Karstenia 25: 1-16. 1985

Floristic notes on Palaearctic (Ascomycetes)

LARRY HULDEN

HULDEN, L. 1985: Floristic notes on Palaearctic Laboulbeniales (Ascomycetes).­ Karstenia 25: 1-16. Floristic notes arc presented on 53 taxa of Laboulbeniales from various parts of the Palaearctic Regwn and one taxon from Taiwan. New species records are given for 33 countries or islands. Special attention is paid to the U.S.S.R., for which there are only scattered records in the literature. The northernmost records of Laboulbeniales are reported from Tit-Ary (Rickia hyperborea Balazuc) and Bulun (Laboulbenia vulgaris Peyritsch), by the Lena River, north of 70°N in Yakutia. Laboulbenia egens Spegazzini, occurring on the carabidicolous genus Tachys (s.Iat.), is recognized as a spec1es separate from L. pedicel/ala Thaxter, occurring on the genera Bembidion (s.lat.) and Dyschirius. The variability of L. pedicel/a/a is discussed. Five new taxa are described: Laboulbenia broscosomae n.sp. on Broscosoma baldense Rosenh. from , L. eubradyce/li n.sp. on Bradyce/lus spp. from many European countnes, L. kobi/ae n.sp. on Neotrechus sururalis ssp. suturalis Schr. from , L. marvinii n.sp. on Bembidion dentellum (Thunb.) and B. starki Schaum from , the Federal Repubhc of and , and L. luxurians subsp. immaculata n.subsp. on Bembidion semipunctatum (Donovan) from Austria and the Leningrad Region in the U.S.S.R. Larry Hulden, Department of Entomology, Zoological Museum, P. Rautatiekatu 13, SF-00100 Helsinki, .

Introduction The material in the present study is primarily Remarks on the distributional data based on the Palaearctic collection in the As far as possible the locality information is given Zoological Museum of Helsinki (MZH). Additional in accordance with modern atlases, because the labels material has been obtained from the insect collections on the old insect material may be rather cryptic unless of the Zoological Museum of Turku, Finland (cave supplemented with information from old diaries, etc. from ), the Zoological Museum of in the Zoological Museum of Helsinki. In the few Bergen, (Coleoptera from the Faeroes), the cases in which the exact locality is not known, the Institute of Zoology and Botany, Academy of Sciences smallest known modern geographical unit is given. of the Estonian S.S. R. in Tartu, U.S.S.R. (Coleoptera In some exceptional cases, like 'Germania' , the text from ), Mr. Bjorn Erling Waage and Dr. is given as such, because the area extends over many Dagfinn Refseth (Carabidae from Norway) and Prof. countries nowadays. Old records from the present Alois Kofler (Coleoptera from Austria). Some Israel with its immediate surroundings are given under specimens collected by myself in Estonia during the Palestine. Second Estonian-Finnish Entomological The Laboulbeniales of the U.S.S.R. have previously Symposium in 1982 are also included. About 25 000 been poorly studied. In this paper distributional data specimens of selected material of Coleoptera are given for 24 taxa from numerous localities. Due (Carabidae, Gyrinidae, Staphylinidae), Diptera to the large area of the country, this means a significant (Ephydridae, Nycteribidae) and Hymenoptera extension of the known distribution of many species. (Formicidae) were studied. As many of the locality names are poorly known and The preparation of the fungus material was done probably difficult to find, a map containing all the as described in Hulden (1983). The fungus material, collecting sites in the U.S.S.R. (and one in Iran) is including types, and the infected insects, except the given in Fig. I. Most of the localities on the labels borrowed specimens, are kept in a separate collection could be interpreted with good accuracy except those in MZH. in the Southern Urals in which G.A. Duske collected Twenty taxa, including the new ones, are illustrated insects (shown as an open circle on the map). in this paper, most of the others in Hulden (1983). The rest of the localities are presumably easy to For Arthrorhynchus see Blackwell ( 1980a, 1980b ), and identify . The Faeroes (belonging to ), Azores for Rhachomyces see Lepesme ( 1942), Balazuc ( 1970) and Madeira () and the Canary Islands () and Rossi ( 1978). are treated as separate units in the material lists. 2 L. Hulden

Review of the species (Middelhoek & Boelens) Scheloske 1969:92. Floristic notes are presented for 54 taxa of the Material examined Laboulbeniales from 37 countries or islands. New Sweden. Skane: Tykarpsgrottan, on Catops fuliginosus species records are given for 33 countries and islands Erichson. Balsbergsgrottan , on Catops fuliginosus and C. nigricans (Spence). The hosts were found in caves in both (marked with an asterisk *). The genera and species localities. are in alphabetical order. The numbers in the country­ species index refer to the species. General distribution Albania *35 Czechoslovakia, Federal Republic of Germany, Finland, Algeria 35 France, Italy, , , Sweden and U.S.S.R. Austria *4, 12, 14, *22, *23, 24, *26, 28, 30, 32, *34, 35, (Hulden 1983). *36, 39,41, *46, *49. Azores *35, *52. Host *11. Catops spp., Catopoides spp. and Sciodrepoides spp. 14, *31, 35. (Coleoptera, Carabidae). Canary Islands *9, 14, *35, *36, *38, *53. Czechoslovakia *7, *9, 15, *21, 35, *37, *39. 3. Cantharomyces orientalis Denmark *12, *36. -Figs. 4a-d Egypt *9 , *27. Cantharomyces orienta/is Spegazzini 1915b:43. Faeroes *14, *30, *35. Variable in size, 130-240 J..Lm in length. Larger Federal Republic of Germany 14, 15, 21, *23, 27, 35. specimens have the basal cells of the perithecium more Finland *II, *16. France 14, *23, 27, 31 , 35, 36, 39, 40, 47. elongated than small specimens. The compound German Democratic Republic *5, *35, 39, *41, *49. antheridium variable, situated on the second Greece *15, 27, *35, *36. appendage cell (Figs. 4 b and d). The differences in *41. size and habitus do not seem to be correlated with Iran • 36. the host species. Ireland *4, *14, *27 , *35, *36, *39. Material examined Italy *6, *II, 35, *36, *43, 44, *45. *Sweden. Uppland: Stockholm, on Carpelimus corticinus Lebanon *29. (Gravenhorst). *. Ticino: Chiasso, on C. opacus Madeira *II, *35. Baudi. U.S.S.R. *Yakut A.S.S.R.: Nikolsk and Ytyk-Haja, Morocco *35. on C. corticinus. Netherlands *15, 27, 35. General distribution Norway *4, • 5, *7, *II, * 12, *14, *24, *25, *35, *37, 50. Algeria, Federal Republic of Germany, Finland, Italy, Palestine *9, * 14, *35 , *36. Netherlands, Poland and U.S.S.R. (Hulden 1983). Poland 12, 35. Romania 35, 39 . Host Spain 7, *9, 35 , 36, Carpelimus spp. (Coleoptera, Staphyli nidae). Sweden 2, *3, *4, *5 , *7, *8, *10, *12, *15, *27, *31 , *35, *37, *39, *41, *49, *54. Switzerland *3 , 4, 14, * 15, 30, 35. 4. Laboulbenia argutoris Syria *35. Laboulbenia argutoris Cepede & Picard 1909:260. Taiwan *I. Material examined Turkey *27, 35. *Austria. Hohe Tauern (Pterostichus diligens (Sturm)). United Kingdom *II, 12, 15, 24, 35, 39. *Ireland. Kerry: Killarney (P. diligens, P. strenuus (Panzer)). U.S.S.R. 3, 4, 7, *10, *II, 12, *13, 14, 15, *16, *17, *18, *Norway. Hordaland: Linga (P. strenuus). *Sweden. Skane: *21 , *22, *24, 27,30, 31, *34, 35, 36, 38, 39,50. Ringsji:in (P. diligens). - Uppland (P. strenuus). Switzerland. Yugoslavia *19, 33, 35,41, 42, *51. Graubunden: Engadin (P. diligens). U.S.S.R. *Estonian S.S.R.: Harju r. , N6mmeveski (P. strenuus). 1. Arthrorhynchus nycteribiae General distribution Laboulbenia nycteribiae Peyritsch 1871:451. Federal Republic of Germany, Finland, France, Hungary, Helminthophana nycteribiae (Peyritsch) Peyritsch Italy, Poland, Romania, Switzerland and U.S.S.R. (Hulden 1873:250. - Arthrorhynchus nycteribiae (Peyritsch) 1983). T haxter 1901 :408. Host Material examined Prerostichus spp. (Coleoptera, Carabidae). *Taiwan. Tainan. On Penicillidiajenynsii (Westw.) (Diptera, Nycteribiidae). No data on the host species of the bat fly. 5. Laboulbenia bradycelli General distribution Laboulbenia bradycelli Balazuc 1974, tn Balazuc Compiled from Ryberg (1947), Balazuc (1971) and Blackwell 1973- 74, vol. 43:15. (1980a): Austria, Burma, Czechoslovakia, Denmark, France, Material examined Hungary, Italy, Kenya, Netherlands, Poland ('East Prussia'), *German Democratic Republic. Eberswalde: Chorin Romania, Spain, Sri Lanka, Sweden, Yugoslavia and (Trichocellus placidus (Gyllenhal)). *Norway. Hordaland: Zambia. . Skogsi:iy (Bradycellus ruficollis (Stephens)). *Sweden. Host Uppland: Stockholm (Trichocellus placidus). Batflies of the genera Eucampsipoda. Nycteribia, Penicillidia and Phthiridium (Diptera, Nycteribiidae). General distribution Finland, France, Poland and U.S.S.R. (Hu1den 1983). 2. Asaphomyces tubanticus Hosr Barbariella tubantica Middelhoek & Boelens in Trichocellus placidus and Bradycellus spp. (Coleoptera, Middelhoek 1949:260. Asaphomyces tubanticus Carabidae). Karstenia 25, 1985 3

