press. My research is focusing on now administrator for the Herbaria’s on- Margriet is also the Managing Editor the systematics of the family Marat- line database (SMASCH), webmaster, and for TJM2 and will have many of the tiaceae, an early evolutionary branch technical developer of the Jepson Online same responsibilities she had during of the ferns. The Marattiaceae has an Interchange and Jepson Horticultural Da- the Desert Manual project. extensive fossil record, and members tabase. In future issues of the Globe, of this group were dominant trees in the Jepson Flora Project staff will Carboniferous coal swamps. The large Tom Rosatti provide updates and short articles tree ferns of this group are long extinct; Scientific Editor discussing taxonomic changes and the extant members of this family, In 1987, Tom moved west from other revisions and additions that while still quite large (with fronds to 7 Michigan to accept the position of Scientif- will be recognized in the Second meters in length in the genus Angiop- ic Editor of The Jepson Manual. Tom also Edition of TJM. We look forward teris), never reach the height nor the contributed many taxonomic treatments to sharing the evolving understand- ecological dominance of the ancestral that had been orphaned late in the project. ing of California’s flora with our tree ferns. This extant lineage of ferns After the Manual was published in 1993, Friends. Stay tuned! Tom became the Project Coordinator for

6 Categories of Giving Friends of the Jepson Herbarium Gifts to support the Second Edition Name(s) ______$25,000 Honor the contributions and founding principles Address ______of W. L. Jepson, former City, State Zip ______Jepson Trustees Lincoln Constance and Robert Or- Telephone / email ______nduff, and former Jepson Curators Rimo Bacigalupi and Lawrence R. Heckard I would like to join the Friends / renew my membership (contribution to the annual fund). $10,000 Support taxonomic efforts in an organizing I would like to support the Second Edition of The Jepson Manual unit of the Manual: with my gift of ______. Ferns, Gymnosperms, Dicots, or Monocots Enclosed is _____ of a total pledge of _____ to be paid over ____ years. $5,000 Support floristic effort for

Please acknowledge me as a sponsor of ______a particular bioregion (indicate genus name e.g., Cypripedium, family name, e.g., Asteraceae, or (Twenty-four listed in other category) by printing my name in The Jepson Manual the Manual) (for gifts of $1,000 or more, see side bar). $2,500 Support taxonomic work in a particular family. Please acknowledge my gift as anonymous. See the Herbarium web site for an up-to-date, My or my spouse’s employer will match this gift. (Please enclose company form) complete list $1,000 Show enthusiasm for This gift is ___ in honor of ___ in memory of ______your favorite genus Please make your check payable to the Friends of the Jepson Herbarium (pledge $200 / 5 years) or charge your gift. Annual Support ___ Visa ___ Mastercard $500 Contribute to the illustra- tion of a new species Account # ______$250 Help accession specimens Exp. Date ______from the backlog Signature ______$100 Support taxonomic research at the species level MAIL TO: $35/$50 Basic membership in Friends The Jepson Herbarium of the Jepson Herbarium 1001 VLSB #2465 University of California Sponsorship opportunities Berkeley, CA 94720 are exclusive and will be available on a first-come, first-served basis. With The information you provide will be used for University business and will not be released unless required by law. A portion of all gifts is used to defray the costs of administering the funds. approval from the donor, gifts at the All gifts are tax deductible as prescribed by law. $1,000 level and above will be ac- knowledged in the front pages of The Jepson Manual. Gifts may be made as one-time payments or as a pledge, payable over 5 years.

7 The Jepson Herbarium supporters and Friends Although housed on the UC Berkeley campus, the Jepson Herbarium is largely supported by grants and pri- vate donations. The Herbarium gratefully acknowledges the following individuals for their generous support of the Annual Fund, the Jepson Flora Project, and the Second Edition of The Jepson Manual.

