Biomechanical Reconstruction of the Appendicular Skeleton in Three
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The Braincase, Brain and Palaeobiology of the Basal Sauropodomorph Dinosaur Thecodontosaurus Antiquus
applyparastyle “fig//caption/p[1]” parastyle “FigCapt” Zoological Journal of the Linnean Society, 2020, XX, 1–22. With 10 figures. Downloaded from https://academic.oup.com/zoolinnean/advance-article/doi/10.1093/zoolinnean/zlaa157/6032720 by University of Bristol Library user on 14 December 2020 The braincase, brain and palaeobiology of the basal sauropodomorph dinosaur Thecodontosaurus antiquus ANTONIO BALLELL1,*, J. LOGAN KING1, JAMES M. NEENAN2, EMILY J. RAYFIELD1 and MICHAEL J. BENTON1 1School of Earth Sciences, University of Bristol, Bristol BS8 1RJ, UK 2Oxford University Museum of Natural History, Parks Road, Oxford OX1 3PW, UK Received 27 May 2020; revised 15 October 2020; accepted for publication 26 October 2020 Sauropodomorph dinosaurs underwent drastic changes in their anatomy and ecology throughout their evolution. The Late Triassic Thecodontosaurus antiquus occupies a basal position within Sauropodomorpha, being a key taxon for documenting how those morphofunctional transitions occurred. Here, we redescribe the braincase osteology and reconstruct the neuroanatomy of Thecodontosaurus, based on computed tomography data. The braincase of Thecodontosaurus shares the presence of medial basioccipital components of the basal tubera and a U-shaped basioccipital–parabasisphenoid suture with other basal sauropodomorphs and shows a distinct combination of characters: a straight outline of the braincase floor, an undivided metotic foramen, an unossified gap, large floccular fossae, basipterygoid processes perpendicular to the cultriform process in lateral view and a rhomboid foramen magnum. We reinterpret these braincase features in the light of new discoveries in dinosaur anatomy. Our endocranial reconstruction reveals important aspects of the palaeobiology of Thecodontosaurus, supporting a bipedal stance and cursorial habits, with adaptations to retain a steady head and gaze while moving. -
By HENRY FAIRFIELD OSBORN and CHARL
VoL. 6, 1920 PALAEONTOLOGY: OSBORN AND MOOK IS RECONSTRUCTION OF THE SKELETON OF THE SAUROPOD DINOSAUR CAMARASA URUS COPE (MOROSA URUS MARSH) By HENRY FAIRFIELD OSBORN AND CHARLES CRAIG MOOK AMERICAN MusEUM or NATURAL HISTORY, NEW YORK CITY Read before the Academy, November 11, 1919 The principles of modern research in vertebrate palaeontology are illustrated in the fifteen years' work resulting in the restoration of the massive sauropod dinosaur known as Camarasaurus, the "chambered saurian.." The animal was found near Canyon City, Colorado, in March, 1877. The first bones were described by Cope, August 23, 1877. The first at- tempted restoration was by Ryder, December 21, 1877. The bones analyzed by this research were found probably to belong to six individuals of Camarasaurus mingled with the remains of some carnivorous dinosaurs, all from the summit of the Morrison formation, now regarded as of Jurassic- Cretaceous age. In these two quarries Cope named nine new genera and fourteen new species of dinosaurs, none of which have found their way into. palaeontologic literature, excepting Camarasaurus. Out of these twenty-three names we unravel three genera, namely: One species of Camarasaurus, identical with Morosaurus Marsh. One species of Amphicaclias, close to Diplodocus Marsh. One species of Epanterias, close to Allosaurus Marsh. The working out of the Camarasaurus skeleton results in both the artica ulated restoration and the restoration of the musculature. The following are the principal characters: The neck is very flexible; anterior vertebrae of the back also freely movable; the division between the latter and the relatively rigid posterior dorsals is sharp. -
A Reexamination of Four Prolacertiforms \Tith Implications for Pterosaur Phylogenesis