Fig. 1. Localities of Laboulbeniales records in the U.S.S.R. Carabidae ). Holotype. Slide L. Hulden 26, in MZH. Collecting data of the host: Italy. Mte. Baldo. 6. Laboulbenia broscosomae Hulden n.sp. _ Fig. 16 Spegazzini ( 1914) reported another species, L. rig ida Crassulus. Matrix: Broscosoma baldense Rosenhauer. Thaxter, from the same host and locality. His Receptacle. Pale yellowish, slightly darkened along microphotograph (Fig. 54) shows that his material the margin of the appendical side. Cells I-II forming represents L. broscosomae. L. broscosomae differs a stalk about 110 ~-tm in length, 25-35 ~-tm in width. from L. rigida in its more rounded and clumsy habitus Cell I distinctly shorter than cell II. Cells III-IV and different host genus (L. rig ida was described from about 60 11m in length, cell III longer than cell IV. Pterostichus). Cell V small, oblique, not connected with cell III. Material examined. See the type. Cell IV nearly isodiametric, about 25 ~-tm in width. 7. Laboulbenia clivinalis Appendages. Insertion cell externally broadly black, Laboulbenia clivinalis Thaxter 1899: 165. situated slightly below middle of the perithecium. Ma1eria/ examined Outer appendage consisting of a simple row of at *Czechoslovakia. Paskau (near Ostrava), on Clivina collaris least seven cells, which are equal in size, 25 ~-tm in (Herbst). *Norway. Hordaland: Linga (Clivina fossor length a nd IS ~-tm in width. Inner appendage (1,.-innaeus)). Spain. Guipuzcoa: I ru n (C. fossor). *Sweden. consisting of two simple rows of cells similar to that Oland: Vickleby (C. fossor): U.S.S.R. *Archangel Region: of the outer appendage, except for the basal cell which Solovetsk Islands (C. fossor). - *Caucasus (C. fossor). is about 13 ~-tm in width. One antheridium visible General dislribulion on the second cell of the inner appendage. T he Algeria, Federal Republic of Germany, Finland, France, appendages 125- 150 ~-tm in length. Hungary, Italy, Philippines, Poland, Spain, United Kingdom Perithecium. Brown, paler near the sutures of the a nd U.S.S.R. (Hulden 1983). cells, blackish around the tip. Habitus robust, Hos1 rounded and stout, tip nearly straight, size about 60 Clivina spp. (Coleoptera, Carabidae). x 120 ~-tm. Spores not observed. 8. Laboulbenia curtipes Total length from base of foot to tip of perithecium Laboulbenia curtipes Thaxter 1892:40. about 250 ~-tm . Maleria/ examined Host. Broscosoma baldense Rosenh. (Coleoptera, *Sweden. Skane: Bokeberg. On Bembidion obliquum Sturm. 4 L. Hulden

General distribution 11. Laboulbenia eubradycelli Hulden n.sp. Fin land, U.S.A. and U.S.S.R. (Hulden 1983). -Figs. 2 (map), 12a-c Ur Laboulbenia bradycelli sed sporis majoribus. Matrix: Host. Bradycellus spp. . Bembidion: subgen. Notaphus and B. dentellum (Thu nberg) Receptacle. Pale yellowish, cell I usu~lly browmsh (Coleoptera, Carabidae). in distal part, cells III-IV to vanable d~gree brownish darkening upwards. Cells I- II formmg a 9. Laboulbenia egens - Figs. 22a- b rather e l ~ ngated stalk, 120-200 11m in length, the Laboulbenia paupercula Spegazzini !9!5b:59, auct. width increasing from about 20 11m at the .b~se to non Thaxter 1891 :269. Laboulbenia egens 25- 50 11m in the upper part. Cell I usually dtstmctly Spegazzini 1918:323. . . shorter than cell II. Cells III-IV 55-70 11m in length, This species has been confused with L. pedtcellata. cell III being slightly longer than cell IV. Cell V rather It grows on Tachys species (Coleoptera, Carabidae), small , not connected with cell III. Cell VI almost while L. pedicel/ala occurs mamly on .the. g~ nera isodiametric or slightly elongated, about 20-35 11m Dyschirius and Bembidion (s.lat.). L. egens IS dtstmctly in width. separable from L. pedicel/ala by tts smaller hp ~ells Appendages. Insertion ce.ll externally sh~rply or (more pointed perithecium), m?re or less dtsttpct diffusely blackish brown, sttuated below mtdd~e of humps on outer side on pentheciUm? usually ~erttcal perithecium. Outer appendage conststmg of a smgle and slightly shorter septu.m IV-V (m L. pedtcellata row of 5-10 cells about equal in length distally it is normally slightly obhque, but thts character ts slightly narrowing. Inner appendage consisting of a not constant) and strongly protruding fi rst cell of small brush of about 5-10 branchlets, each outer appendage (see Fig. 22b). The last-mentioned terminating in an antheridium. Width of basal cell character is typical of L. picardii Maire and L. tachyts os inner appendage 112 to 3/4 of that of basal cell Thaxter, also occurring on Tachys spp, but t h~se of outer appendage. Length of outer appendage about species, do not have septum IV-V connected wtth 100- 150 J.Lm, inner appendage shorter and not septum III-IV. usually reaching tip of perithecium. L. tachyis has been reported from Japan on Tachys Perithecium. Brown, the sutures of the cells darker. laetificus Bates (Sugiyama 1973), from Hungary on Black spots on lip cells. Perithecium straight but tip Tachyta nana (Gyllenhal) (Banhegyi 1949) and from slightly oblique. Size 40-70 X 85-125 J.Lm. Spores Taiwan on Tachys klugii Nietner (Sugiyama 1978). about 3 X 45 11m. The illustration in Sugiyama ( 1973), however, Total length from base of foot to tip of perithecium represents L. egens. The two latter references lack 230-370 J.Lm . . . illustrations, but the reports probably also represent Host. Bradycellus harpalinus (Audtnet-Servtlle), B. L. egens. Balazuc's ( 1973- 74, 1978, 1982) reports of caucasicus Chaudoir, B. ruficollis (Stephens) L. pedicel/ala from France, Madagascar and (Coleoptera, Carabidae). Probably also many other Guadeloupe on Tachys (s. lat.) also seem to represent Bradycellus species. The parasite occurs on vanous L. egens, cf. illustration in Balazuc ( 1982). parts of the host. Material examined. Holotype. Slide L. Hulden I, in MZH. On *Canary Islands. (Spain), Gran Canaria: Arucas (Tachys Bradycellus caucasicus. Collecting data of the host: haemorrhoidalis (Ponza)). - Tenerife (T. haemorrhordalts). Finland. Ab: Lohja, 7.V.l893 Luther. *Czechoslovakia. Paskau (T. quadrisignatus (Duftschmtd)). *Egypt. Cairo. (T. lucasi Du~a l , T . .Jumigat us ssp. soci~s In Hulden (1983) this species occurring on Schaum). - Luxor (T. lucasi). *Palestme. Jencho (T. lucasi). Bradycellus spp. was wrongly interpreted as *Spain. Cordoba (T. lucasi). Laboulbenia polyphaga Thax.ter. . Laboulbe.ma eubradycelli diffe rs from that spectes tn tts la~ger ~ 1ze , General distribution Compiled from the literature and my own records (see larger lip cells, angular habitus of the pe~1thecmm comments above): Canary Islands, Czechoslo\akia, Egypt, and more strongly pigmented thallus. It ts closely France, Guadeloupe, Hungary, Japan, Madagascar, related to L. bradycel/i, described by Balazuc ( 1974, Palestine, Spain and Taiwan. in 1973-74). L. bradycelli seems to be much more common on Trichocellus placidus Gyllenhal) than on Host Bradycellus spp. It is possible tha~ some matenal of Tachys (s.lat.) spp. (Coleoptera, Carabidae). L. bradycelli studied by Balazuc ~n fact. belongs to the new species. L. bradycelli ~~ dtfferenttated by tts 10. Laboulbenia elaphri smaller size (130-200 J.!ffi agamst 230-37.0 Mm for Laboulbenia elaphri Spegazzini 1915a:464. L. eubradycel/i), cells I and II about equ.al tn len~th, Material examined the somewhat greenish tinge of the dark ptgmentatwn, *Sweden. Uppland: Stockholm, on Elaphr~s cupreus simpler inner appendage, longer cells of outer Duftschmid. *U.S.S.R. Estoman S.S. R.: SeadJarv, on E. appendage, basal cell of oute.r appendage more than cupreus. twice the size of basal cell of mner appendage, flatter General distribution cell VI more oblique tip of perithecium and slightly Belgium, Denmark, Federal Republic of Germany, Finland, smalle; spores (about 38 11m against about 45 11m France, Hungary and Poland (Hulden 1983). for L. eubradycelli). . The new species varies somewhat tn the shape of Host . cupreus and£. riparius (Coleoptera, Carabtdae). the perithecium, which usually has an angular Karstenia 25, 1985 5

(P. atrorufus). *Sweden. Uppland: Lidingo (P. atrorufus). -Gotland: Klinte. (P. alrorufus). United Kingdom. Scotland. Moray, Forres (P. assimilis Chaudoir). U.S.S.R. Estonian S.S.R.: Paldiski, Vaike-Pakri s. (P. atrorufus). - *Pskov Region: Pechory (P. atrorufus).