$25,000 Penstemon Paul Doty Roderic & Cathy Park Karen Markos Daniel Drake Nemophila Ecological Concerns Inc. $5,000 (gifts or pledges) Betsy Ringrose Virginia Fletcher Elizabeth Crispin Astragalus Leslie D. Gottlieb Georgie Robinett Steve Schoenig Helix Environmental* In memory of Jim Robinett Mimulus Susan H. Hossfeld Jake Sigg Willard Huyck $2,500 Eriogonum Richard Ibarra* Family Sponsor J. Giles Waines William W. Lester III Anonymous James G. Little & Linda K. Persson Compositae $500 Michael D. Luther Paul Licht Arthur Morley $1,500 Helen & J. Fraser Muirhead Elaine Plaisance & Jim Duncan Roderic & Cathy Park Desi & Karen Zamudio Donald Potter Roger & Marian Reeve $1,000 (gifts or pledges) $101 - $499 In memory of Wayne Roderick Robert J. Berman Flo Anderson Bruce & Alice Ronald Beth Burnside Stephen Buckhout & Stella Yang Irma Jo Salo Toni Corelli Gerald & Buff Corsi In memory of Margaret Williams Patrick Creehan Claire Englander Jessie Schilling In memory of Yvonne Doreen Creehan Norman E. Graham Alan & Joan Smith Katherine Cuneo Jeffrey & Judy Greenhouse Robert & Jean Soost Frank W. Ellis Virginia W. Havens John Sugrue Jane Hicks Lawrence Janeway Thomas Reid Associates, Inc.* In memory of Rob Schonholtz Dwight L. Johnson John M. Tucker Natalie Hopkins Neal Kramer Katherine F. Mawdsley & William F. Beth Lowe Corbin Up to $99 McCoy J. W. McKibben Mario Abreu In memory of Patrick Elvander L. Maynard Moe Lowell Ahart S. B. Meyer John W. Reynolds Verna Arnest In memory of Robert Ornduff Felice Pope Linda M. Barker In memory of Robert L. Pope Peter Slattery Heath Bartosh James Payne Smith Jr. Robert Stephens & Julie Packard Irene G. Stewart Camille Barr* Genus sponsors Bob Battagin Chris Davidson $100 Susan Bazell Paeonia J. K. Anderson Rolf W. Benseler Frank W. Elllis In memory of Esperance Jim & Alma Benson* Fritillaria Rebecca & Calvin Anderson-Jones Rosetta B. Bentley John Game David & Ann Bauer Roland Bergthold Erythronium Kathleen A. Becker F. Thomas Biglione Verne Garcia Berkeley Garden Club Edith Black* Arctostaphylos June Bilisoly Charles E. Blair Adeline L. Black Robert Garner Larry Blakely* (plus a matching gift from IBM) Richard & Trisha Burgess William Brace Pinus Coeta Chambers Anne Bradley Jeffrey & Judy Greenhouse Sarah Chaney & Dan Richards Cypripedium Ronald L. Clendenen* Thomas E. Bradner Jim Loughlin Sally Davis* Leslie Burkhart* 8 Eva Buxton Noel & Patricia Holmgren David Popp Brad Carter* William A. Howell Ruth Porter Barbara Castro* Daisie Huang* Jennifer Potts* Stephanie Changaris & Bruce Bonar Layne Huiet Thomas B. Powers & Noreen Stimac Sally Chess Virginia Hunt Robert Preston Oscar F. 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Schoolcraft Jan Dobak Ann Lambrecht In memory of Vern Oswald Heidi Dobson Michele Lee Howard Schorn Kim Earll Wilde Legard Michael Shephard* Margery Edgren Barbara Leitner Dennis Sheridan Phyllis Ellman Evelyne & David Lennette* Dave Silverman & Cindy Hopkins Oscar Enstrom Edmond Leong Magen Solomon Robert Fischer Aaron Liston Veva Stansell Diana Fish R. John Little Mary E. Stanton Margaret Flesher Larry Loeher & Grace Murayama* Peter Stekel W. J. Frank William Lupfer* In memory of May Stekel Sharon Franklet* Stephanie MacDonald* Laura Stockton Greg Gallaugher* Marta Mackenzie Ronald M. Stork Peter Garcia Lillian & Alex Maksymowicz W. Larry Sward John Game Katie Martin Chas Talbot Tamara Gedik* Roger D. Mays The New York Botanical Garden Lawrence Giles & Elizabeth Guy* Mary Ann McCrary Lynn R. Thomas Judith Graves* Sheila McMahon Robert F. Thorne Philip Greer Linda Bea Miller Yulan Tong Paul Grunland Alan E. Mohr Lorie Topinka Ann Gurbaxani* Peggy Moore Mehrey Vaghti* Susanne Haffner Reid Moran Linda Ann Vorobik Louise Hallberg Laura Moran* Christine Walden H. David Hammond Randall Morgan* Charlotte Waldman Mary Ann Hannon Gilbert Muth Georgie Waugh Clare B. Hardham Marcile Nevins* James Weigand Gloria Harris* Ivana Noell Cynthia Weiner In memory of John Harris Richard O’Donnell Cherie L. R. Wetzel Harvard University, Botany Library Nobu- George J. Omi In memory of Lawrence Heckard katsu Hasebe Bruce Orr Catherine Whiteside & Peter Mikkelsen Joyce Hawley Steve Patterson* Meghan Williams* Janell Hillman Joel T. Perlstein Hartmut Wisch* Kenneth R Himes Luellen Pierce* Carl Wishner Pete Holloran Barbara Pitschel George A. Yatskievych Ellen Holmes & Aaron Schusteff Joan Pont * New Members 9 Friends of the Jepson Herbarium Nonprofit Organization The Jepson Globe, Vol. 15 No. 1 1001 Valley Life Sciences Building #2465 U.S. Postage PAID University of California, Berkeley University of California Berkeley, CA 94720-2465

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Jepson Herbarium Public Programs

June 2004 July 2004 September 2004 Spring Mountains, Nevada Mt. Lassen Flora Vegetation Mapping *WL only June 17 - 20 July 22 - 25 September 8 - 12

For more information or to register for a class, please contact Cynthia Perrine at the Jepson Herbarium; phone: (510) 643-7008, email: [email protected]. Please visit our Web site at: http://ucjeps.berkeley.edu/jepwkshp.html