Rìvista Italiana di Paleontologia e Stratigrafia Dicembre 2000 I--r4-""l*-I-."-''* 1 A REEXAMINATION OF FOUR PROLACERTIFORMS \TITH IMPLICATIONS FOR PTEROSAUR PHYLOGENESIS DAVID PETERS ReceìterJ October 23, 1999; accepted October 20, 200A Kqt uorcls: Pterosauria, Prolacertiiormes (Reprilia, Diapsida), Traditionally the answer has been rhat prerosaurs Phyìogeny, Cladisric an:ìy.is. are archosaurs (Romer 1956); the sister group of the Riassunto . Tradizionalmente gli prerosauri venir.ano considerati Dinosauria, ScleromochÌus a.nd Lagosuclcws/Maraswchus come appartenenti agli Archosaurifomes e molti specìalistì contempo_ (Benton 1985, 1990, 1999; Padian 1984; Gauthter 1984, ranei considerano gli pterosauri quali sisrer groups di Lagosuchus, 1986; Sereno 1991, 1994; Kellner 1996); or perhaps Schleromochlus e dei Dinosauria. La nuova analisi filogenerica qui pro- archosauriformes close posta merte in discussione queste affinirà jn quanto tutte le presunte to prorerosuchids and eryrhro- sinapomorfie che collegherebbero gli Pterosauria con gli Archosauri_ suchids (Bennett 1996a), chiefly because prerosaurs formes o con gli pterosaurìa, Ornìthodira mancano in realtà negli have a prominent anrorbiral fenestra and a suite of other oppure sono condivise anche da alcuni taxa di prolacertiformi. ll archosaur-like characrers almosr entirely recente riesame degli olotipi dt confined to the Cosesaurus a,Liceps, Longisquama ìnsig_ hind nis e di Sharovipteryx mìrabi/ìs suggeriscono che molti caratteri potreb- limb (Bennert 1996a). Although Benton (1982, bero venire interpretati in maniera diversa rispetto alle precedenti L984) initially indicated that the prerosauria are descrìzioni. I risultati di molteplici analisì cladistjche suggeriscono che archosauromorphs and the sister-group ro all other questi tre prolacertìformi enigmatici, uniramente a Langobardìsawrws, archosauromorphs, later work (Benton 1985, 1.990, recentemente descritto, costituirebbero i sister taxa degli prerosauri, in base ad un insieme di sinapomorfie di nuova identificazione. -
A New Clade of Basal Early Cretaceous Pygostylian Birds and Developmental Plasticity of the Avian Shoulder Girdle
A new clade of basal Early Cretaceous pygostylian birds and developmental plasticity of the avian shoulder girdle Min Wanga,b,1, Thomas A. Stidhama,b, and Zhonghe Zhoua,b,1 aKey Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China; and bCenter for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Beijing 100044, China Contributed by Zhonghe Zhou, August 16, 2018 (sent for review July 16, 2018; reviewed by Stephen L. Brusatte and Gareth Dyke) Early members of the clade Pygostylia (birds with a short tail Systematic Paleontology ending in a compound bone termed “pygostyle”)arecriticalfor Aves Linnaeus, 1758; Pygostylia Chiappe, 2002; Jinguofortisidae understanding how the modern avian bauplan evolved from fam. nov. (SI Appendix, SI Text); Jinguofortis perplexus gen. et sp. nov. long-tailed basal birds like Archaeopteryx. However, the cur- rently limited known diversity of early branching pygostylians Holotype obscures our understanding of this major transition in avian A complete and articulated skeleton with feathers is housed at evolution. Here, we describe a basal pygostylian, Jinguofortis the Institute of Vertebrate Paleontology and Paleoanthropology perplexus gen. et sp. nov., from the Early Cretaceous of China (IVPP) under the collect number IVPP V24194 (Fig. 1 and SI that adds important information about early members of the Appendix, Figs. S1–S7 and Table S1). short-tailed bird group. Phylogenetic analysis recovers a clade (Jinguofortisidae fam. nov.) uniting Jinguofortis and the enig- Etymology matic basal avian taxon Chongmingia that represents the second The generic name is derived from “jinguo” (Mandarin), referring earliest diverging group of the Pygostylia. -
Reconstruction of the Skeleton of The
