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Catalogue of Latvian Silphidae (Insecta: Coleoptera)

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Acta Biol. Univ. Daugavp. 19 (1) 2019 ISSN 1407 - 8953 CATALOGUE OF LATVIAN SILPHIDAE (INSECTA: COLEOPTERA) Andris Bukejs, Marina Janovska Bukejs A., Janovska M. 2019. Catalogue of Latvian Silphidae (Insecta: Coleoptera). Acta Biol. Univ. Daugavp., 19 (1): 17 – 29. A critical annotated catalogue of Silphidae species recorded in Latvia is presented. In total, 20 species belonging to 8 genera and two subfamilies are confirmed for Latvia. A complete bibliography of Latvian large carrion beetles is reviewed and analysed. The brief research history of Latvian Silphidae is presented. The Latvian silphid fauna is represented by five chorotypes. Species with wide distribution are prevailing: Asiatic-European (8 species or 40.0%), Palaearctic (4 species or 20.0%), and Centralasiatic-European (4 species or 20.0%). Key words: large carrion beetles, Silphidae, Latvia, fauna, catalogue, biogeography, biodiversity.. Andris Bukejs, Marina Janovska. Daugavpils University, Institute of Life Sciences and Technology, Vienības Str. 13, Daugavpils, LV - 5401, Latvia, E-mail: [email protected], [email protected] INTRODUCTION The first information on large carrion beetles Silphidae is divided into two subfamilies from the present territory of Latvia was Silphinae and Nicrophorinae, and contains about published in the second half of the 18th century 200 species in 15 genera worldwide (Bouchard by J.B. Fischer (1778, 1784, 1791). In the et al. 2011, Ślipiński et al. 2011). The family first edition of his monograph Versuch“ einer has a worldwide distribution but is predominant Naturgeschichte von Livland”, which introduced in Holarctic (temperate regions) (Sikes 2005, the natural history of Livland (Fischer 1778), 2008). The most of silphids are necrophagous three species of Silpha were mentioned: Silpha and feed on carrion; members of few genera (e.g. littoralis L. [Necrodes littoralis (Linnaeus)]; Ablattaria Reitter, Dendroxena Motschulsky Silpha seminulum L. [according to Perreau and Phosphuga Leach) are specialised (2004), it is Agathidium seminulum (Linnaeus, predators of small invertebrates (snails, slugs 1758) (Leiodidae)]; and Silpha aquatica L. and caterpillars) or phytophagous (genus [according to Hansen (2004), it is Helophorus Aclypea Reitter) (Nikolajev & Kozminykh aquaticus (Linnaeus, 1758) (Helophoridae)]. 2002, Sikes 2005, 2008, Pushkin 2015). After some additions (Fischer 1784), in the Large carrion beetles perform vital ecosystem second edition of his monograph, Fischer function of contributing to the breakdown and (1791) already were mentioned eight Silpha recycling of organic matter within the terrestrial species. Of them three species only belong to ecosystem (Peck 1990, Kalinova et al. 2009, Silphidae, namely S. littoralis L. [Necrodes Dekeirsschieter et al. 2011). littoralis (Linnaeus)], Silpha oscura L., and S. 17 Bukejs A., Janovska M. vespillo L. [Nicrophorus vespillo (Linnaeus)]. second edition of the monograph Seidlitz (1887- Other five listed Silpha species belongs to 1891), 12 species of Silphinae and 7 species of another Coleoptera families: Silpha aquatica L. Nicrophorinae were mentioned. [according to Hansen (2004), it is Helophorus aquaticus (Linnaeus, 1758) (Helophoridae)]; At the beginning of the 20th century, another Silpha ferruginea L. [according to Kolibáč checklist of Baltic Coleoptera was published (2007), it is Peltis ferruginea Linnaeus, 1758 (Rathlef 1905). In this list, 19 species of (Trogossitidae)]; Silpha quadripustulata L. Silphidae were reported for Latvia. [according to Jelínek & Audisio (2007), Silpha quadripustulata Linnaeus, 1761 is synonym of A few other papers containing faunal data