General distribution Holarctic, in known from Austria, Federal Republic of Germany, Finland, France, Greece, Hungary, Italy, Poland, Romania, Spain, S\\itzerland, United Kingdom, U.S.S.R. (Leningrdd Region, Karelian A.S.S.R., Estoman S.S.R., Latvian S.S.R.) (Balazuc et al. 1983, Hulden 1983). Host Patrobus atrorufus is presumably the main host, but L. fasciculata also occurs on many other speciCs of Patrobus and related genera (Coleoptera, Carabidae). 13. Laboulbenia fennica Figs. 2-3. - Distribution of 2) Laboulbenia eubradycelli Laboulbenia fennica Hulden 1983:54. and 3) L. polyphaga in Eastern Fennoscandia. Material examined *U.S.S.R. Estonian S.S.R.: Vastse-Kuuste, Loot\ ina appearance, but may a lso be more rounded (filled ( marinus Gyllenhal). with spores) or rather narrow. The spores are usually General dislribution about 45 ILm in length, but in one case they were Previously reported only from Finland (Huldi:n 1983). 42 1-1m and in another about 48 1-1m. These probably represent different strains within the species. . Host. Gyrinus spp. G. aeratus Stephens being the main host 111 The distribution of Laboulbenia eubradyce/11 and F inland (Coleoptera, Gyrinidae). L. polyphaga in Eastern Fennoscandia are given in Figs. 2 and 3 because of the previOus confusiOns m Laboulbenia fl agellata Hulden (1983). 14. Labou/benia flagellata Peyri tsch 187 3:247. Material examined Malerial examined *Belgium. Brabant: Tervuren (Bradycellus harpalinus). Austria. Tirol. (Agonum moestum (Duftschmid)). - • Finland. Ab: Uusikaupunki (B. rujicollis, B. caucastcus). Burgenland: Neusiedler See (A. thoreyi Dejean): Bulgaria. Lohja (B. caucasicus). Vihti (B . . caucasicus). Turku (B. Rila Mountais. (Agonum antennarium (Duftschm1d. Canary caucasicus). N: Hango, Tvlirmmne (B. ruf tcollts: B. Islands (Spain). Gomcra (A . marginatum (Linnaeus)). caucasicus). Helsinki (B. caucasicus). Espoo. (B. caucastcus). *Faeroes (Denmark). Vagar: Si:irvagur (Patrobus Tuusula (B. caucasicus) Sjundea. (B. caucasicus). - Ta: septentrionis Dejean). - Suduroy: Ven (P._ septentrionis). Hartola (B. caucasicus). Ylojlirvi (B. coucasicus). Plilkline Federal Republic of Germany. Hamburg(A . mtcans (N1colat)). (B. caucasicus). Hattula (B. caucastcus). --: Sa: Joutseno(B. France. Allier: Vichy (A. moestum (Duftschmid)). *Ireland. caucasicus).-Sb. Vehmersalm1 (B. caucastcus)._- Kb: Knee Kerry: Killarney (A . muelleri (Herbst)). *Norway. T roms: (B. caucasicus). - Ok: Ristijlirvi (B .. car:castcus). -. Ks: Finsnes (Patrobus septentrionis). - Sor-Trondelag: Melhus Kuusamo (B. caucasicus) . - L1: UtSJOki (B. caucastcus) . (A . micans). *Palestine. Kafr Kanna (A. nigrum Dejean). *Italy. 'Parco Abruzzo' (B. caucasicus). *Madeira. Switzerland. Solothurn: Dornach (A. viduum (Panzer), A. (Portugal). On B. harpalinus. • Norway. Hordaland: Skogsoy muelleri). U.S.S.R. Moscow (A . Jhoreyi). - Krasnoyarsk (B.harpa/inus). Sor-Trondelag: Agdenes (B. caucastcus) (A. bicolor Dejean). - Novorossiysk (A. nigrum). - Tuva *United Kingdom. Scotland: Moray, Forres (B. caucastcus). A.S.S.R.: Soldan (A. thoreyi). *U.S.S.R. Leningrad Region: Kamennogorsk (Antrea). Segeza. Uslanka.-Karelian A.S.S.R.: Sortavala, P1rkmtsa, General distribution Ahvenjlin i. Kendjlirvi. SemSJlin 1. All records on B. Nearly cosmopolitan (Hulden 1983). caucasicus. Host. Mainly Agonum spp. (Coleoptera, Carabidae). 12. Labou1benia fasciculata Laboulbenia jasciculata Peyritsch 1873:248. 15. Laboulbenia giardii In Finland this species occurs only m the south Labou/benia giardii Cepede & Picard 1909:258. (Hulden 1983). It is interesting to note that on the Material examined Norwegian coast it reaches as far north as the.Tro.ms Federal Republic of Germany. Niedersachsen: Helmstedt. region, thus very exactly followmg the distnbutwn (Dicheirotrichus gustavi Crotch). *Greece. Attika: Elevsis (D. of Patrobus atrorufus (Strom). obsolelus (Dejean)). *Netherlands. Texel: De Koog (D. gustavi). *Sweden. Bohusllin: Koster (D. gustavi). Material examined *Switzerland. Valais: Saxon (D. gustavi). United Kingdom. Austria. Klirnten: Koralpe (Patrobus slyriacus Chaudoir). - Scotland: Moray, Forres (D. gustavi) . U.S.S.R. *Novosibirsk Tirol (P. septentrionis Dejean). - Steiermark: Turnau (P. Region: Karasuk steppe (D. obsoletus). - *Archangel styriacus). *Denmark. Sjalland: Dyreha,·en (P.atrorufus Region: Solovetsk Islands (D. guslavi). - Ural Mts., no (Strom)). *Norway. Hordaland: Askoy, Gjerstad. Hjeltnes. locality (Dicheirotrichus sp.). Ullensvang. Nordheimsund. Linga (P. atrorufus) . Ystanes (Synuchus viva/is (JIIiger)). - More og Romsdal: Molde (P. General distribution atrorufus). Sjoholt (P. atrorufus, P. asstmths). - Sor­ Finland, France, 'Germania', Hungary, Italy, United T rondelag: Trondheim. Melhus. Rognes. (P. atrorufus). - Kingdom and U.S.S.R. (Karelian A.S.S.R., Tomsk Region), Troms: Finsnes (P. septentrionis). Poland: Katowice: Cleszyn (Hulden 1983). 6 L. Hulden

Host Perithecium. Blackish brown, black spot on inner Dicheirotrichus spp. (Coleoptera, Carabidae). side near the yellow tip. Size 40-45 X 135-140 J.Lm. Only upper fo urth of perithecium free, abruptly 16. Laboulbenia gyrinicola narrowing towards the slightly outwards bent tip. Laboulbenia gyrinicola Spegazzini 1914:34. Spores about 5 X 55 J.Lm in size. ,\1aterial examined Total length from base of foot to tip of perithecium *Finland. AI: Jomala ( Gyrinus natator (Linnaeus)). *U.S.S.R. Estonian S.S.R.: Tartu r., Vahi (G. substriatus Stephens). 330-350 J.Lm. - P6ha r., Taevaskoja (G. natator. G. marinus Gyllenhal). Host. Neotrechus suturalis ssp. sutura/is Schaufuss (Coleoptera, Carabidae). The parasite was found on General disTribution the pronotum of the host. Algeria, Belgium, Czechoslovakia, France, Hungary, Italy, Holotype. Slide L. H ulden 27, in MZH. Collecting Poland, Romania, Sweden, Switzerland, Yugoslavia (Hulden data of the host: Yugoslavia. Bosnia-Hercegovina: 1983). Records from the United Kingdom and Spain possibly Kobila Glava. concern L. fennica. Relationship with other species unclear. L. shanorii Hosr Banhegyi, described from the same host, is quite Gyrinus spp. and Aulonogyrus spp. (Coleoptera, Gyrinidae). different (see Fig. 14). T he exact structure of the appendages is not certain; the illustration in Fig. 15 17. Laboulbenia hastiana is only schematic. Laboulbenia hastiana Hulden 1983: 55. Material examined. This species is closely related to L. vulgaris and can See the type. be regarded as one extreme in the vulgaris comples. I still prefer to treat it as a separate species, as it 20. Laboulbenia lecoareri (Balazuc) Hulden stat. nov. seems to be confined to Bembidion hasti. Laboulbenia subterranea subsp. lecoareri Balazuc 1974, in Balazuc 1973-74, vol. 43:306. Material examined T his taxon is well delimitated and it is quite safe to *U.S.S.R. Archangel Region: Kanin (Bembidion hasti treat it as a separate species. It is apparently Sahlberg). monophagous on Trechus micros (Herbst). T~e General distribution Norwegian specimens were smaller (330-470 J.Lm m Finland (Hulden 1983) length) than the French material (525 J.Lm in length according to Balazuc 1973-74) but otherwtse they Host are similar. Bembidion hasti (Coleoptera, Carabidae). Material examined *Norway. Sor-Trondelag: Kviil (Trechus micros). 18. Laboulbenia ishikawai - Fig. II Labou/benia ishikawai Sugiyama 1973:52. General distribution Material examined Federal Republic of Germany, France, Hungary, Poland, *U.S.S.R. Khabarovsk Region, Amur River (Diplous United Kingdom (Balazuc 1973-74). depressus (Gebler)). Host Trechus micros (Herbst) (Co1eptera, Carabidae). General distribution Japan (Sugiyama 1973). 21. Laboulbenia luxurians - Fig. 17 Laboulbenia luxurians Peyritsch 1873:248. Host Diplous spp. (Coleoptera, Carabidae). This species was originally described from Austria on Bembidion varium (Oliver) (Coleoptera, 19. Laboulbenia kobilae Hulden n.sp. Carabidae). Later authors have reported it from many - Figs. 15a-b regions and numerous host species (Thaxter 1896, Piceocoloratus. Matrix: Neotrechus suturalis Baumgartner 1923, Siemaszko & Siemaszko 1928, Schaufuss. Briedis 1932, Colla 1934, Banhegyi 1949, Maire 1920). Receptacle. Yellowish, with strong blackish broen My material of Labou/benia from B. dentellum pigmentaiton from the upper part of cell l upwards. (Thunberg), a species reported as host for L. /uxurians, Cells I-II forming a stalk 180-200 J.Lm in length, is different from that found on B. varium and it is evenly broadening from 20 J.Lm to 40 J.Lm in the upper described as L. marvinii n.sp. in this paper. Since it part. Cells III a nd IV equal in size, 20 X 60 J.Lm. is probable that these two Labou/benia species have Cell V small, triangular, about 15 J.Lm in height. Cell been mixed under the name L. luxurians, I think it VI triangular-isodiametric, 25- 30 J.Lm in width. is best to give an improved description of L. /uxurians Appendages. Insertion cell externally black, according to material from B. varium. appendage cells yellow. Basal cell of outer appendage Receptacle. Pale brown, cell I nearly hyaline in basal rather robust, 25-30 J.Lm in length, width at base part and with a black spot in upper part on appendage about 18 J.Lm and in upper part about 10 J.Lm, size side (not mentioned by previous authors!). Small, of the second cell about 10 X 15 J.Lm, third cell broken dark and rounded-elongated dots on cells I- VI and near the base. Basal cell of inner appendage almost base of perithecium. Cells I-II forming a stalk 80- isodiametric, 10 J.Lm in width. Inner appendage 90 J.Lm in length, broadening upwards from 15 J.Lm branching dichotomously twice, just reaching tip of to about 35 J.Lm in width. Cells I and II about equal perithecium. in length or cell II slightly shorter than cell I. Cells Karstenia 25, 1985 7

10 b

Figs. 4- ll. - 4: Cantharomyces orienta/is, a)-c) on Carpelimus corticinus, d) on C. opacus. Mature specimens from a) Ytyk-Haja (Yakut A.S.S.R.), b) Nikolsk (Yakut A.S.S.R.), c) immature specimen from Ytyk-Haja, d) base of thallus showing p osition of compound antheridium (Switzerland). - 5: Rickia wasmannii on Myrmica sabuleti (Yugosla\ ia). - 6: Stigmatomyces ptilomyiae on Atissa pygmaea (Azores), a) mature specimen, b) immature specimen, c) apex of perithecium. - 7: Stigmatomyces trianguliapicalis on Parydra coarctata (Canary Islands), a)-b) two mature specimens in different views. - 8: Laboulbenia thaxteri on Asaphidion jlavipes (Latvian S.S.R.). - 9: Laboulbenia picardii on Tachys bisulcatus (Lebanon). - 10: Laboulbenia nebriae on rujescens from a) Krasnoyarsk Region, b) United Kingdom. - II: Laboulbenia ishikawai on Dip/ous depressus (Khabarovsk Region).