VoL. 6, 1920 PALAEONTOLOGY: OSBORN AND MOOK IS RECONSTRUCTION OF THE SKELETON OF THE SAUROPOD DINOSAUR CAMARASA URUS COPE (MOROSA URUS MARSH) By HENRY FAIRFIELD OSBORN AND CHARLES CRAIG MOOK AMERICAN MusEUM or NATURAL HISTORY, NEW YORK CITY Read before the Academy, November 11, 1919 The principles of modern research in vertebrate palaeontology are illustrated in the fifteen years' work resulting in the restoration of the massive sauropod dinosaur known as Camarasaurus, the "chambered saurian.." The animal was found near Canyon City, Colorado, in March, 1877. The first bones were described by Cope, August 23, 1877. The first at- tempted restoration was by Ryder, December 21, 1877. The bones analyzed by this research were found probably to belong to six individuals of Camarasaurus mingled with the remains of some carnivorous dinosaurs, all from the summit of the Morrison formation, now regarded as of Jurassic- Cretaceous age. In these two quarries Cope named nine new genera and fourteen new species of dinosaurs, none of which have found their way into. palaeontologic literature, excepting Camarasaurus. Out of these twenty-three names we unravel three genera, namely: One species of Camarasaurus, identical with Morosaurus Marsh. One species of Amphicaclias, close to Diplodocus Marsh. One species of Epanterias, close to Allosaurus Marsh. The working out of the Camarasaurus skeleton results in both the artica ulated restoration and the restoration of the musculature. The following are the principal characters: The neck is very flexible; anterior vertebrae of the back also freely movable; the division between the latter and the relatively rigid posterior dorsals is sharp. -
The Princeton Field Guide to Dinosaurs, Second Edition
MASS ESTIMATES - DINOSAURS ETC (largely based on models) taxon k model femur length* model volume ml x specific gravity = model mass g specimen (modeled 1st):kilograms:femur(or other long bone length)usually in decameters kg = femur(or other long bone)length(usually in decameters)3 x k k = model volume in ml x specific gravity(usually for whole model) then divided/model femur(or other long bone)length3 (in most models femur in decameters is 0.5253 = 0.145) In sauropods the neck is assigned a distinct specific gravity; in dinosaurs with large feathers their mass is added separately; in dinosaurs with flight ablity the mass of the fight muscles is calculated separately as a range of possiblities SAUROPODS k femur trunk neck tail total neck x 0.6 rest x0.9 & legs & head super titanosaur femur:~55000-60000:~25:00 Argentinosaurus ~4 PVPH-1:~55000:~24.00 Futalognkosaurus ~3.5-4 MUCPv-323:~25000:19.80 (note:downsize correction since 2nd edition) Dreadnoughtus ~3.8 “ ~520 ~75 50 ~645 0.45+.513=.558 MPM-PV 1156:~26000:19.10 Giraffatitan 3.45 .525 480 75 25 580 .045+.455=.500 HMN MB.R.2181:31500(neck 2800):~20.90 “XV2”:~45000:~23.50 Brachiosaurus ~4.15 " ~590 ~75 ~25 ~700 " +.554=~.600 FMNH P25107:~35000:20.30 Europasaurus ~3.2 “ ~465 ~39 ~23 ~527 .023+.440=~.463 composite:~760:~6.20 Camarasaurus 4.0 " 542 51 55 648 .041+.537=.578 CMNH 11393:14200(neck 1000):15.25 AMNH 5761:~23000:18.00 juv 3.5 " 486 40 55 581 .024+.487=.511 CMNH 11338:640:5.67 Chuanjiesaurus ~4.1 “ ~550 ~105 ~38 ~693 .063+.530=.593 Lfch 1001:~10700:13.75 2 M. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
Neural Spine Bifurcation in Sauropods Palarch’S Journal of Vertebrate Palaeontology, 10(1) (2013)
Wedel & Taylor, Neural Spine Bifurcation in Sauropods PalArch’s Journal of Vertebrate Palaeontology, 10(1) (2013) NEURAL SPINE BIFURCATION IN SAUROPOD DINOSAURS OF THE MORRISON FORMATION: ONTOGENETIC AND PHYLOGENETIC IMPLICATIONS Mathew J. Wedel* & Michael P. Taylor# *Corresponding author. College of Osteopathic Medicine of the Pacific and College of Podiatric Medicine, Western University of Health Sciences, 309 E. Second Street, Pomona, California 91766-1854, USA. [email protected] #Department of Earth Sciences, University of Bristol, Bristol BS8 1RJ, UK. [email protected] Wedel, Mathew J. & Michael P. Taylor. 2013. Neural Spine Bifurcation in Sauropod Dinosaurs of the Morrison Formation: Ontogenetic and Phylogenetic Implications. – Pal- arch’s Journal of Vertebrate Palaeontology 10(1) (2013), 1-34. ISSN 1567-2158. 