Silpha quadripunctata Linnaeus, 1758 and this on large carrion beetles were published at the taxon belong to sap beetle genus Glischrochilus beginning of the 20th century (Heyden 1903, Reitter, 1873, G. quadripunctatus Linnaeus, Lindberg 1932, Lackschewitz & Mikutowicz 1758 (Nitidulidae)]; Silpha sabulosa L. 1939). [according to Iwan & Löbl (2008), it is Opatrum sabulosum Linnaeus, 1760 (Tenebrionidae)]; Four silphid species, namely Necrodes and Silpha seminulum L. [according to Perreau littoralis (Linnaeus), Nicrophorus germanicus (2004), it is Agathidium seminulum (Linnaeus, (Linnaeus), N. vespillo (Linnaeus), and N. 1758) (Leiodidae). vespilloides Herbst, were recorded in Latvian storehouses (Vītols 1941). In “Beschreibungen der Provinz Kurland” (Groschke 1805), the author mentioned Silpha Few species were mentioned as pest of vespillo L. [Nicrophorus vespillo (Linnaeus)], agriculture (Tomsons 1939-1940, 1940, Danka Silpha quadripustulata L. [according to 1950, Smarods & Liepa 1956). Jelínek & Audisio (2007), belong to genus Glischrochilus Reitter, 1873 (Nitidulidae), Fragmentary information on Latvian Silphidae see comment above], Silpha littoralis L. can also be found in Trauberga (1957), [Necrodes littoralis (Linnaeus)], Silpha atrata Brammanis (1930), Stiprais & Varzinska (1985), L. [Phosphuga atrata (Linnaeus)] and Silpha Barševskis (1988a, 1988b, 1997), Barševskis rugosa L. [Thanatophilu rugosus (Linnaeus)]. & Savenkovs (1991), Cibuļskis (1995, 1997), Telnov (1996), and Cinītis (1997). In the monograph devoted to the beetle fauna of “Ostseeprovinzen” (Fleischer 1829), six large In monograph “The Beetles of Eastern Latvia” carrion beetles were mentioned: Dendroxena (Barševskis 1993), faunal data on fifteen silphid quadrimaculata (Scopoli) [listed as Silpha species are presented. quadripunctata], Necrodes littoralis (Linnaeus) [as Silpha], Silpha tristis Illiger, Thanatophilu As for specially protected nature territories, dispar (Herbst) [as Silpha], Nicrophorus 16 species of Silphidae are known from Silene investigator Zetterstedt [as Necrophorus], Nature Park (Barševskis 2002), and 17 species – and Nicrophorus vespillo (Linnaeus) [as from Gauja National Park (Kalniņš et al. 2007). Necrophorus]. During the last decades, Gailis & Vilks (2001), At the end of the 19th century, Seidlitz’s Spuņģis (2002), Barševskis et al. (2004, 2008, monograph “Fauna Baltica” (Seidlitz 1872- 2009), Telnov et al. (2005, 2010), Bukejs 1875, 1887-1891) was published. In this (2006), Telnov & Salmane (2015), and Roze monograph, keys to families, genera and & Barševskis (2018) have also provided species of the Baltic Coleoptera (including fragmentary faunal data on Latvian large carrion potential ones) were presented. Unfortunately, beetles. not all data on localities were provided. In the 18 Catalogue of Latvian Silphidae (Insecta: Coleoptera) The catalogues of Latvian Coleoptera (Telnov ANNOTATED CATALOGUE OF et al. 1997, Telnov 2004) list 19 species of LATVIAN SILPHIDAE Silphidae. Family Silphidae Latreille, 1806 Two papers especially devoted to Latvian Subfamily Silphinae Latreille, 1806 Silphidae: Šmits (1975) present faunal data on 19 species, but Murd & Barševskis (2007) given Genus Aclypea Reitter, 1885 faunal data on 16 silphid species. =Blitophaga Reitter, 1885 The aim of the current work is to critically 1. Aclypea opaca (Linnaeus, 1758) summarize previous bibliographical data, to = hirta Herbst, 1783 compile the annotated catalogue of Latvian = tomentifera Reitter, 1907 Silphidae according to modern systematics, = tomentosa Villers, 1789 and to provide biogeographical analysis of the = villosa Naezén, 1792 Latvian fauna of large carrion beetles. = villosa Reitter, 1887 Precht 1818 (Silpha); Seidlitz 1872-1875 (Silpha), 1887-1891 (Silpha); Rathlef 1905 MATERIAL AND METHODS (Blitophaga); Tomsons 1939-1940, 1940; Ozols 1948; Danka 1950; Eglītis 1954; Smarods & Taxa listed in the current catalogue are arranged Liepa 1956; Trauberga 1957; Ozols 1963, 1973; in tribes and subfamilies. The order of higher taxa Šmits & Spuris 1966; Šmits 1975; Barševskis follows the Catalogue of Palaearctic Coleoptera, 1988a, 1988b, 1993 (Blitophaga), 2002; Telnov volume 2, chapter Silphidae (Růžička & et al. 1997; Silfverberg 2004, 2010; Růžička Schneider 2004; Růžička 2015) and Bouchard & Schneider 2004; Telnov 2004; Murd & et al. (2011), synonymy and nomenclature for Barševskis 2007; Kalniņš et al. 2007; Růžička the species-rank taxa were used in accordance 2015. with Sikes et al. (2002), Růžička & Schneider (2004), Růžička (2015). Species and genera are 2. Aclypea undata (O.F. Müller, 1776) listed alphabetically. = cancellata Gmelin, 1790 = kindermanni Faust, 1877 The species certainly recorded from Latvia = reticulata Fabricius, 1787 are marked with index number. Synonyms and = verrucosa Faldermann, 1835 references follow species names. For some Precht 1818 (Silpha reticulata); Seidlitz 1872- species a Note is presented. 1875 (Silpha), 1887-1891 (Silpha); Rathlef 1905 (Blitophaga); Tomsons 1940; Trauberga The general distribution of the species was 1957; Ozols 1963, 1973; Šmits 1975; Barševskis analysed according to Lafer (1989), Peck 1993, 2002; Telnov et al. 1997; Silfverberg & Miller (1993), Peck (2001), Nikolajev & 2004, 2010; Růžička & Schneider 2004; Telnov Kozminykh (2002), Sikes et al. (2002), Růžička 2004; Murd & Barševskis 2007; Růžička 2015. & Schneider (2004), Pushkin (2015), Růžička (2015), Ghahari & Háva (2015), and Çiftçi et al. Note: Insufficiently known species, known 2018. The classification of chorotypes follows from few localities. the one suggested by Taglianti et al. (1999). Dendroxena Abbreviations of chorotypes: OLA – Holarctic, Genus Motschulsky, 1858 =Xylodrepa Thomson, 1859 PAL – Palaearctic, ASE – Asiatic-European, Dendroxena quadrimaculata CAE – Centralasiatic-European, and WPA – 3. (Scopoli, West
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    Evolution of Uniparental Genetic Systems in Dermanyssine Mites (Acari: Mesostigmata) A THESIS SUBMITTED FOR THE DEGREE OF DOCTOR OF PHILOSOPHY OF THE UNIVERSITY OF LONDON by Robert Hamish Cruickshank Molecular Biology Unit, Department of Zoology, The Natural History Museum, Cromwell Road, South Kensington, LONDON SW7 5BD, England, UK Department of Biology, The Galton Laboratory, University College London, 4 Stephenson Way, LONDON NW l 2HE, England, UK September 1997 ProQuest Number: 10045787 All rights reserved INFORMATION TO ALL USERS The quality of this reproduction is dependent upon the quality of the copy submitted. In the unlikely event that the author did not send a complete manuscript and there are missing pages, these will be noted. Also, if material had to be removed, a note will indicate the deletion. uest. ProQuest 10045787 Published by ProQuest LLC(2016). Copyright of the Dissertation is held by the Author. All rights reserved. This work is protected against unauthorized copying under Title 17, United States Code. Microform Edition © ProQuest LLC. ProQuest LLC 789 East Eisenhower Parkway P.O. Box 1346 Ann Arbor, Ml 48106-1346 DEDICATION This thesis is dedicated, with love, to my wife Alice who I met and married whilst working on this project and who has put up with so much to help me get it finished. "The meaning of this degree is that the recipient of instruction is examined for the last time in his life, and is pronounced completely full. After this, no new ideas can be imparted to him. " Stephen Leacock "Sunshine Sketches of a Little Town” ACKNOWLEDGEMENTS This thesis could not have been completed without the guidence, enthusiasm and patience of my supervisor Dr.