HI- V forming a compact group of celts, 40- 45 J.Lm appendage irregularly triangular; branches (2-4 in length, 35 J.Lm in width. Cell III rather flat and together) arisi ng from several isodiametric cells on obliquely placed against base of perithecium, the basal celL Branches of inner appendage directed narrowly connected with celt V. Cell V nearly inwards on each side of perithecium, probably spherical, about 20 J.Lm in width. Cell VI rather flat. terminating in antheridia. Septa II- III and II- VI about equal in length. Perithecium. Pale brown-olivaceous brown, not or Appendages. Insertion cell externally very weakly only slightly darker than receptacle. A slightly darker pigmented with brown, darkest a t lower margin and shade around the perithecium below the blunt tip. near perithecium. Basal celt of outer appendage rather Perithecium slightly inwards curved. Size 30-40 X robust, pale brown, more strongly pigmented along 80- 100 J.Lm. Spores at least 30 J.Lm in length, but outer margin; second celt nearly black along outer not yet ripe; according to Thaxter (1896), they are margin. Outer appendage seems to be strongly about 50 1-Lm in length. branched from second cell upwards. Basal cell of inner Total length from base of foot to tip of perithecium 8 L. Hulden about 200-210 ,urn. Insertion cell externally narrowly darkened. Host. Bembidion varium (Olivier) (Coleoptera, Perithecium. Olivaceous brown, darker than Carabidae). The parasite grows on the pronotum and receptacle, the septa of the wall cells darker brown, el ytra of the host. brown-black spot below the tip on outer side. It is possible that L. luxurians is more restricted Perithecium straight, fairly symmetrical, tip narrow. in its choice of host species than has previously been Size about 60 X 120 /-lm. Spores 55-60 1-lm in length. realized. For the separation of L. luxurians from L. Total length from base of foot to tip of perithecium marvinii, see under the latter species. Some forms of 260- 290 1-lffi . L. pedicellata may resemble L. luxurians but differ Host. Bembidion starki Schaum and B. dentellum distinctly in the arrangement of cells III-V (see Figs. (Thunberg) ( B. jlammulatum Clairville) 23a- g). (Coleoptera, Carabidae). The parasite grows on the The specimens illustrated by Spegazzini ( 19 14, Figs. elytra of the host. 18a- b) are not L. luxurians althought occurring on Holotype. Slide L. Hulden 29, in MZH. Collecting B. varium; they possibly represent young L. vulgaris. data of the host (Bembidion dentellum): Federal Material examined Republic of Germany. Hamburg, 5. VII .1949. *Czechoslovakia. : Cenke (B. varium). Federal I have named this species in honour of my friendly Republic of Germany. Liineburg (B. varium). *U.S.S.R. guinea-pig, Marvin, who likes to gnaw my Kazachstan: Kazalinsk by Syr Darya (B. varium). illustrations. General distribution The new species differs from L. luxurians in its larger According to records of occurrences on Bembidion varium, size, the difference in size between septa 11- 111 and found at least in Algeria (Maire 1920), Austria (Peyritsch II-VI, more pointed perithecium, slightly larger 1873), Hungary (Banhegyi 1949) and Italy (Colla 1934). spores and possibly less branched appendages. It is Reports from other host species must be rechecked. probably closely related to the Holarctic L. fumosa Thaxter, occurring on Platynus spp. and Agonum spp. 22. Laboulbenia luxurians subsp. immaculata Hulden (Coleoptera, Carabidae). The latter species is more n.subsp. -Fig. 18 strongly pigmented, and has more upright appendage Sine maculis. Matrix: Bembidion semipunctatum branches and quite different host genera. (Donovan). L. fumosa was reported from Switzerland on Resembles Laboulbenia luxurians but lacks the Bembidion dentellum and B. conforme Dejean by distinct black spot in upper part of cell I. The thallus Baumgartner (1923). These records probably concern is also slightly smaller, about 135-190 1-lffi in length. L. marvinii. Many European records of L. luxurians Spores 37- 40 1-lm in length. from various Bembidion species may refer to either Host. Bembidion semipunctatum (Donovan) L. luxurians or L. marvinii. (Coleoptera, Carabidae). Material examined Holotype. Slide L. Hulden 28, in MZH. Collecting *Austria. Wildshut. (B. starki). *Federal Republic of data of the host: U.S.S.R. Leningrad Region: Germany. Hamburg (B. dentellum). *France. Seine & Oise: Molodeznoe (Vammeljoki), leg. W. Hellen. Cerney (B. dentellum). Although this morph is very similar to L. luxurians, I think it is quite safe to regard it as a separate 24. Laboulbenia nebriae subspecies, because the difference seems to be Laboulbenia nebriae Peyritsch 187 1:455. - L. alpestris constant. Specimens of L. luxurians from three remote Picard 19 13:552. - Figs. !Oa- b localities were also very uniform in habitus (see under Material examined Austria. Hohe Tauern: Gr. Glockner Pass, Hochtor, altitude that taxon). 2500 m (Nebria atrata Dejean). *Norway. Si:ir-Tri:indelag: Material examined Melhus (Nebria rufescens (Strom)). United Kingdom. *Austria. Vienna. On Bembidion semipunctatum. *U.S.S.R. Shetland. (N. mfescens). *U.S.S.R. Krasnoyarsk Region: Leningrad Region: Moloddnoe (Vammeljoki). On B. Abakan (N. rufescens). semipunctatum. General distribution Alaska (Aleutian Islands), Algeria, Austria, Canada, France, 23. Laboulbenia marvinii Hulden n.sp. Hungary, Italy, Morocco, Poland, Romania, Switzerland, - Figs. 19a-d Umted, U.S.A., Yugoslavia (Hincks 1960, Stadelmann & Ut Laboulbenia luxurians sed major. Matrix: Bembidion Poelt 1962, Balazuc 1973, 1973-74). spp. Host Receptacle. Pale brown-olivaceous brown, lighter Nebria spp. (Coleoptera, Carabidae). near the base. Small round or transversely elongated dark dots on cells II-VI. Cells 1-11 forming a stalk 25. Laboulbenia notiophili 100- 130 1-lm in length, broadening upwards from Laboulbenia notiophili Cepede & Picard 1909:259. 20 /-lffi to 45-50 ,urn. Cells III-V about equal in Material examined size. Cell V nearly spherical. Cells III and V narrowly *Norway. Hordaland: Aski:iy ( biguttatus connected. Septum 11- 111 distinctly shorter than (Fabricius)). septum II- VI. General distribution Appendage. The appendages (unfortunately more Federal Republic of Germany, Finland, France, Hungary, or less damaged) resemble those of L. luxurians, but Poland, Spain, Switzerland (Hulden 1983). seem to be Jess branched; when young the branches Host are directed inwards towards the perithecium. Notiophilus spp. (Coleoptera, Carabidae). Karstenia 25, 1985 9

Figs. 12- 19. - 12: Laboulbenia eubradycelli n.sp. a) on Bradycellus ruficol/is (Finland), b) holotype on Bradycellus caucasicus (Finland), c) spore (from holotype). - 13: Laboulbenia rougetii on Brachinus crepitans (Austria). - 14: Laboulbenia shanorii, immature specimen on Neotrechus suturalis ssp. otiosus (Yugoslavia).- 15: Laboulbenia kobilae n.;p. on Neotrechus suturali s ssp. suturalis (Yugoslavia), a) holotype, b) schematic illustration of the appendages.- 16: Laboulbenia broscosomae n.sp. on Broscosoma baldense (Italy), holotype. - 17: Laboulbenia luxurians on Bembidion varium (Czechoslovakia). - 18: Laboulbenia luxurians subsp. immaculata n. subsp. on Bembidion semipunctatum (Leningrad Region), holotype. - 19: Laboulbenia marvinii, a) and d) on Bembidion starki (Austria), b) and d) young specimens. - Scale: 40 J.lm (Fig. 12c), IOOJ.Lm (Figs. 16-19) and 200 J.lm (Figs. 12a- b, 13-15).

