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HAROLD F. DE LISLE, LYNN R. G. RAW,

DANIEL A. MELNIKOV

AGAMIDAE

A CATALOG OF RECENT

HAROLD F. DE LISLE, LYNN R. G. RAW,

DANIEL A. MELNIKOV

© Harold F. De Lisle, Daniel Melnikov, Lynn R.G. Raw, 2016 Email: [email protected]

Cover photo: Psammophilus dorsalis, male in breeding colours, Hyderabad, , by JM Garg Back cover: kingie, Hann River area, Queensland, , photo by Stephen Zozaya

CONTENTS

INTRODUCTION …...... 1

CATALOG …...... 13

ACKNOWLEGEMENTS …...... 99

REFERENCES . …...... 105

TAXON INDEX …...... 143

INTRODUCTION

Family

Iguaniens (part), Cuvier, 1817, Le règne distribué d’après son organisation, pour servir de base a l’historire naturelle des animaux et d’introduction a l’anatomie comparée . Vol. 2. Les , les poissons, les mollusques et les annélides. Déterville, Paris

Ascalabotae (part), Merrem, 1820, Versuch eines Systems der Amphibien I (Tentamen Systematis Amphibiorum). J. C. Krieger, Marburg, 191 pp.

Stellionidae (part), Gray 1825, A synopsis of the genera of reptiles and Amphibia, with a description of some new species. Annals of Philosophy , 10: 196

Agamidae Gray, 1827 , A Synopsis of the Genera of Saurian Reptiles in which some new Genera are indicated, and the others reviewed by actual Examination. Philosophical Magazine, London, 2 (2): 57

Iguaniens acrodontes Duméril & Bibron, 1837, Erpetologie Générale ou Histoire Naturelle Complete des Reptiles. Vol.4. Libr. Encyclopédique Roret, Paris

Uromasticidae Theobald, 1868, Catalogue of the reptiles of British Burma, embracing the provinces of Pegu, Martaban, and Tenasserim; with descriptions of new or little-known species. Journal of the Linnean Society , London, Zool., 10: 34

The large family Agamidae contains Some Agamid is adapted to almost some 450 species, which are mostly every type of terrestrial habitat. From sea confined to Asia, Australia, and . level to the top of tropical mountain; from Before the recent split of the family hot desert (Sahara) to cold desert (Gobi). Iguanidae, the two generas were often Some are arenicolous, many are saxicolous , compared, with the Iguanidae largely found many more are arboreal. A few are almost in the Western Hemisphere. semiaquatic, but none are semifossial . Perhaps the major anatomical Most Agamids are insectivores, but a difference between the two families is in few are omnivorous. A very few are dentition. Agamids have acrodont and herbivores. heterdont dentition, a character they only All Agamids are oviparous. Most share with the Chamaeleonidae. Dermal species lay leathery eggs with the oval shape appendages such as crests, dewlaps, and that are typical of most eggs, while a even wing-like mebranes ( Draco ) are often few genera ( ) lay spindle-shaped present. is the rule, with eggs. There are few enigmatic and the males often having brilliant colors and chrestomatic examples of viviparous more pronounced dermal appendages. Agamids. They are extraordinary The eye of Agamids is provided with representatives of different subfamilies. complete lids and the pupil is always round. Live-bearing are found in Some species have no tympanum while from high elevations of others have large exposed tympanum. Only Tibet and Himalaya ( Ph. vlangalii , Ph. a few genera have femoral pores typical of theobaldi , Ph. putjatai and others) , and . The tongue is rather short and broad. lowland desert of Takla-Makan ( Ph. Limbs are well developed. Tail is usually forsyhtii ). Viviparous are long and tough ceyloniaca and Harpesaurus borneensis . another with G . robinsonii, and a third with Recent phylogenetic studies have ). found that the genera Moloch and At present (2019) there are 53 genera render paraphyletic . in the Family Agamidae. This includes is non-monophyletic. Also, several monotype genera, many of which are (G. robinsonii is only part of the morbid thirst for naming new distantly related to other Gonocephalus ) and genera in the last two decades (Kaiser et al., Japalura (with species distributed 2013). There are also ditype genera among three distantly related clades, including one allied with Ptyctolaemus, .

