CURRICULUM VITAE

E. Alan Verde Maine Maritime Academy Corning School of Ocean Studies Castine, ME 04420 (207) 326-2124 [email protected]

EDUCATION

Ph.D. 1993 Florida Institute of Technology Biological Sciences M.S. 1987 Walla Walla University Biological Sciences B.S. 1984 Walla Walla University Biological Sciences

AREAS OF COMPETENCY IN TEACHING

Biological Statistics Invertebrate Zoology Marine Ecology Coral Reef Biology Marine Biology Plant Physiology Cellular Biology Cellular Physiology Physiological Ecology Physiology

TEACHING EXPERIENCE

Maine Maritime Academy (2004-Present)

Professor of Marine Biology

Departmental Courses

Cell Biology (BI 308) Marine Ecology (BI 322) Scientific Diving (OS 300) General Biology I (BI 101) General Biology II (BI 102) Organic Chemistry (CH 310) Biology of Symbiosis (BI 321) Tropical Marine Biology (OS 325) Research Project Preparation (OS 400) Research Project Presentation (OS 401) Physiology of Marine Organisms (BI 301) Ocean Studies Seminar (OS I, II, III, IV, V) Service Courses

Chemical Principles (CH 301) Chemical Principles (labs of CH 301) Engineering Chemistry (labs of CH 352) Introduction to Oceanography (labs of OC 101)

Rosario Beach Marine Laboratory (1999)

Visiting Assistant Professor Marine Ecology

Florida Institute of Technology (1988-1995)

Visiting Assistant Professor Marine Ecology Invertebrate Zoology

Graduate Teaching Assistant

Biometry Marine Biology Marine Ecology Biological Statistics Invertebrate Zoology Comparative Vertebrate Anatomy

University of Arkansas (1987-1988)

Graduate Teaching Assistant Anatomy and Physiology Comparative Animal Physiology

Walla Walla University (1981-1987)

Graduate Teaching Assistant Cell Physiology General Biology General Ecology Plant Physiology Animal Physiology Anatomy and Physiology

Undergraduate Teaching Assistant General Biology General Chemistry Animal Physiology Anatomy and Physiology

2 Other Teaching Experience

Marine Biology and Marine Science for K-12 at The Living Seas, Walt Disney World Company, FL. Marine Biology and Biological Oceanography for Melbourne and Palm Bay, FL, Elementary Schools. Marine Biology and Marine Science for elementary students at Holland Christian School, Holland, MI. Invited Speaker for OR and WA High School Career Day Programs (Careers in Marine Biology). Interpretive Marine Intertidal Lectures for the Washington State Parks and Recreation.

RESEARCH EXPERIENCE

Ongoing Research Co-Investigator: In collaboration with Drs. Ann Cleveland and Ray Lee, we investigate the uptake and transfer of 13C and 15N stable isotopes between the intracellular algae (Symbiodinium sp.), anemone Heteractis crispa, and its clownfish Amphiprion clarkii and A. perideraion. Complimentary experiments in both laboratory and field conditions are conducted at the Silliman University Marine Laboratory, Dumaguete, Philippines.

Ongoing Research Co-Investigator: In collaboration with Dr. James Nestler, we investigate the diel patterns of behavior, migration, and metabolic rates, and both nitrogen and phosphorus cycling of both tropical (Pearsonothuria graeffei and Holothuria edulis) and temperate (Parastichopus californicus) sea cucumbers, as a function of environmental temperature. Tropical studies occur at the Silliman University Marine Laboratory, Dumaguete, Philippines whereas temperate studies occur at the Rosario Beach Marine Laboratory, Anacortes, WA.

Ongoing Research Co-Investigator: In collaboration with Drs. Kirt Onthank and Lloyd Trueblood, we investigate various aspects of the biology, ecology, and metabolic physiology of the red octopus, Octopus rubescens, as a function of lowered pH conditions and reduced oxygen availability at the Rosario Beach Marine Laboratory, Anacortes, WA.

2016 Co-Investigator: In conjunction with Taylor Grace (MMA Class of 2015), the sensory modes used by the red Pacific octopus ((Octopus rubescens) to select between scallops () that had () on them or not were investigated at the Rosario Beach Marine Laboratory, Anacortes, WA. The results of the study were presented (and published in the Proceedings) at the American Academy of Underwater Sciences conference at the University of Rhode Island, from 19-24 September 2016.