  • Burying Beetles

    Burying Beetles

    Colorado Insect of Interest Burying Beetles Scientific Name: Nicrophorus spp. Order: Coleoptera (Beetles) Family: Silphidae (Carrion Beetles) Identification and Descriptive Features: The burying beetles are moderate-large sized (25-35 mm), brightly colored beetles marked with Figure 1. A burying beetle, Nicrophorus orange-red and black. Their antennae are investigaror, feeding on a piece of meat. Photograph pronouncedly club-like. courtesy of Joseph Berger/IPM Images Distribution in Colorado: Ten species of burying beetles are known from Colorado (Table 1) and one or more representatives can be found throughout the state. Life History and Habits: Burying beetles have unusual life histories that involve extraordinary involvement of adults of both sexes in larval rearing. Although variations may occur among the different Nicrophorus species, the general outline follows. Rearing of larvae begins upon location of a recently dead rodent, bird or other small carrion item. The adults are acutely sensitive to odors Figure 2. Burying beetle, Nicrophorus associated with early decay and the first beetle marginatus. Photograph courtesy of Insects in arriving at the carcass - either male or female - will Kansas. guard it and await a mate. When a member of the opposite sex arrives the pair will then inter the carcass, a process that can usually be completed in a few hours. Once the food item has been successfully covered it is further prepared by stripping all skin, fur and/or feathers and gathering the flesh into a pear-shaped ball. The flesh is chewed and covered with salivary and anal secretions that help to preserve it from decay. A rearing chamber is then dug out where eggs are laid.
  • Coleoptera, Silphidae)

    Coleoptera, Silphidae)

    African InvertebratesDetermination 58(2): 1–10 (2017) of species and instars of the larvae of the Afrotropical species... 1 doi: 10.3897/AfrInvertebr.58.12966 RESEARCH ARTICLE http://africaninvertebrates.pensoft.net Determination of species and instars of the larvae of the Afrotropical species of Thanatophilus Leach, 1817 (Coleoptera, Silphidae) Claire A. Daniel1, John M. Midgley1,2,3, Martin H. Villet1 1 Southern African Forensic Entomology Research Laboratory, Department of Zoology and Entomology, Rhodes University, P.O. Box 94, Grahamstown, 6140 South Africa 2 Department of Entomology and Arachnology, Albany Museum, Somerset Street, Grahamstown, 6139 South Africa 3 Current address: KwaZulu-Natal Mu- seum, Private Bag 9070, Pietermaritzburg, 3200, South Africa Corresponding author: John M. Midgley ([email protected]) Academic editor: P. Stoev | Received 29 March 2017 | Accepted 3 May 2017 | Published 17 May 2017 http://zoobank.org/75D5D52B-CB56-4B33-B4AE-02AEBBC1515B Citation: Daniel CA, Midgley JM, Villet MH (2017) Determination of species and instars of the larvae of the Afrotropical species of Thanatophilus Leach, 1817 (Coleoptera, Silphidae). African Invertebrates 58(2): 1–10. https://doi.org/10.3897/ AfrInvertebr.58.12966 Abstract Thanatophilus micans and T. mutilatus have significance for forensic entomology. Their larvae are therefore described and a key is provided for identifying the larvae of Afrotropical Silphidae based on morphologi- cal characters. It is shown that seven common species of Thanatophilus can be distinguished by a 360 bp mtDNA sequence from the cytochrome oxidase I gene. Keywords Thanatophilus, Silpha, Silphidae, taxonomic key, forensic entomology, Africa Copyright Claire A. Daniel et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.