34 pages + 25 figures, 2 tables. Keywords: sauropod, vertebra, neural spine, ontogeny, Morrison Formation AbsTrAcT It has recently been argued that neural spine bifurcation increases through ontogeny in several Morrison Formation sauropods, that recognition of ontogenetic transforma- tion in this ‘key character’ will have sweeping implications for sauropod phylogeny, and that Suuwassea and Haplocanthosaurus in particular are likely to be juveniles of known diplodocids. However, we find that serial variation in sauropod vertebrae can mimic on- togenetic change and is therefore a powerful confounding factor, especially when deal- ing with isolated elements whose serial position cannot be determined. When serial po- sition is taken into account, there is no evidence that neural spine bifurcation increased over ontogeny in Morrison Formation diplodocids. Through phylogenetic analysis we show that neural spine bifurcation is not a key character in sauropod phylogeny and that Suuwassea and Haplocanthosaurus are almost certainly not juveniles of known diplodo- cids. -
Re-Description of the Sauropod Dinosaur Amanzia (“Ornithopsis
Schwarz et al. Swiss J Geosci (2020) 113:2 https://doi.org/10.1186/s00015-020-00355-5 Swiss Journal of Geosciences ORIGINAL PAPER Open Access Re-description of the sauropod dinosaur Amanzia (“Ornithopsis/Cetiosauriscus”) greppini n. gen. and other vertebrate remains from the Kimmeridgian (Late Jurassic) Reuchenette Formation of Moutier, Switzerland Daniela Schwarz1* , Philip D. Mannion2 , Oliver Wings3 and Christian A. Meyer4 Abstract Dinosaur remains were discovered in the 1860’s in the Kimmeridgian (Late Jurassic) Reuchenette Formation of Moutier, northwestern Switzerland. In the 1920’s, these were identifed as a new species of sauropod, Ornithopsis greppini, before being reclassifed as a species of Cetiosauriscus (C. greppini), otherwise known from the type species (C. stewarti) from the late Middle Jurassic (Callovian) of the UK. The syntype of “C. greppini” consists of skeletal elements from all body regions, and at least four individuals of diferent sizes can be distinguished. Here we fully re-describe this material, and re-evaluate its taxonomy and systematic placement. The Moutier locality also yielded a theropod tooth, and fragmen- tary cranial and vertebral remains of a crocodylomorph, also re-described here. “C.” greppini is a small-sized (not more than 10 m long) non-neosauropod eusauropod. Cetiosauriscus stewarti and “C.” greppini difer from each other in: (1) size; (2) the neural spine morphology and diapophyseal laminae of the anterior caudal vertebrae; (3) the length-to-height proportion in the middle caudal vertebrae; (4) the presence or absence of ridges and crests on the middle caudal cen- tra; and (5) the shape and proportions of the coracoid, humerus, and femur. -
A New Camarasaurid Sauropod Opisthocoelicaudia Skarzynskii Gen
MAGDALENA BORSUK-BIALYNICKA A NEW CAMARASAURID SAUROPOD OPISTHOCOELICAUDIA SKARZYNSKII GEN. N., SP. N. FROM THE UPPER CRETACEOUS OF MONGOLIA (plates 1-14) Abstract. - An almost complete postcranial skeleton lacking cervicals of Opisthocoelicaudia skarzynskii gen. n., sp. n. (Sauropoda, Camarasauridae) from the Upper Cretaceous Nemegt Formation, Gobi Desert , is described and figured. The reconstruction of the muscle system and sternum as well as the restoration of the whole animal is made. It is shown that Opisthocoelicaudia was a straight backed sauropod with the tail carried in a horizontal position. The neck is supposed to have been of medium length (about 5 m) and was carried low. The possibility of habitual assuming a tirpodal position is suggested by the opisthocoelous structure of the ant erior caudals. The importance of some osteologic features of sauropods for the understanding of their attitudes as well as for the systematics is discussed. It is argued that the length of neural spines depends on both the curvature of the back-bone and the length of the neck and tail in sauropods. Forked neural spines are indicative ot the habitual lowering of the neck, or even of the low carrying of the neck, if the anterior dorsals lack traces of the nuchal ligament insertion. Some titanosaurid characters of Opisthocoelicaudia are regarded as progressive ones in sauropods, whereas its camarasaurid features seem to indicate a true relationship in spite of their highly behavioural character. CONTENTS Page Introduction. 6 Description ... 8 Vertebral column 9 Thoracic ribs . 18 Sternum . 19 Pectoral girdle 22 Fore limbs . 24 Pelvic girdle . 32 Hind limbs . -
