have visual pigments that let Flying Colours them detect ultraviolet wavelengths. Some males have ultraviolet markings on their wings that they use to attract females.

There are no colours in the dark! Butterflies’ splendid hues show up Subtle ultraviolet markings when light hits the thousands of tiny scales that make up the distinctive markings on their wings.

Couleurs

papillons Hypolimnas bolina Phoebis philea Faboulous Great eggfly bolina Orange-barred sulphur

Linden Gledhill is a biochemist and Scales photographer who uses a microscope with an attached camera and a technique called focus stacking to capture the stunning beauty of wings at a unique . This lets him reveal the subtle nuances of the colours and different shapes of the scales, and the fascinating patterns they create.

Please don’t touch any of the butterflies or . Rothschildia jacobaeae Jacob’s silkmoth

Magnification Some butterflies have bare patches on their x 25 x 125 x 25 x 25 x 25 wing membranes. By letting light through, these transparent “windows” change the Left to right ’s shape and fool rhipheus • Hypolimnas dexithea • potential predators. Morpho zephyritis • Argema mittrei • blumei Attacus atlas Cobra

% trickery 100 Transparent Dryas julia Dryadula phaetusa Heliconius melpomene Papilio lowi Papilio rumanzovia Troides rhadamantus Julia Banded orange Postman Asian swallowtail Rumanzovia swallowtail Small troides

Papilio thoas Caligo sp. Danaus chrysippus Danaus plexippus Argema mimosae Idea leuconoe Thoas swallowtail Owl butterfly African monarch Monarch African moon Moth Rice paper

The scales on some butterflies’ wings get their colours from the plants that the fed on. The colours of the different scales combine to create patterns on A pigment mosaic the wings.

Morpho helenor Morpho polyphemus Siproeta stelenes Graphium agamemnon Blue morpho White morpho Banded peacock Malachite Tailed jay André Sarrazin As light strikes the internal and external structures of the scales on these butterflies’ wings, the wavelengths that reflect off them produce shimmering colours. Optical effects Rothschildia jacobaeae, jacobaeae, Rothschildia Mr Clean1982 • Dennis • Carol Hunt • Jas • Jim, the Photographer • Stéphane LeTirant • David Marquina Reyes • Reyes Marquina • David LeTirant • Stéphane Hunt • Jas Jim, the Photographer Mr Clean1982 • Dennis Carol Small photos: Beek van • Anneke • Anne Toal Strater Smith • Tim Seeger • Brad • Stuart Ruellan • André Sarrazin Mensen • Gabriela Rene F. photo: Large