26. Laboulbenia paradoxa - Figs. 20a-b 27. Laboulbenia pedicellata - Figs. 23a-g Laboulbenia paradoxa Spegazzini 1915b:58. Laboulbenia pedicellata Thaxter 1892:44. The parasite occurring on the genus Tachys s. lat. and My material of this species is slightly overmature and previously interpreted by several authors as L. the tip of the perithecium is somewhat damaged. The pedicellata actually belongs to L. egens Spegazzini (see size of the spores is 5 X 62 1-Lm. The habitus of the under that species). The remaining material om appendages is very characteristic. The species was Bembidion s.lat. and Dyschirius spp. is still very described from Bembidion nitidu/um (Marsham). heterogeneous. In some cases the general Material examined morphological characters seem to be very similar, but *Austria. Karnten, on Bembidion modestum (Fabricius). considerable variation occurs in spore size. In others General distribution distinctly different morphs can be found on the same Italy (Spegazzini 1915b). host individuai.In certain cases, however, parasites Host on a particular host collected from very remote Bembidion modestum and B. nitidulum (Coleoptera, localities may be nearly identical. This has convinced Carabidac). me that L. pedicellata s.lat. consists of many tax-a_. 10 L. Hulden

Although the separate morphs are not yet safely specimen from B. inoptatum represents a very short separable, I here present characters for some diffuse form. (Figs. 23d- e). morph-groups that may help future studies on this Within the groups occur morphs which are very problem. In the list of material L. pedicellata is still uniform on certain host taxa collected from localities interpreted in a broad sense. in different regions. These possible represent separate Morph-group 1. Cells I- II forming a slender, evenly species. The morphs in group 3 seem to be more broad stalk with little pigmentation, perithecium uniform on Dyschirius species than on Bembidion rather small and narrow. Spore size 35 .urn (on species, a fact which makes it seem possible that this Bembidion quadrimaculatum) 40 .urn (B. doris) and material could be separated from the rest of L. 45- 50 .um (B. gifvipes). In typical cases the total pedicellata. More investigations are needed before length of the thallus is about 300 J.lffi, but on B. gilvipes definite conclusions can be made. I also found short specimens, which may look like Material examined members of the next morph-group, but differing in Austria. Hofgastein (Dyschirius angustatus (Ahrens)). - the evenly broad stalk. The typical L. pedicel/ala , Vienna (Bembidion octomaculatum (Goeze), B. as described by Thaxter ( 1892), belongs to this group. semipunctatum (Donovan)). - Donau-Au (B. inoptatum Schaum). Egypt. Luxor (D. lafertei Putzeys). Federal Republic (Fig. 23f, see also Fig.77d in Hulden 1983). of Germany. Hamburg (B. guttula (Fabricius), D. thoracicus Morph-group 2. Cells I-II forming an only slightly (Rossi), D. obscurus (Gyllenhal)). - Einbech (B. biguttatum pigmented stalk, which broadens strongly upwards, (Fabricius)). - Sylt (B. aeneum Germar).- Ellwangen (B. perithecium short and broad appearing swollen), lunulatum (Fourcroy)). - Zeit berg bei Li.ineburg (B. varium septum ll-III distinctly shorter than septum II­ (Olivier)). France. Gascogne (B. articulatum (Panzer)). - VI, spores about 35-40 .um in length, total length Allier: Vichy (B. articulatum).- 'LaChapelle' (B. azurescens of thallus at most about 170 .urn. Specimens from (Dalla Torre)). Greece. Elevsis (B. irico/or Bedel). - Crete: Bembidion quadrimaculatum (Finland and Siberia) Canea (B. lunulatum). *Ireland. Kerry: Killarney (B. harpaloides Audinet-Serville). Netherlands. Amsterdam (B. were 150-170 .urn in length, and from B. azurescens aeneum). *Sweden. Sk

fOb

100 ~m

23 f

23 d

Figs. 20-23. - 20: Laboulbenia paradoxa, a) mature specimen with slightly broken perithecium, b) spore.- 21: Laboulbenia vulgaris on Pelophila borealis (Krasnoyarsk Region). - 22: Laboulbenia egens, a) on Tachys haemorrhoidalis (Canary Islands), b) on Tachys fumigatus spp. socius (Egypt). - 23: Laboulbenia pedicel/ata s.lat. on a) Bembidion aeneum (Sweden), b) B. aeneum (Federal Republic of Germany), c) B. iricolor (Greece), d) B. inoptatum (Austria), e) B. guliula (Fed eral Republic of Germany), f) and g) B. quadrimaculatum (Irkutsk Region).

Host Host Tachys bisulcatus (Coleoptera, Carabidae). Calathus, Acupalpus and some other genera (Coleoptera, Carabidae). 30. Laboulbenia polyphaga - Fig. 3 (map) Laboulbenia polyphaga Thaxter 1893:165. 31. Laboulbenia pseudoroasei Laboulbenia pseudomasei Thaxter 1899: 196 Material examined Austria. Obertilliach (Cala thus melanocephalus (Linnaeus)). Material examined *Faeroes (Denmark). Vagur: Sorvagur (C. fuscipes (Goeze)). *Bulgaria. Rila (Pterostichus strenuus (Panzer)). France. *Norway. More og Romsdal: Molde (C. micropterus f.yrenees: Porte Puymorens (P. strenuus). *Sweden. Skane: Duftschmid). Switzerland. Munstertal, Santa Maria (C. Angelholm (P. strenuus). - Gotland: Klinte (P. strenuus). melanocephalus). U.S.S.R. *Archangel Region: Mesen (C. U.S.S.R. *Archangel Region: Mesen (P. brevicornis (Kirby)). melanocephalus). - *Caucasus (C. melanocephalus). - *Caucasus (P. strenuus). - *Yakut A.S.S.R.: Bardeljak (P. ochoticus Sahlberg). Agrafena (P. ochoticus). Shigansk General distribution (P. ochoticus var. mutator Poppius). Tungus-Haja (P. Cosmopolitan, in Europe Known from Austria, Belgium, brevicornis). - 'Sibir. or.' (P. empetricola Dejean). the Federal Republic of Germany, Finland, France, General distribution Hungary, Italy, the Netherlands, Poland, Portugal, Federal Republic of Germany, Finland, France, Hungary, Switzerland, the U.S.S.R. (Leningrad Region, Latvian Italy, Mongolia, Poland, Switzerland, U.S.S.R. (Karelia.J S.S.R.) (Hulden 1983). A.S.S.R.) (Hulden 1983). 12 L. Hulden