2015 Co-Investigator: In conjunction with Jillian Perron (MMA Class of 2015), various aspects of prey handling capabilities of the red Pacific octopus (Octopus rubescens) were conducted at the Rosario Beach Marine Laboratory, Anacortes, WA. The results of the study were presented (and published in the Proceedings) at the American Academy of Underwater Sciences conference at Key Largo, FL from 28 September to 3 October, 2015. A manuscript is to be submitted to the journal Marine Biology or Journal of Experimental Marine Biology and Ecology.

2011 Co-Investigator: In conjunction with Elizabeth Chase (MMA Class of 2012), various aspects of both the biology and ecology of the red Pacific octopus (Octopus rubescens) were conducted at Admiralty Bay, WA. The population densities at Admiralty Bay and the choice of food type and den material were investigated and the results of the study were presented (and published in the Proceedings) at the American Academy of Underwater Sciences conference at Portland, ME.

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1998-2004 Postdoctoral Research Associate: Utilized molecular methods to elucidate the communicative interactions between the algal symbionts (zooxanthellae and zoochlorellae) and host anemone, Anthopleura elegantissima, both in the adult and larval stages. In addition, the effects of various heavy metals, at the ecological and cellular levels of interaction, on the symbiosis of A. elegantissima and the tropical coral, Pocillopora damicornis, were also investigated. Both field and laboratory studies were conducted at the Hatfield Marine Science Center (Oregon State University) and at the Rosario Beach Marine Laboratory (Walla Walla University), WA.

1996 Co-Investigator: Investigated the feeding and physiological energetics between two clownfish , Amphiprion melanopus and Premnas biaculeatus, and their host anemone, Entacmaea quadricolor at the Christensen Research Institute, Madang, Papua New Guinea.

1995-1996 Co-Investigator: Examined the behavior and food preferences of two species of damselfish, Stegastes planifrons and S. dorsopunicans, as a function of territory at the Smithsonian Tropical Research Institute Marine Laboratory, San Blas, Panama.

1994 Research Assistant, National Parks Service: Documented and quantified both the flora and invertebrate fauna in the Colorado River Basin to identify the basic food sources of the various aquatic inhabitants of the Grand Canyon system.

1991-1993 Principal-Investigator: Investigated the daily carbon flux in the jellyfish Cassiopea xamachana in the Florida Keys. The carbon flow between the zooxanthellae symbiont and host medusae was compared as a function of jellyfish size and seasonal environmental change.

1992 Doctoral Dissertation Research: “The effects of temperature, light, season, and body size on the photosynthesis and respiration of zooxanthellae or zoochlorellae symbiotic within Anthopleura elegantissima (Brandt).” Investigated how different environmental and biological variables regulate the carbon transfer between the algal symbiont and anemone host.

1991 Participant, Joint U.S.-Israel Coral Workshop: “Nutrient limitation in the symbiotic association between zooxanthellae and reef-building corals.” Hosted by Hawaii Institute of Marine Biology. Examined the effects of elevated ammonium concentrations on the productivity, respiration, and algal symbionts of the coral Pocillopora damicornis.

1991 Co-Investigator: Investigated the differences in electron flow through Photosystem-I and Photosystem-II of zooxanthellae and zoochlorellae symbionts freshly isolated from their host anemone, Anthopleura elegantissima at Brookhaven National Laboratories, NY. A custom-made “pump-and-probe” fluorometer was operated to compare algal photosystem excitation and quenching rates.

1989-1991 Scientific Research Diver at The Living Seas, Walt Disney World Company: Investigated the sonar acuity of Atlantic Bottlenose dolphins to size, shape, and composition of various objects to elucidate dolphin cognition, spatial memory, and potential for communication with humans via iconic symbols.

1989 Research Assistant at the Space Research Institute: Conducted experimental and histological studies involving the morphological changes in the neurons of the mouse inner ear as a function of both ageing and diet restriction. 4

1987 Research Technician for Biosonics: Conducted hourly hydroacoustic measurements on juvenile salmon migrating down the Snake River, WA through the hydroelectric turbines and spillbays.

1987 Master’s Thesis Research: “A comparative analysis of the carbon budgets of the sea anemone Anthopleura elegantissima (Brandt) symbiotic with zooxanthellae or zoochlorellae.” The carbon flow between the algal symbionts and anemone host was investigated as a function of aerial exposure to simulate intertidal environmental conditions.