The Scapulocoracoid of an Early Triassic Stem−Frog from Poland
The scapulocoracoid of an Early Triassic stem−frog from Poland MAGDALENA BORSUK−BIAŁYNICKA and SUSAN E. EVANS Borsuk−Białynicka, M. and Evans, S.E. 2002. The scapulocoracoid of an Early Triassic stem−frog from Poland. Acta Palaeontologica Polonica 47 (1): 79–96. The scapulocoracoid of Czatkobatrachus polonicus Evans and Borsuk−Białynicka, 1998, a stem−frog from the Early Tri− assic karst locality of Czatkowice (Southern Poland), is described. The overall type of scapulocoracoid is plesiomorphic, but the subcircular shape and laterally oriented glenoid is considered synapomorphic of Salientia. The supraglenoid fora− men is considered homologous to the scapular cleft of the Anura. In Czatkobatrachus, the supraglenoid foramen occupies an intermediate position between that of the early tetrapod foramen and the scapular cleft of Anura. The cleft scapula is probably synapomorphic for the Anura. In early salientian phylogeny, the shift in position of the supraglenoid foramen may have been associated with an anterior rotation of the forelimb. This change in position of the forelimb may reflect an evolutionary shift from a mainly locomotory function to static functions (support, balance, eventually shock−absorption). Laterally extended limbs may have been more effective than posterolateral ones in absorbing landing stresses, until the specialised shock−absorption pectoral mechanism of crown−group Anura had developed. The glenoid shape and position, and the slender scapular blade, of Czatkobatrachus, in combination with the well−ossified joint surfaces on the humerus and ulna, all support a primarily terrestrial rather than aquatic mode of life. The new Polish material also permits clarifica− tion of the pectoral anatomy of the contemporaneous Madagascan genus Triadobatrachus. -
Dinosaur Species List E to M
Dinosaur Species List E to M E F G • Echinodon becklesii • Fabrosaurus australis • Gallimimus bullatus • Edmarka rex • Frenguellisaurus • Garudimimus brevipes • Edmontonia longiceps ischigualastensis • Gasosaurus constructus • Edmontonia rugosidens • Fulengia youngi • Gasparinisaura • Edmontosaurus annectens • Fulgurotherium australe cincosaltensis • Edmontosaurus regalis • Genusaurus sisteronis • Edmontosaurus • Genyodectes serus saskatchewanensis • Geranosaurus atavus • Einiosaurus procurvicornis • Gigantosaurus africanus • Elaphrosaurus bambergi • Giganotosaurus carolinii • Elaphrosaurus gautieri • Gigantosaurus dixeyi • Elaphrosaurus iguidiensis • Gigantosaurus megalonyx • Elmisaurus elegans • Gigantosaurus robustus • Elmisaurus rarus • Gigantoscelus • Elopteryx nopcsai molengraaffi • Elosaurus parvus • Gilmoreosaurus • Emausaurus ernsti mongoliensis • Embasaurus minax • Giraffotitan altithorax • Enigmosaurus • Gongbusaurus shiyii mongoliensis • Gongbusaurus • Eoceratops canadensis wucaiwanensis • Eoraptor lunensis • Gorgosaurus lancensis • Epachthosaurus sciuttoi • Gorgosaurus lancinator • Epanterias amplexus • Gorgosaurus libratus • Erectopus sauvagei • "Gorgosaurus" novojilovi • Erectopus superbus • Gorgosaurus sternbergi • Erlikosaurus andrewsi • Goyocephale lattimorei • Eucamerotus foxi • Gravitholus albertae • Eucercosaurus • Gresslyosaurus ingens tanyspondylus • Gresslyosaurus robustus • Eucnemesaurus fortis • Gresslyosaurus torgeri • Euhelopus zdanskyi • Gryponyx africanus • Euoplocephalus tutus • Gryponyx taylori • Euronychodon