Host mountains (B. tibiale (Duftschmid). B. millerianum, B. eques Pterostichus spp. and Anisodacty/us spp. (Coleoptera, ssp. combustum Menetries). Canary Islands (Spain). Gran Carabidae). Canaria (B. subca!!osum Wollaston). - Gomera (B. atlanticum ssp. phobon Netolitzky). - Tenerife (B. fortunatum Wollaston, B. at/anticum ssp. phobon). 32. Laboulbenia rougetii - Fig. 13 Czechoslovakia. Bohemia: Pardubice (B. fluviatile Dejean). Laboulbenia rougetii Montagne & Robin in Robin Without locality (B. ascendens Daniel, B. nitidulum 1853:622. (Marsham)). - Moravia: Paskau (B. tricolor). - Slovakia: Tatra, Banska Bystrica (B. difflcile ssp. haeneli Netolitzky). Material examined Fatra, Krivar\ (Duvalius microphthalmus ssp. microphthalmus Austria. Neusiedler See (Brachinus crepitans (Linnaeus)). (Miller)). Silica (D. microphthalmus ssp. hungaricus Csiki). General distribution Zadiel dol. (B. subcostatum ss p.javurcovae Fassati). *Faeroes Holarctic, Hong Kong (Stadel mann & Poelt I 962, Sugiyama (Denmark). Suduroy: Nes (T obtusus Erichsen). Federal 1973, Balazuc 1973-74). Republic of Germany. Hamburg (B. tetracolum ssp. Host tetracolum Say, B. punnctulatum ssp. punctulatum Drapiez, Brachinus, Chlaenius and some other related genera B. bipunctatum (Linnaeus)). Hanau, Main (B. tetracolum ssp. tetracolum). Marburg (B. fluviatile Dejean). Einbech (B. (Coleoptera, Carabidae). biguttatum (F.tbricius)). Baden-Wi.i rtenberg, Ulm (B. schueppeli Dejean). Bonn (B. tetraco/um ssp. tetracolum). 33. Laboulbenia shanorii Bayrischen AI pen (B. conjorme Dejean). France. (B. Laboulbenia shanorii Banhegyi 1960:54. kuesteri Schaum). Corsica , Vizzarona, 1200 m (B. - Fig. 14 (immature specimen) geniculatum ssp. dilutipes Deville). Corsica, Ajaccio (B. Material examined obtusus ssp. tethys Netolitzky). Allier, A brest (B. punctulatum Yugoslavia. Montenegro: Krivosije (Neotrechus suturalis ssp. ssp. punctulatum). Allier, Vichy (B. coeruleum Audinet­ otiosus Obenb.). Serville, B. azurescens H. Wagner). Paris (B. coeruleum). Strasbourg (B. prasinum (Duftschmid)). Beaugency (B. General distribution punctulatum ssp. punctulatum). Polminhac (B. punctulatum Albania, Yugosla\ia (Banhegyi 1960). ssp. punctulatum). Gennevil/iers, Seine (B. tetracolu m ssp. tetracolum). 'Gallia' (B. genei ssp. illigeri Netolitzky). Host •German Democratic Republic. Leipzig (B. biguttatum). Neotrechus spp. (Coleoptera, Carabidae). *G reece. Corfu (B. nigropiceum (Marsham)). Lesbos (B. punctulatum ssp. punctulatum). Nauplia (B. tetraco/um ssp. tetracolum ). *Jreland. Kerry: Killarney (B. tetracolum ssp. 34. Laboulbenia thaxteri - Fig. 8 tetracolum, B. harpaloides). - Cork: Mallow (B. tetracolum Laboulbenia thaxteri Cepi:de & Picard 1908:260. ssp. tetracolum). Italy. (T. rufulus Dejean). Sardinia, Tempio (B. elongatum ssp. elongatum Dejean). Mte Baldo Material examined (T. longobardus, T. sinuatus ssp. sinuatus Schaum). Bergamo, *Austria. Graz (A. flavipes). *U.S.S.R. Latvian S.S. R.: Sigulda (Asaphidionflavipes (Linnaeus)). Pellegrino (B. ascendens) . Val Giudica (T. longulus ssp. danie/i Holdhaus). Bolzano, Tires (B. tricolor) . Mte Maggiore General distribution (Anophthalmus schmidti ss p. flachi Winkler). Val Camonica, Algeria (Maire 1920), France (Cepede & Picard 1908), Greece Edolo (B. geniculatum Heer). Val Camonica, Oglia (B. (Balazuc 1973-74), Italy (Colla 1934), Poland (Siemaszko tricolor). Cuneo, Crissolo (B. longipes). Genova, Ronco & Siemaszko 1928), Switzerland (Baumgartner 1951). Scrivia (B. ascendens). •Madeira (Portugal). Paul da Serra (B. obtusum ssp. tethys). Porto Santo (B. atlanticum ssp. Host atlanticum). *Morocco. Haute Atlas: Arround (B. Asaphidion spp. (Coleoptera, Carabidae). atrocoeruleum Stephens, B. sicu/um ssp. siculum Dejean, B. coeruleum. B. da/matinum ssp. dudichi Csiki). Amizmiz (B. coeruleum). Reraia (B. coeru/eum). Netherlands. Rotterdam 35. Laboulbenia vulgaris - Fig. 21 (B. tetracolum ssp. tetracolum). •Norway. Hordaland: Laboulbenia vulgaris Peyritsch 1873:248. Gjerstad (B. bruxellense). *Palestine. Galilee (B. tetracolum The illustrated specimen from Pelophila borealis is ssp. tetracolum). Poland. Koszalin (B. biguttatum). a morph with exceptional appendages. It very Kroscienko (B. lomnickii Netolitzky). Romania. Transylvania p robably belongs to the L. vulgaris complex. (Duvalius procerus Putzeys). Spain. Madrid. (B. paulinoi Heyden). Sierra Nevada (B. atrocoeruleum). *Sweden. Skane: Material examined Arlov (B. tetracolum ssp. tetracolum). Ven (B. stephensi *Albania. Korab (Bembidion balcanicum Apfel beck). Algeria. Crotch). - Vastergotland: Alingsas (B. mannerheimi Constantine (B. quadripustu!atum Audinet-Serville). Austria. Sahlberg). Switzerland. Mi.instertal (B. tricolor, B. conforme. Salzburg: Bad Gastein (B. stomoides Dejean). Schafberg B. complanatum Heer, B. ruflcorne Sturm, B. andrea ssp. (Trechus wagncri Ganglbauer). - Tirol: Reutte (B. longipes bualei Duval). Unter-Engadin, Zernez (B. andreae ssp. bualei, Daniel). Lienz (B. longipes). Without locality (B. tricolor B. geniculatum). Chiasso, Tessin (B. coeruleum). Solothurn, (Fabricius)). - Karnten: Eisenkappel (B. millerianum Dornach (B. ascendens. B. andreae ssp. bualei, B. decorum Heyden). Vorarlberg: Riezlern (B. ruflcorne Sturm).-Ober­ ssp. decorum, B. punctulatum ssp. punctulatum ). Wallis, bsterreich: Wildshut (B. stark!. Schaum). Ostermiething (T. Gletsch Furka (B. ruficorne). Uri, Schachen-Tal (B. discus (Fabricius)). - Nieder-Osterreich: Raxalpe (T. ovatus millerianum Heyden). Uri, Oberalppass (B. incognitum Putzeys). - Vienna (B. seminpunctatum (Donovan), B. ~li.iller). Wallis, Bivuntal (B. complanatum). No locality (B. lunatum (Duftschmid)).- Steiermark: Ad mont (B. conforme complanatum). *Syria. Sanamein (B. quadripus tulatum Dejean, B. testaceum ssp. testaceum (Duftschmid), B. tricolor Audinet-Serville, B. tetracolum ssp. tetracolum). Turkey. Fabricius). Schneealm (T. ovatus). Graz (B. schueppeli Ephesos (B. praeustum ). Hermes River (B. tetracolum ssp. Dejean). Dackstein , Gutenberghaus (T. glacialis Heer). tetracolum, B. moschatum). Smyrna (B. dalmatinum ssp. Gesause (B. ruflcorne. T. ovatus). - Hohe Tauern: Hallbri.inn dalmatinum). Tschakit-Thal (B. andreae ssp. caucasicola (T. hampei Ganglbauer). Salesenwald (T. glacialis). *Azores. Netolitzky). Baba Dagh (B. cordicol/e Duval, B. praeustum). Flores (B. harpaloides Audinet- Serville). Bulgaria. Rila United Kingdom. Scotland, Dumfricsshire (B. decorum ssp. Karstenia 25, 1985 13 decorum). Shetland (T. obw sus). Orkney (B. tibiale General distribution (Duftschmid), B. tetraco/um ssp. tetracolum). U.S.S.R. Finland, Haiti, Jamaica, Poland, U.S.S.R. (Leningrad *Archangel Region: Mesen (B. guttula). Pjosa (B. andreae Region, Karelian A.S.S.R., Murmansk Region) (Hulden ssp. polonicum Muller, B. semipunctatum (Donovan)). - 1983). *Armenia: Zsachkadzor (B. testaceum ssp. parallelipenne Chaudoir, B. kei/bachi Muller, B. tibia/e, B. geniculatum Host Heer). - *Azerbayjan: Lenkoran (B. andreae ssp. Oxytelus laqueatus (Coleoptera, Staphylinidae) in Europe quadriflammeum Reitter). - *Turkmenistan: Ashkhabad (B. {Huldi:n 1983). tetracolum ssp. tetracolum). - *Kirgizia: Dschil-Aryk (B. giganteum Sahlberg, B. insidiosum Solsky). - *Tomsk 38. Monoicomyces sanctae-helenae Region: Schukova (B. tetracolum ssp. tetraco/um). - Monoicomyces sanctae-helenae Thaxter 1900:413. *Krasnoyarsk Region: Dudinka (Pelophila borealis Paykull). The present material represents the typical morph (cf. Osnatschennaja (B. petrosum ssp. petrosum Gebler). - *Yakut A.S.S.R.: Olekminsk (B. hirmocaelum Chaudoir). Huldlm 1983). Yakutsk (B. bruxellense Wesmael). Ytyk-Haja (B. poppii Material examined Netolitzky). Shigalovo (B. hirmocae/um). Shigansk (B. U.S.S.R. Southern Urals, locality unknown (Oxytelus piceus bruxel/ense. B. obscurellum (Motschulsky)). Batylim (Linnaeus)). - *Yakut A.S.S.R.: Olekminsk (0. piceus). (Trichocellus mannerheimi ssp. mannerheimi (F. Sahlberg)). Bulun (B. difforme Motschulsky). Yugoslavia. Carniolia (B. General distribution elongatum ssp. tarsicum Peyron). Carniolia, Cerna prst Algeria, Cameroon, Finland, 'Germany', Italy, Malaysia (Anophthalmus bohimiensis Ganglbauer). Dalmatia (B. (Sarawak), Poland, St. Helena, Taiwan, U.S.S.R. (Leningrad dalmatinum ssp. dalmatinum Dejean). Montenegro, Uruja Region, No\'osibirsk Region) (Huldi:n 1983). (B. tricolor). Bosnia-Herzegovina (B. ruficorne). Bosnia­ Herzegovina, Vran Plan ina (B. reiseri ssp. reiseri Apfel beck, Host B. reiseri ssp. vranense Apfelbeck). Oxytelus spp. (Coleoptera, Staphylinidae). 39. Peyritschiella protea General distribution Cosmopolitan (Hulden 1983). Peyritschiella protea Thaxter 1900:427. All records on Anotylus rugosus (Fabricius). Host Material examined Bembidion (s.lat.) and some related genera (Coleoptera, *Austria. Marburg. *Czechoslovakia. Paskau. France. Seine Carabidae). & Oise: Cerney. German Democratic Republic. Lubars. *Ireland. Kerry: Killarney. Romania. T ransylvania. *Sweden. 36. Misgomyces dyschirii Skane: Lomma. K~ngsmarken . Bjared. - Halland: Saro. Misgomyces dyschirii Thaxter 1900:443. - Vastergotland: Orby. Skovde. United Kingdom. Surrey. Material examined - NW Wales. -Shetland. U.S.S.R. *Archangel Region: • Austria. Neusiedler See (Dyschirius extensus Putzeys, D. Archangel. pusillus (Dejean)). *Canary Islands (Spain). Gran Canaria: General distribution Pt. Maspalomas (D. pusillus). *Denmark. Sjalland: Algeria, Federal Republic of Germany, Finland, France, Saltholmen (D. salinus Schaum). France. Allier: Vichy (D. German Democratic Republic, Hungary, Netherlands, nitidus (Dejean)). 'Germania'. No locality, on D. intermedius Poland, Romania, United Kingdom, U.S.A., U.S.S.R. Putzeys. *Greece. Corfu (D. salinus). *Iran. Lotfabad (D. (Leningrad Region, Karelian A.S.S.R.) (Balazuc 1973, politus (Dejean)). *Ireland. Kerry: Killarney (D. globosus Hulden 1983). (Herbst)). Italy. Ancona reg.: Ancona (D. digitatus (Dejean)). Host - Basilicata reg.: San Basilio (D. /aeviusculus Putzeys). - Anotylus spp., Oxy telus spp., and some related genera Lago Trasimeno (D. a/pica/a Ganglbauer). *Palestine. Kison, (Coleoptera, Staphylinidae). near Haifa (D. rufoaeneus Chaudoir).- Jordan, near Jericho (D. rufoaeneus). Spain. Algecir<~s (D. numidicus Putzeys). U.S.S.R. Leningrad (D. globosus). - *Estonian S.S.R.: 40. Rhachomyces aphaenopsis Haapsalu r., 'Nucko' (D. globosus). Tartu r., Tartu (D. Rhachomyces aphaenopsis Thaxter 1905:314. globosus). - *Latvian S.S.R.: Sigulda (D. globosus). Jugla Material examined (D. g/obosus).- *: Odessa (D. substriatus ssp. priscus France. Ariege: Grotte Daubert (Aphaenops cerberus Muller). Bogdo (D. nitidus). -*Turkmenistan: Michailovsk (Dieck)). (D. sa/inus). Tedzhen (D. nitidus). - *Buryatian A.S.S.R.: Ubukunsk (D. globosus). General distribution France (Balazuc 1970), Spain (Balazuc et al. 1983). General distribution Algeria, Federal Republic of Germany, Finland, France, Host Hungary, Italy, Japan, Netherlands, Poland, Romania, Aphaenops spp. Coleoptera, Carabidae). Spain , Switzerland, United Kingdom, U.S.A., U.S.S.R. (Leningrad Region) (Bechet & Bechet 1960, Balazuc et al. 1983 , Hulden 1983). 41. Rhachomyces furcatus Host Acanthomyces furcatus Thaxter 1893: 177. Dyschirius spp. (Coleoptera, Carabidae). Rhachomyces furcatus (Thaxter) Thaxter 1895:468. Material examined 37. Monoicomyces furcatus Austria. Nieder-Ostcrreich: Alps ( Othius brevipenn is Kraatz). Monoicomycesfurcatus Thaxter 1931:41. - Steiermark: Graz (0. brevipennis). *German Democratic Material examined Republic. Freienwalde ( 0 . angustus Stephens). *Hungary. No *Czechoslovakia. Bohemia ( Oxytelus laqueatus (Marsham)). locality indicated (0. crassus Motschulsky). *Sweden. Skane: *Norway. Dovre: Fokstua (0. laqueatus) . *Sweden. Oland: Balsbergsgrottan ( 0. punctulatus (Goeze)). Yugoslavia. Vickleby ( 0 . laqueatus). Bosnia & Herzegovina: Mostar (0. lapidicola Kiesenwetter). 14 L. Hulden