1986 Co-Investigator: The cardiac responses of the rattlesnake, Crotalus viridis, were analyzed during successive stages of prey tracking, immobilization, and swallowing and during courtship and fighting. The heart rates were characterized using surgically implanted subdermal electrodes that triangulated the heart.

1984 Research Assistant: Conducted studies on the carbon flow between symbiotic zooxanthellae and their cnidarian host at the Inter-University Institute for Marine Science, Eilat, Israel. The following coral(s) and scyphozoan were studied as a function of depth: Stylophora pistillata, Millepora dichotoma, Xenia sp., Heteroxenia sp., and Cassiopea andromeda.

Grants

2016 MMA Faculty Development Grants (Anemone Research and Octopus presentation at meeting; $11,309) 2015 MMA Faculty Development Grants (Clownfish/Anemone and Octopus Research; $8,160) 2014 MMA Faculty Development Grants (Sea Cucumbers presentation at meeting; $3,028) 2014 MMA Faculty Development Grant (Sea Cucumbers and Octopus Research; $2,960) 2013 MMA Faculty Development Grant (Clownfish/Anemone and Sea Cucumbers Research; $6,358) 2013 MMA Human Resources Tuition Grant (PADI Scuba Instructor certification, Philippines; $2,601) 2012 MMA Faculty Development Grants (DAN Instructor certification at meeting; $2,115) 2011 MMA Faculty Development Grants (Clownfish/Anemone presentations at two meetings; $5,324) 2010 MMA Faculty Development Grant (Clownfish/Anemone and Sea Cucumbers Research; $5,906) 2009 MMA Faculty Development Grant (Sea Cucumbers Research; $1,781) 2009 MMA Faculty Development Grant (Presentation at meeting; $1,623) 2008 MMA Faculty Development Grant (Presentations at two meetings; $3,447) 2007 Maine Aquaculture Innovation Center Grant (Polyculture: Bivalves and Sea Urchins; $22,000). 2006 MMA Faculty Development Grant (Sea Cucumbers; Anemone Symbiosis Research; $3,093) 2006 MMA Faculty Development Grant (Sea Cucumbers; Starfish Research; $1,745) 2005 MMA Faculty Development Grant (Anemone Symbiosis; Sea Cucumbers Research; $1,474) 2004 PADI American Project AWARE Foundation Grant (Sea Cucumbers Research; $2,000) 2003 PADI American Project AWARE Foundation Grant (Sea Cucumbers Research; $1,000) 2003 PADI American Project AWARE Foundation Grant (Clownfish/Anemones Research; $1,000) 5 2002 PADI Asia Pacific Project AWARE Foundation Grant (Clownfish/Anemones Research; $1,000) 1993 Houston Underwater Club SeaSpace Scholarship (Anemone Symbiosis Research; $2,000) 1992 Houston Underwater Club SeaSpace Scholarship (Anemone Symbiosis Research; $2,000) 1992 Walla Walla University Graduate Marine Scholarship (Anemone Symbiosis Research; $2,000) 1991 Walt Disney World Company Research Grant (Jellyfish Symbiosis Research; $16,000) 1990 Houston Underwater Club SeaSpace Scholarship (Anemone Symbiosis Research; $2,000) 1990 Sigma Xi Grants-in-Aid of Research (Anemone Symbiosis Research; $500)

Publications and Presentations

2016 Grace, T. J. W. and E. A. Verde. Prey preference and feeding behavior of Octopus rubescens on hastata (scallop) encrusted with incrustans (). Lobel, L. K and Lombardi, M. R. (eds). Proceedings of the American Academy of Underwater Sciences. 35:58-73.

2015 Verde E. A., Cleveland A., Lee R. W. Nutritional exchange in a tropical tripartite symbiosis II: direct evidence for the transfer of nutrients from host anemone and zooxanthellae to anemonefish. Mar Biol. 162:2409-2429.

2015 Perron, J. and E. A. Verde. Octopus rubescens’ Prey Handling Procedures are Influenced by Bivalve Shell Thickness and Adductor Muscle Strength. Lobel, L. K. (eds). Proceedings of the American Academy of Underwater Sciences. 34:78-94.

2014 Nestler, J. R., L. Cole, L. Dann, C. Leggitt, E. Manley, R. Reeve, A. Tan, E. A. Verde. Don’t Poop Where You Eat: Location of Sea Cucumber Egesta in a Coral Reef Environment. Proceedings of the American Academy of Underwater Sciences Diving for Science, Diving for Science 2014. Dauphin Island, AL. pp 1-7.