Acanthocercus Fitzinger, 1843 (13 species) Physignathus Cuvier, 1829 (1 species) Gray, 1831 (11 species) Storr, 1982 (8 species) Daudin, 1802 (47 species) Psammophilus Fitzinger, 1843 (2 species) Wagler, 1830 (4 species) Pseudocalotes Fitzinger, 1843 (23 species) Peters, 1864 (2 species) Pseudocophotis Manthey and Grossmann. 1997 (2 Kaup, 1827 (12 species) species) Bufoniceps Arnold, 1992 (1 species) Fitzinger, 1843 (6 species) Calotes Cuvier, 1817 (23 species) Ptyctolaemus Peters, 1865 (2 species) Ceratophora Gray, 1835 (4 species) Wells & Wellington, 1984 (1 species) Chelosania Gray, 1845 ( 1 species) Salea Gray, 1845 (3 species) Chlamydosaurus Gray, 1825 (1 species) Sarada Deepak, Karanth and Giri, 2016 (3 species) Complicitus Manthey & Grossmann, 1997 (1 species) Sitana Cuvier, 1829 (13 species) Cophotis Peters, 1861 (2 species) Thaumatorhynchus Parker, 1924 (1 species) Blyth, 1860 (2 species) Cuvier, 1817 (13 species) Witten, 1984 (2 species) Tropicagama Melville et al., 2019 (1 species) Cristidorsa Wang et al., 2018 (2 species) Peters, 1863 (15 species) Fitzinger, 1843 (29 species) Uromastyx Merrem, 1820 (18 species) Dendragama Doria, 1888 (3 species) Boulenger, 1895 (4 species) Diploderma Hallowell, 1861 (24 species) Gray, 1842 (22 species) Draco Linnaeus, 1758 (39 species) Gonocephalus Kaup, 1825 (16 species) Wells & Wellington. 1984 (2 species). Harpesaurus Boulenger, 1885 (5 species) Hydrosaurus Kaup, 1828 (3 species) Hypsicalotes Manthey & Denzer, 2000 (1 species) Hypsilurus Peters, 1867 (16 species) Intellagama Wells & Wellington, 1985 (1 species) Japalura Gray. 1853 (7 species) Gray, 1845 (20 species) Leiolepis Cuvier, 1829 (9 species) Lophocalotes Günther, 1872 (2 species) Lophognathus Gray. 1842 (2 species) Günther, 1872 (3 species) Lyriocephalus Merrem, 1820 (1 species) Malayodracon Denzer, 2015 (1 species) Mantheyus Ananjeva and Stuart, 2001 (1 species) Moloch Gray, 1841 (1 species) Monilesarus Pal et al, 2018 (4 species) Wagler, 1830 (3 species) Phoxophrys Hubrecht, 1881(5 species) Phrynocephalus Kaup, 1825 (31 species)

Many authors (Anajeva, 2011) subdivide the Southeast Asian Draconinae Fitzinger, 1826 agamids into six subfamilies: (1) with numerous Indian and Southeastern Uromastycinae Theobald, 1868 with the genera (mostly arboreal or semi-arboreal) genera Uromastyx and Saara ; (2) and (6) Afro-West-Asian Agaminae Spix, Leiolepidinae Fitzinger, 1843 with the 1825. Leiolepis ; (3) Wagler, 1830 The exact position of many genera is with all Australian and New Guinean still debated among taxonomists. species; (4) Hydrosaurinae K аuр, 1828 with the genus Hydrosaurus ; (5) South- and

This is the phylogeny according to Pyron, Burbrink and Wiens, 2013( Pyron et al. BMC Evolutionary 2013)