General distribution General distribution Algeria, Austria, 'Equatorial Africa', Federal Republic of Algeria, ?Hungary, ?Yugoslavia (Balazuc 1970). Germany, Fmland, France, Poland, United Kingdom, U.S.S.R. (Kareltan A.S.S.R.), ?Yugoslavia (Frank 1982 Host Hulden 1983). ' Duvalius spp. (Coleoptera, Carabidae). 46. Rhachomyces philonthinus Host Othius spp. (Coleoptera, Staphylinidae). Rhachomyces philonthinus Thaxter 1900:435. Material examined *Austria. Ost-Tirol: in the vicinity of Lienz altitude 1800m 42. Rhachomyces hypogaeus and 2100m (Philonthus varians (Paykull)). ' Acanthomyces hypogaeus Thaxter 1893:177. Rhachomyces hypogaeus (Thaxter) Thaxter 1895:467. General distribution Material examined Largely distributed in Europe, Africa, Asia and North Y~goslavia. 'Gradah Hohle' (Typhlotrechus bilimeki ssp. Amenca (Balazuc, 1973-74). In Europe recorded from billmek1 (Sturm)). -. Without locality information, Belgium, the Federal Republic of Germany, France, on T btlimek1 ssp. likanensis Schauf. 'Silna Radino pee' Hungary, Italy, the Netherlands, Poland Switzerland and (T. bilimeki ssp. kiesenwetteri Schmidt). the United Kingdom (Collart I 945, Hincks,l960, Stadel mann & Poelt 1962, Balazuc 1973-74, Rossi 1975). General distribution Austria, Yugoslavia (Thaxter 1896, Banhegyi 1960). Host Philonthus spp. (Coleoptera, Staphylinidae). Host Typhlotrechus bilimeki sspp. (Coleoptera, Carabidae). 47. Rhachomyces stipitatus 43. Rhachomyces hypogaeus subsp. richardii Rhachomyces stipitatus Thaxter 1900:438. Rhachomyces hypogaeus subsp. richardii Balazuc Rhachomyces stipitatus var. pallidus Maire 1912:196. 1973(42):256 in Balazuc 1973-74. Illustrated in - Rhachomyces capucinus Thaxter 1931:276. Balazu 1970. Synonymy according to Rossi (1978). Described on Speotrechus mayeti (Ab.) (+ sspp.) from Material examined France. Tarn-et-Garonne: Grotte de Capucin (Duvalius France. lespesi Fairmaire). Material examined *Italy. Trento: 'Monte Pasi', on Speotrechus knauthi General distribution Gangl bauer. Algeria, France, Greece, Italy (Balazuc 1970, Rossi 1978).

General distribution Host France (Balazuc 1970). Duvalius spp. (Coleoptera, Carabidae). Host 48. Rhachomyces tenenbaumii Speotrechus spp. (Coleoptera, Carabidae). Rhachomyces tenenbaumii Siemaszko & Siemaszko 44. Rhachomyces maublancii 1928:205. Rhachomyces maublancii Lepesme 1942:73 (nomen The . species was described on Thalassophilus nudum). - Rhachomyces maublancii Lepesme ex W. longzcorms (Sturm) from Poland. Both T. longicornis and T w_hzlet Wollaston are free-living Rossi 1978:329. occurnng m motst habitats. The former is largely Material examined Italy. Cima Tombea (Duvalius baldensis ssp. tombeanus dtstnbuted 10 Europe and the latter restricted to the Ganglbauer). Bergamo: Monte Alben (Duva!ius winkleri Canar~ Islands. A t~ird SJ?ecies, T. breuili (Jeanne!), Jeanne!). ts confmed to c~ves 10 Spam. The tdenttty of possible parasttes on thts spectes would be of great interest· General distribution so far none has been found. ' Italy (Rossi 1978), Hungary (Banhcgyi 1949). Material examined *Canary Islands (Spain). La Palma: Caldera (Thalassophilus Host while! Wollaston). - Gomera: El Cedro (T. whilei). Duvalius spp. (Coleoptera, Carabidae). General distribution Austria, France, Poland (Banhegyi 1964). 45. Rhachomyces ?peyerimhoffii Rhachomyces peyerimhoffii Maire 1912:197. Host Described from Algeria on Duva!ius lapiei Thalassophilus spp. (Coleoptera, Carabidae). (Peyenmhoff). Balazuc ( 1970) questioned the European records from Hungary and Yugoslavia (cf. 49. Rhachomyces vayssierei Rhachomyces vayssierei Lepesme 1942:65. Banhegyi 1960). My material consists of only one Material examined specimen, which lacks the perithecium. It resembles *Austria. Tirol: Hohe Tauern, Gasteiner Tal (Trechus the illustration of R. peyerimhoffii presented by quadristriatus (Schrank)). *German Democratic Republic. Lepesme (1942, Fig. IV:!). Total length of the Leipzig (Trechus obtusus Erichson). *Sweden. Oland: Hogby receptacle 255 J.Lm . (T. quadristriatus). Material examined *Italy. Brescia: Val Camonica, Monte Glisento (Duvalius General distribution longhii Comolli). Bulgaria, France, Hungary (Banhegyi 1960). Karstcnia 25, 1985 15

Host Finland, Hungary, Italy, Netherlands, Portugal, Spain, Trechus spp. (Coleoptera, Carabidae). United Kingdom, U.S.A. , U.S.S.R. (Leningrad Region, Murmansk Region), Yugoslavia (Hulden 1983). 50. Rickia hyperborea Rickia hyperborea Balazuc 1980:216. Host The find of Rickia hyperborea in T it-Ary in Yakutia Quedius spp. (Coleoptera, Staphylinidae). is the northernmost record of Laboulbeniales Acknowledgements. I am grateful to the following persons presently known. The species probably occurs all over for providing material or information essential ot this study: the range of .\1.icralymma marinum and its closest Prof. Harald Haberman (Tartu, Estonian S.S.R.), Dr. Lita relatives. Greve. Jensen (Bergen, Norway), Prof. Aloi s Kofler (Lienz, Material examined Austna), Mr. Bjorn Erling Waage (Bergen, Norway), Dr. Norway. Tra ms: Kvesmenes (Micralymma marinum Strom). Dagfinn Refseth (Trondheim, Norway) and Mr. Seppo Maalselv (M. marinum). Hillesoya (M. marinum). U.S.S.R. Koponen (Turku, Finland). Mr. Olav Aas (Bergen, Norway), *Yakut A.S.S. R.: Tit-Ary, Lena River, about 72°N (.'vi. who has also investigated Laboulbeniales on the insects brevilingue ssp. dicksoni Maklin). collected by Bjorn Erling Waage, kindly gave me permission to publish the results in this connection. General distribution France, Norway, U.S.S.R. (Murmansk Region, Magadan Region) (Hulden 1983). References Host Balazuc, J. 1970: Commentaires sur les Rhachomyces .'vficralymma spp. (Coleoptera, Staphylinidae). parasites des Trechidae, avec descri ptions de formes nouvelles (Ascomy~etes , Laboulbeniales). - Ann. Soc. 51. Rickia wasmannii -Fig. 5 Ent. Fr. (N.S.) 6(3). 677-699. 1971: Notes sur les Laboulbeniales 3. Rectifications, Rickia wasmannii Cavara 1899: 173. synonymies et mises au point. - Bull. Soc. Linn. Lyon Material examined 40: 211 -216. *Yugoslavia. : Podcetrtek (Myrmica sabuleti 1973: Ebauche d'une flare des Laboulbeniales de Meinert). Roumanie (Ascomycetes). - Livre du Cinquantenaire de l'lnstitut de Speleologie 'Emile Racovitza , Bucarest: General distribution 463-477. Austria, Federal Republic of Germany, France, Italy, 1973-74: Laboulbcniales de France.- Bull. Soc. Linn. Luxemburg, Spain, Switzerland, Uni ted Kingdom (Scotland) Lyon 42: 244-256,280-285,43: 12-21,57-64,73- (Balazuc 1973- 74, Balazuc et al. 1982). 79,253--262,295,--315, 346--368. Host 1978: Laboulbeniales (Ascomycetes) de Ia region Myrmica spp. (Hymenoptera, Formicidae). FranfYaise Antilles-Guyane. - Bull. Soc. Linn. Lyon 47 : 488-500. 1980: Laboulbeniales nouvelles (Ascomycetes), parasites 52. St igmatomyces ptilomyiae - Fig~. 6a-c de Coleopteres et de Di£teres. - Bull. Mus. Nat. Hist. Stigmatomyces ptifomyiae T haxter 1931: 154. Nat. 4•ser. (section B) 2(2): 209-219. 1982: Laboulbeniales (Ascomycetes) de Madagascar, des Material examined Comores et des Mascareignes. - Bull . Soc. Linn. Lyon *Azores (Portugal). Fayal: Horta (Atissa pygmaea Haliday). 51:6-27. - Terceira: Angra do Heroismo (A. pygmaea). Balazuc, J., Espadaler, X. & Girbal, J. 1983: Laboulbenials General distribution (Ascomiccts) Iberiques 2. Naves aportacions.- Collnea Grenada, Italy, Jamaica (Thaxter 1931, Rossi & Cesari Rossi Bot. 14: 39- 42. 1979). Banhegyi, J . 1949: Les Laboulbeniales de Ia Transylvanie. - Index Horti Bot. Univ. Budapest 7: 93- 101. Host 1960: Contributions it Ia connaissance des Laboulbeniales Atissa (Ptilomyia) spp. (Diptera, Ephydridae). de Ia pen ins~le des Balkans.- Ann. Univ. Sci. Budapest. Sect. Bwl. 3. 49- 67. 1964: Notes sur quelques Laboulbeniales de Ia Pologne. 53. Stigmatomyces trianguliapicalis - Figs. 7a-b -Ann. Univ. Sci. Budapest. Rolando Eotvos nomin., Stigmatomyces trianguliapicalis Majewski 1972:234. Sect. Bioi. 7: 19-27. Baumgartner, R. 1923: Contribution it !'etude des Material examined Laboulbeniales de Ia Suisse. -Jahrb. Philosoph. •Canary Islands (Spain). Gran Canaria: Los Lagunetas Fakultat II Univ. Bern 3: 257-265. (Parydra coarctata (Haliday)). - 195 1: Resultats de mes dernieres recherches sur les Laboulbeniales (Champignons sur insectes vivants). - General distribution Mitt. Naturforsch. Ges. Bern. Neue Folge 8: xxx-xxxiii. Italy, Poland (Majewski 1972, Ross i & Cesari Rossi 1979). Bechet, M. & Bechet, I. 1960: Ciuperci parazite ~i sa{>rofite Host pe i ~ secte ( l ). - Studia Universitatis B abe~ - Bolya 1. (Ser. Parydra spp. (Diptera, Ephydridae). 2) 2. 95-103. Blackwell, M. 1980a: Incidence, host specificity, distribution and morphological variation in Arthrorhynchus 54. Symplectromyces vulgaris nycteribiae and A. eucampsipodae Teratomyces vulgaris Thaxter 1900:431. (). - Mycologia 72: 141- 158. Symplectromyces vulgaris (Thaxter) T haxter 1908:3 15. 1980b: Developmental morphology and taxonomic characters of Arthrorhynchus nycteribiae and A. Material examined eucampsipodae (Laboulbeniomycetes).- Mycologia 72: •Sweden. Skane: Balsbergsgrottan ( Quedius mesomelinus 159-168. (Marsham)). Tykarpsgrottan (Q. mesomelinus). Cavara, F. 1899: Di una nuova Laboulbeniacea: Rickia Wasmannii , nov.gen. e nov.spec. - Malpighia 13: 173- General distribution 188. Cepede, C. & Picard, F. 1908: Observations biologiques sur Belgium, Bengal, Canada, Federal Republic of Germany, les Laboulbeniacees et d iagnoses sommaires de quelques 16 L. Hulden