2011 Cleveland A., Verde E. A., Lee R. W. Nutritional exchange in a tropical tripartite symbiosis: direct evidence for the transfer of nutrients from anemonefish to host anemone and zooxanthellae. Mar Bio 158:589-602

2011 Chase, E. R. and E. A. Verde. Population density and choice of den and food made by Octopus rubescens collected from Admiralty Bay, Washington, in July 2011. Proceedings of the American Academy of Underwater Sciences, Diving for Science 2011. Dauphin Island, AL. pp 110-116.

2007 Verde, E. A. and L. R. McCloskey. A comparative analysis of the photosynthesis and respiration of zooxanthellae or zoochlorellae symbiotic in the temperate clonal anemone Anthopleura elegantissima (Brandt). III. Seasonal effects of natural light and temperature on photosynthesis and respiration. Mar. Biol. 152:775-792.

2007 Mitchelmore, C. L., E. A. Verde, and W. M. Weis. Uptake and partitioning of copper and cadmium in the coral Pocillopora damicornis. Aquat. Tox. 85:48-56

2007 Wheeling, R. J., E. A. Verde, J. R. Nestler. Diel cycles of activity, metabolism, and ammonium concentration in tropical holothurians. Mar. Biol. 152:297-305

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2003 Mitchelmore, C. L., E. A. Verde, A. H. Ringwood, and V. M. Weis. Differential accumulation of heavy metals in the sea anemone Anthopleura elegantissima as a function of symbiotic state. Aquat. Tox. 64:317-329.

2002 Verde, E. A. and L. R. McCloskey. A comparative analysis of the photosynthesis and respiration of zooxanthellae or zoochlorellae symbiotic in the temperate clonal anemone Anthopleura elegantissima (Brandt). II. Effect of light intensity. Mar. Biol. 141:225-239.

2002 Weis, V. M., E. A. Verde, and W.S. Reynolds. Characterization of a Short Form Peridinin- Chlorophyll-Protein (PCP) cDNA and Protein from the Symbiotic Dinoflagellate Symbiodinium muscatinei (Dinophyceae) from the Sea Anemone Anthopleura elegantissima (Cnidaria). J. Phycol. 38:157-163.

2002 Weis, V. M., E. A. Verde, A. Pribyl, and J. S. Schwarz. Aspects of the larval biology of the sea anemones Anthopleura elegantissima and A. artemisia. Invert. Biol. 121:190-201.

2001 Verde, E. A. and L. R. McCloskey. A comparative analysis of the photosynthesis and respiration of zooxanthellae or zoochlorellae symbiotic in the temperate clonal anemone Anthopleura elegantissima (Brandt). I. Effect of temperature. Mar. Biol. 138:477-489.

1998 Verde, E. A. and L. R. McCloskey. Production, respiration, and photophysiology of the mangrove jellyfish Cassiopea xamachana: effect of jellyfish size and season. Mar. Ecol. Prog. Ser. 168:147-162.

1996 Verde, E. A. and L. R. McCloskey. Carbon budget studies of cnidarian anemones-evidence in support of some assumptions. J. Exp. Mar. Biol. Ecol. 195:161-171.

1996 McCloskey, L. R., T. G. Cove, and E. A. Verde. Symbiont expulsion from the anemone Anthopleura elegantissima (Brandt)(Cnidaria; Anthozoa). J. Exp. Mar. Biol. Ecol. 195:173-186.

1996 Verde, E. A. and L. R. McCloskey. Photosynthesis and respiration of two species of algal symbionts in the anemone Anthopleura elegantissima (Brandt)(Cnidaria; Anthozoa). J. Exp. Mar. Biol. Ecol. 195:187-202.

1994 Hayes, W. K., E. A. Verde and F. E. Hayes. Cardiac responses during courtship, male-male fighting, and other activities in rattlesnakes. J. Tennessee Acad. Sci. 69:7-9.

1990 Park, J. C., K. C. Cook and E. A. Verde. Dietary restriction slows the abnormally rapid loss of spiral ganglia neurons in C57BL-6 mice. Hearing Res. 48:275-280.

Published Abstracts

2016 Grace, T. J. W. and E. A. Verde. Prey preference and feeding behavior of Octopus rubescens on (scallop) encrusted with Myxilla incrustans (sponge). Proceedings of the American Academy of Underwater Sciences. University of Rhode Island.