Abbreviations of museums holding the types are as follows

AMNH - American Museum of Natural History, Division of Vertebrate Zoology (), 79th Street and Central Park West, New York, New York 10024, USA. ANSP - Academy of Natural Sciences, Department of Herpetology, 19th and the Benjamin Franklin Parkway, Philadelphia, Pennsylvania 19103, USA. ANWC – Australian National Wildlife Collection, Canberra, Australia BA – Insitute of Rare and Plants, Sichuan Teachers College, Nanchong, Sichuan Province 637702, BMNH – The Natural History Museum, Department of Zoology, Cromwell Road, London SW7 5BD, United Kingdom. BPBM - Bernice P. Bishop Museum, Department of Zoology, 1355 Kalihi Street, Honolulu, Hawaii 96818, USA. BSP - Bureau Science (destroyed in WWII). CAS - California Academy of Sciences, Department of Herpetology, Golden Gate Park, San Francisco, California 94118, USA. CIB - Chengdu Institute of Biology, Academia Sinica, P.O. Box 416, Chengdu, Sichuan, P.R. China. DB.ULPGC - Departamento de Biología, Universidad de Las Palmas de Gran Canaria, Calle Juan de Quesada, 30, 35001 Las Palmas de Gran Canaria, Las Palmas, Spain FMNH - Field Museum, Division of Amphibians and Reptiles, Roosevelt Road at Lake Shore Drive, Chicago, Illinois 60605, USA. GNHM - Göteborgs Naturhistoriska Museum, Museivägen 10, 413 11, Göteborg, Sweden HLMD - Hessisches Landesmuseum, Darmstadt, Germany. HUJ - Hebrew University, Department of Zoology, Jerusalem, Israel. IEBR - Institute of Ecology and Biological Resources, No 18 Hoang Quoc Viet Street, Hanoi, IZANU - Zoological Institute. Ukrainian National Academy of Sciences, Kiev, Ukraine KIZ - Kunming Institute of Zoology, Chinese Academy of Sciences, 32 East Jaio Chang Road; 650223, Kunming, Yunnan, China KUZ - Department of Zoology, Graduate School of Science, Kyoto University, Sakyo, Kyoto 606-8502, Japan. HNM - Hungarian National Museum, Budapest, Múzeum körút 14-16, 1088 Hungary LSUHC - La Sierra University Herpetological Collection, La Sierra University, Riverside, California MBL - Museu Bocage, Universidade de Lisboa, Lisbon, Portugal [destroyed by fire in March 1978] MCZ - Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts 02138, USA MHNCF - Musée d'histoire naturelle, 2300 La Chaux-de-Fonds, Switzland MHNP - Museum National dHistoire Naturelle, Laboratoire des Amphibiens et Reptiles, 25 Rue Cuvier, 75005 Paris, France MMK - McGregor Museum, Egerton Rd, Kimberley 8301 , Cape Province, MNCN - Museo Nacional de Ciencias Naturales, Calle José Gutiérrez Abascal, 2, 28006, Madrid, Spain MSNG - Museo Civico di Storia Naturale di Genova Giacomo Doria, Via Brigata Liguria 9, 16121 Genova, Italy MTKD - Staatliche Naturhistorische Sammlungen Dresden, Museum für Tierkunde, Königsbrücker Landstrasse 159, 01109 Dresden, Germany MZB - Museum Zoologicum Bogoriense, Jl. Raya Juanda 3, Bogor, , MZUT - Universita di Torino, Museo Zoologico, Torino, Italy NCBS – National Centre for Biological Sciences, Bangalore, GKVK Campus, Bellary Rd, Bengaluru, Karnataka 560065, India NHMB - Naturhistorisches Museum, Augustinergasse 2, 4001 Basel, Switzerland NMBE - Naturhistorisches Museum der Burgergemeinde Bern, Bernastrasse 15, 3005 , Bern, Switzerland NMK - National Museums of , P.O. Box 78420-00500 Ngara Rd, Nairobi, Kenya NMP6V - Department of Zoology, National Museum, Vinohradská 1, 110 00 Prague 1, Czech Republic NMSL - Sri Lanka National Museum, PO Box 854, Colombo 07, Sri Lanka. NMU - Nanjing Medical University, Nanjing, China NMV - National Museum of Victoria, Melbourne, Victoria, Australia NMW - Naturhistorisches Museum, Zoologische Abtheilung, Postfach 417, Burgring 7, 1014 Wien, Austria. NNU - i Nanjing Normal University, Jiangsu, Nanjing, Gulou, China NRC/ZSI - Northern Regional Centre /Zoological Survey of India, 218, Kaulagarh Road, Dehra Dun- 248 195, Uttrakhand, India ntre /Zoological Survey of Indiag Normal University NRM - Naturhistoriska Rijkmuseet, Section for Vertebrate Zoology, Roslagsvägen 120, Box 50007, 104 05 Stockholm 50, Sweden. NTNUB - Institute of Zoology, National Taiwan University, No. 1 Sec. 4 Roosevelt Road, Taipei, Taiwan OMNH - Osaka Museum of Natural History, Nagai Park, Higashi-sumiyoshi-ku, Osaka 546, Japan. OUM - Oxford University Museum of Natural History, Parks Rd, Oxford OX1 3PW, United Kingdom