especes nouvelles. - Comptes Rendus Assoc. Fran9. - 1918: Observaciones microbiol6gicas. - An. Soc. Cient. Avancement Sci, 36°sess., Reims 1907: 778-784. Argent., Buenos Aires 85: 311-323. 1909: Contribution a Ia biologic et a Ia systematique Stadelmann, M. & Poelt, J. 1962: Zur Kenntnis der des Laboulbeniacees de Ia flore fran9aise. - Bull. Sci. mittele~ropaischen Laboulbeniales. - Ber. Bayer. Bot. Fr. Belg. 42: 247-268. Ges. 35. 120-132. Colla, S. 1934: Laboulbeniales. Flora Italica Sugiyama, K. 1973: Species and genera of the Laboulbeniales Cryptogamica I (Fungi 16): 1-157. (Ascomycetes) in Japan.- Ginkgoana 2: 1-97. Collart, A. 1945: A propos des Laboulbeniacees. - Les - 1978: The Laboulbeniomycetes of eastern Asia (2). On Naturalistes Belges 26: 98-103. eight species from Japan and Formosa including two Frank, J.H. 1982~ The parasites of the Staphylinidae new species of Rickia.-J. Japan. Bot. 53: 154-160. (Coleopt~ra). -:- Techn. Bull. Agric. Exp. St. Inst. Food Thaxter, R. 1891: Supplementary note on North American Agnc. Sc1. Umv. Fla. 824. 1-118. . - Proc. Amer. Acad. Arts & Sci. 25: Hincks, W.D. 1960: Notes on the Laboulbeniales. - The 261 - 270. Naturalist 1960: 97- 102. 1892: Further additions to the North American species Hulden, L. 1983: Laboulbeniales (Ascomycetes) of Finland of Laboulbeniaceae. - Proc. Amer. Acad. Arts & Sci. and adjacent parts of the U.S.S.R.- Karstenia 21: 31- 27: 29- 45. 136. 1893: New species of Laboulbeniaceae from various Lepesme, P. 1942: Revision des Rhachomyces palearctiques localities. - Proc. Amer. Acad. Arts & Sci. 28: 156- (Laboulbeniaceae). - Bull. Soc. Mycol. Fr. 58: 57- 188. 80. 1895: Notes on Laboulbeniaceae, with descriptions of Maire, R. 1912: Contribution a !'etude des Laboulbeniales new species. - Proc. Amer. Acad. Arts & Sci. 30: 467- de !'Afrique du Nord. - Bull. Soc. d'Hist. Nat. Afr. 481. Nord 4(9): 194- 199. 1896: Contribution towards a monograph of the 1916: Deuxieme contribution a !'etude des Laboulbeniaceae.- Mem. Amer. Acad. Arts & Sci. 12: L~b~ulbeniales de I'Af~ique du Nord. - Bull. Soc. 187-429. d Htst. Nat. Afr. Nord 7. 6-39. 1899: Preliminary diagnoses of new species of 1920: Troisieme contribution a !'etude des Laboulbeniaceae 1. - Proc. Amer. Acad. Arts & Sci. Laboulbeniales de !'Afrique du Nord. - Bull. Soc. 35: 153-209. d'Hist. Nat. Afr. Nord 41: 123-138, 143-158, 159- 1900: Preliminary diagnoses of new species of 170. Laboulbeniaceae 2. - Proc. Amer. Acad. Arts & Sci. Majewski, T. 1972: Rare and new Laboulbeniales from 35: 407-450. Poland 2.- Acta Mycol. 8: 229-237. 1901: Preliminary diagnoses of new species of - 1973: Rare and new Laboulbeniales from Poland 3. - Laboulbeniaceae 3. - Proc. Amer. Acad. Arts & Sci. Acta Mycol. 9: 111-124. 36: 395-414. Middelhoek, A. 1949: Laboulbenaiaceae in Nederland 3. - 1905: Preliminary diagnoses of new species of Nederl. Kruidk. Arch. 56: 249-260. Laboulbeniaceae 6. - Proc. Amer. Acad. Arts & Sci. Peyritsch, J. 1871: Ober einige Pilze aus der Familie dcr 41 :301-318. Laboulbenien. - Sitz. Ber. Kaiser!. Akad. Wiss. Math.­ 1908: Contribution towards a monograph of the Naturwiss. Klasse 64(1): 441-458. Laboulbeniaceae 2. - Mem. Amer. Acad. Arts & Sci. - 1873: Beitrage zur Kenntnis der Laboulbenien. - Sitz. 13: 217-469. Ber. Kaiser!. Akad. Wiss. Math.-Naturwiss. Klasse 68(1): 1931 : Contribution towards a monograph of the 227-254. Laboulbeniaceae 5. - Mem. Amer. Acad. Arts & Sci. Picard, F. 1913: Contribution a !'etude des Laboulbeniacees 16: 1-435. d'Europe et du nord de !'Afrique. - Bull. Soc. Mycol. Accepted for publication France 29: 503-571. Robin, C. 1853: Histoire naturelle des vegetaux parasites on September 15, 1984 qui croissent sur l'homme et sur les animaux vivants. - 704 pp. Paris. Errata Rossi, W. 1975: Su alcune Laboulbeniali (Ascomycetes) In 'Laboulbeniales (Ascomycetes) ofFinland and adjacent parts nuove per l'Italia.-Giorn. Bot. Ita!. 109: 71-85. of the U.S.S.R.', Karstenia 21: 31-136, there were some - 1978: Sui Rhachomyces (Ascomycetes, Laboulbeniales) disturbing errors and misprints, which should be corrected: parassiti dei Duvahus italiani (Coleoptera, Carabidae, page 48 left column line 49: for Figs. 30 read Figs. 40 Trechini).- Int. J. Speleol. 10: 323-330. page 55 left column line 26: for Agonum viduum (Panzer) Rossi, W. & Cesari Rossi, G. 1979: Su alcune species di read Agonum moestum Duftschmid. The same change Stigmatomyces (Ascomycetes, Laboulbeniales) parassite di Ditteri italiani. -Boll. Mus. Civ. Venezia 30: 13- should be made for Figs. 101 a-d page 109. This correction 18. does not affect the statistics for A. viduum (pp. 123 and Ryberg, 0. 1947: Studies on bats and bat parasites, especially 131), but A. moestum should be added to the host-parasite with regard to Sweden and other neighbouring countries list on p. 131 and to the parasite-host list on p. 135. of the North. - 330 pp. Stockholm. .. page 56 right column line 24: for Fig. 88b read 98b Scheloske, H.-W. 1969: Bearage zur Biologic, Okologie und page 59 left column line 62: for Fig. 8lb read 9lb Systematik der Laboulbeniales (Ascomycetes) unter page 59 right column line 1: for Fig. 8la read 91a besonderer Beriicksichtigung des Parastt-Wirt­ page 61 and elsewhere: for Monoicomyces oxytelis read Verhaltnisses.- Parasitol. Schriftenr. 19: 1- 176. Monoicomyces oxyteli Siemaszko, J. & Siemaszko, W. 1928: Owadorosty polskie page 67 right column line 40: for Yliirj arvi read Ylojarvi i palearktyczne (Laboulbeniales J>O ionici et palaearctici). page 80 line 3: for italicus read halipli - Polskie Pismo Entomol. 6: 188-21 I. page 103 line 1: for b) immature read c)immature Spegazzini, C. 1912: Contribuci6n al estudio de las page 121: Thaxter 1899-1912 are misplaced under Wize Laboulbeniomicetas argentinas. - An. Mus. Nac. Hist. 1929 Nat. Buenos Aires 23: 167-244. 1914: Primo contribute alia conoscenza delle page 124 right column line 3: for fungus read fuscus Laboulbeniali italiani.-Redia 10: 21-75. page 130 line 24 (Elaphrus cupreus): add Laboulbenia elaphri !915a: Laboulbeniali ritrovati nelle collezioni di alcuni page 132 line 65 (P. debilis): add (I) Dichomyces hybridus musei italiani. -An. Mus. Nac. Hist. Nat. Buenos Aires page 133 line 28: for elonngatus read elongatulus 26: 451 -511. page 135 column line 19: for Agonum septentrionis read 1915b: Segunda contribuci6n a! conocimiento de las Patrobus septentrionis Laboulbeniales italianas. - An. Mus. Nac. Hist. Nat. pages 46, 11 I, 124, 132, 135: Choleva septentrionis Jeanne! Buenos Aires 27 : 37-74. should be interpreted as Choleva lederiana Reitter