7 2015 Perron, J. and E. A. Verde. Octopus rubescens’ Prey Handling Procedures are Influenced by Bivalve Shell Thickness and Adductor Muscle Strength. Proceedings of the American Academy of Underwater Sciences. Key West, FL.

2014 Nestler, J. R., L. Cole, L. Dann, C. Leggitt, E. Manley, R. Reeve, A. Tan, E. A. Verde. Don’t Poop Where You Eat: Location of Sea Cucumber Egesta in a Coral Reef Environment. 33rd Proceedings of the American Academy of Underwater Sciences Diving for Science. University of Alaska Southeast, Sitka, AK.

2013 Verde, E. A., A. Cleveland, and R. Lee. Nutritional exchange in the anemonefish-anemone symbiosis: What Nemo didn’t tell you! 1st Joint International Scientific Diving Symposium, Curacao.

2012 Verde, E. A., A. Cleveland, and R. Lee. Nutritional exchange in the anemonefish-anemone symbiosis: What Nemo didn’t tell you! 12th International Coral Reef Symposium (ICRS), Cairns, Australia.

2012 Verde, E. A., A. Cleveland, and R. Lee. Nutritional exchange in the anemonefish-anemone symbiosis: What Nemo didn’t tell you! 7th International Symbiosis Society (ISS) Congress, Jagiellonian University, Krakow, Poland.

2012 Cleveland A., H. R. Jackson, E. A. Verde. Who is minding the anemone? Gender and species comparisons of territorial behavior in two sympatric anemonefish species (Amphiprion clarkii and A. perideraion). Evolutionary and Ecological Ethology of Fishes conference, University of Windsor, Ontario, Canada.

2011 Chase, E. R. and E. A. Verde. Population density and choice of den and food made by Octopus rubescens collected from Admiralty Bay, Washington, in July 2011. 30th American Academy of Underwater Sciences Diving for Science Symposium, Darling Marine Center, Walpole, ME and Portland, ME.

2009 Verde, E. A., A. Cleveland, and R. Lee. Nutritional Exchange in the Anemonefish/Anemone Symbiosis: Photosynthetically-fixed carbon is translocated to host fish. 6th International Symbiosis Society (ISS) Congress, University of Wisconsin, Madison, WI.

2008 Cleveland, A., E. A. Verde, and R Lee. Nutritional Exchange in the Anemonefish/Anemone Symbiosis: Photosynthetically-fixed carbon is translocated to host fish. 11th International Coral Reef Symposium, Fort Lauderdale, FL.

2008 Croston, H. L, H. G. Stence, M. G. Devin, R. J. Peacock, and E. A. Verde. Scallops and Urchins: Potential culture partners? 100th National Shellfish Association Conference in Providence, RI.

2006 Nestler J. R., R. J. Wheeling, and E. A. Verde. Diel cycles of activity, metabolism, and nutrient production in tropical sea cucumbers. 12th International Echinoderm Conference, University of New Hampshire.

2006 Cleveland A., E. A. Verde. Nutritional exchange in the clownfish/host anemone symbiosis: Stable isotope analysis demonstrates it is a tripartite exchange. Annual Meeting of the American Society of Ichthyologists and Herpetologists, New Orleans, LA.

8 2006 Cleveland A., E. A. Verde, and R. Lee. Nutritional exchange in the clownfish/host anemone symbiosis: Stable isotope analysis demonstrates nutrient exchange is bi-directional. 5th International Symbiosis Society Congress, University of Vienna, Austria.

2005 Yost D.M., E. A. Verde, and R. Lee. Differences in inorganic carbon and nitrogen assimilation as a function of symbiont type in the temperate sea anemone Anthopleura elegantissima. Wiley Exposition, Washington State University, Pullman, WA.

2005 Cleveland A., H. R. Jackson, and E. A. Verde. Behavioral comparisons of two sympatric clownfish species (Amphiprion clarkia and A. perideraion) as symbionts of the anemone Heteractis crispa. Annual Meeting of the American Society of Ichthyologists and Herpetologists, Tampa, FL.

2003 Cleveland A., E. A. Verde, and R. Lee. Investigating the physiological basis for the clownfish/host anemone symbiosis: Do resident fish provide their host with nitrogen? 4th International Symbiosis Society Conference, Halifax, Nova Scotia, Canada.

1995 Verde, E. A. The effects of temperature, light, season, and body size on the photosynthesis and respiration of zooxanthellae and zoochlorellae symbiotic within Anthopleura elegantissima (Brandt). Dissertation Initiative for the Advancement of Limnology and Oceanography.

1983 E. A. Verde, Hayes, W. K. Cardiac responses during courtship, male-male fighting, and other activities in rattlesnakes. Annual Meeting of the American Society of Ichthyologists and Herpetologists, Victoria, Canada.

Manuscripts in Preparation

Verde, E. A. and L. R. McCloskey. A comparative analysis of the photosynthesis and respiration of zooxanthellae or zoochlorellae symbiotic in the temperate clonal anemone Anthopleura elegantissima (Brandt). IV. Effect of anemone size and season. To be submitted to Mar. Biol. 23 pp., 10 figs.

Cleveland, A. and E. A. Verde. Specific dynamic action and resting metabolism in two species of herbivorous damselfishes (Stegastes). To be submitted to J. Exp. Mar. Biol. Ecol. 17 pp., 2 figs., 1 table.

Thesis and Dissertations

1993 Verde, E. A. The effects of temperature, light, season, and body size on the photosynthesis and respiration of zooxanthellae and zoochlorellae symbiotic within Anthopleura elegantissima (Brandt). Doctoral Dissertation. Florida Institute of Technology.

1987 Verde, E. A. A comparative analysis of the carbon budgets of the sea anemone, Anthopleura elegantissima (Brandt) symbiotic with zooxanthellae or zoochlorellae. M.S. Thesis. Walla Walla University.

1984 Verde, E. A. The carbon budgets of the sea anemone, Anthopleura xanthogrammica (Brandt) symbiotic with zoochlorellae. B.S. Thesis. Walla Walla University.

9 Invited Seminars

2016 Department of Biological Sciences, Auburn University, Auburn, AL

2015 CIEE Research Station, Bonaire, Netherland Antilles

2015 Corning School of Ocean Studies (Maine Maritime Academy), Castine, ME.

2012 Shannon Point Marine Center (Western Washington University), Anacortes, WA.

2010 Darling Marine Center (University of Maine, Orono), Walpole, ME.

2009 Taklong Marine Laboratory (University of the Philippines), Guimaras, Philippines.

2007 Silliman University Marine Laboratory (Silliman University), Dumaguete, Philippines.

2005 Rosario Beach Marine Station (Walla Walla University), Anacortes, WA.

2004 Environmental Science Department (Alaska Pacific University), Anchorage, AK.

2004 Silliman University Marine Laboratory (Silliman University), Dumaguete, Philippines.

2004 Corning School of Ocean Studies (Maine Maritime Academy), Castine, ME.

2003 Department of Biological Sciences (Washington State University), Pullman, WA.

2000 Department of Applied Biology (Hiroshima University), Hiroshima, Japan.

2000 Ishigaki Tropical Research Station (Seikai National Fisheries Institute), Ishigaki, Japan.

1999 Department of Biological Sciences (Lewis and Clark College), Portland, OR.

1998 Department of Zoology (Oregon State University), Corvallis, OR

1997 Department of Biological Sciences (Loma Linda University), Loma Linda, CA.

1996 Department of Biological Sciences (Northern Arizona University), Flagstaff, AZ.

Membership in Professional Societies

Divers Alert Network International Symbiosis Society American Physiological Society International Society for Reef Studies American Academy of Underwater Sciences Professional Association of Diving Instructors Society for Integrative and Comparative Biology 10 Licenses/Certifications

1979 YMCA SCUBA Diver 1982 NAUI Advanced Open Water Diver 1995 Scientific Diver (AAUS Protocols at the Smithsonian Tropical Research Institute) 2005 PADI Nitrox Diver 2005 Scientific Diver (AAUS Protocols at Western Washington University) 2009 PADI Rescue Diver 2009 PADI Dive Master 2012 DAN Pro Instructor for: CPR: Health Care Provider Basic Life Support: CPR and First Aid Diving First Aid for Professional Divers Diving Emergency Management Provider Emergency Oxygen for Scuba Diving Injuries First Aid for Hazardous Marine Life Injuries On-Site Neurological Assessment for Divers 2013 PADI Open Water SCUBA Instructor

2016 PADI Sidemount Diver

2016 TDI-AUP Open Circuit Equipment Service Technician

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