PMNH - Pakistan Museum of Natural History, Garden Avenue, Shakarparian, Islamabad 44000, Pakistan PNM - Philippine National Museum, Taft Ave, 1000 Manila, Philippines QM - Queensland Museum, Gregory Terrace, Fortitude Valley, Brisbane, Queensland 4006, Australia. RMNH - Naturalis (Nationaal Natuurhistorisch Museum; formerly Rijksmuseum van Natuurlijke Historie), Postbus 9517, 2300RA Leiden, The Netherlands SAM - South African Museum, P.O. Box 61, Cape Town 8000, Western Cape Province, South Africa. SAMA - South Australia Museum, North Terrace, Adelaide, South Australia 5000, Australia SHHS -Fuhlrott -Museum, Auer Schulstraße 20 42103, Wuppertal, Germany SIZ – Shaanzi Institute of Zoology, Shaanxi Normal University Xi'an, Shaanxi, 710062, China SMF - Forschungsinstitut und Natur-Museum Senckenberg, Senckenberg-Anlage 25, 6000 Frankfurt-am-Main 1, Germany SMK - Sarawak Museum, Pengarah Jabatan Muzium Negeri Sarawak, Jalan Tun Abang Haji Openg, 93566, Kuching, Sarawak, 93000 , East SMNS - Staatliches Museum für Naturkunde, Rosenstein 1, 70191 , Stuttgart, Germany SRNC - Suzhou Railway Teacher’s College (now Suzhou University of Science and Technology), Department of Biology, 1701 BinheRoad Suzhou 215011, Jiangsu, China SYSU - Sun Yat-sen University , 135 Xingang West Rd, Haizhu, Guangzhou, Guangdong, China THNHM - National Science Museum, Natural History Museum, Technopolois, Klong 5, Klong Luang, Patumthani 12120, Thailand TMP - Transvaal Museum, P.O. Box 413, Pretoria 0001, Transvaal, South Africa TINHC - Thailand Institute of Scientific and Technological Research, 196 Phahonyothin Road, Chatuchak, Bangkok 10900 Thailand TNHC - Texas Natural History Collections, Texas Natural Science Center, Texas Memorial Museum, University of Texas at Austin, Austin, Texas USNM - National Museum of Natural History, Division of Amphibians and Reptiles, Washington, D.C. 20560, USA UUZM - Uppsala University Museum of , Zoology section, Norbyvägen 16, 752 36 Uppsala , Sweden WAM - Museum, Francis Street, Perth, Western Australia 6000, Australia WHT - Wildlife Heritage Trust, 95 Cotta Road, Colombo 8, Sri Lanka. ZDKU - Museum of Natural History, V. N. Karazin Kharkiv National University , Kharkiv, Ukraine ZFMK - Zoologisches Forschungsinstitut und Museum Alexander Koenig, Herpetologische Abteilung, Adenauerallee 150-164, 5300 Bonn-1, Germany ZISP - Zoological Institute, Russian Academy of Sciences, St. Petersburg 199034, Russia ZMA - Universiteit van Amsterdam, Zoologisch Museum, Plantage Middenlaan 53, Amsterdam: C, Netherlands. ZMB - Universität Humboldt, Zoologisches Museum, Invalidenstrasse 43, 10115 Berlin, Germany ZMH - Zoologisches Institut und Museum, Universität Hamburg, Martin-Luther-King Platz 3, D-2000 Hamburg 13, Germany ZMLU - Zoologiska museet, Lunds universitet, Helgonavägen, 223 62, Lund, Sweden ZMMU - M. V. Lomonosov Moscow State University, Zoological Museum, Ul. Herzena 6, 103009 Moscow K-9, Russia. ZMTU - Zoologiceskogo Muzei (Museum of Zoology), Tomsk State University, Tomsk, Russia ZMUC - Universitets København, Zoologisk Museum, Universitetsparken 15, 2100 København, Denmark ZRC - Zoological Reference Collection, Department of Zoology, University of Singapore, Bukit Timah Road, Singapore 10 ZSI - Zoological Survey of India, Herpetology Section, Fire-Proof Spirit Building, 27 JL Nehru Road, Kolkata (formerly Calcutta), 700 016, India ZSI/SRC - Zoological Survey of India, 130 Santhome High Road, Chennai 600 028, India ZSM - Zoologisches Staatssammlung, Münchhausenstrasse 21, 8000 München 21, Germany ZUTC - Zoology Museum of University of Tehran, Karaj, Tehran, Iran ZUUG - Zoologisches Museum der Georg-August Universität Göttingen, Göttingen, Niedersachen , Germany

Family Agamidae Gray, 1827 Gray, J.E. 1827. A Synopsis of the Genera of Saurian Reptiles in which some new Genera are indicated, and the others reviewed by actual Examination. Philosophical Magazine, London, 2 (2): 54-58 22 genera

Acanthocercus Fitzinger, 1843 Fitzinger, L. 1843. Systema Reptilium , fasciculus primus, Amblyglossae. Braumüller et Seidel, Wien: 106 pp. Type species: Stellio cyanogaster Rüppell, 1835, by original designation

Acanthocercus adramitanus (Anderson, 1896) Agama adramitana Anderson , 1896:31

D Fischer Type locality: between Makalla and the Hadramut Valley, Yemen Holotype : BMNH 1897.3.11.47 . Distribution: Oman, Yemen, southwestern Saudi Arabia

Acanthocercus annectens (Blanford, 1870) Agama annectans Blanford, 1870:455 Type locality: Sooroo (= Suru Pass, ) Holotype: BMNH 1869.11.4.12 Distribution: , Kenya, , Djibouti

Acanthocercus atricollis (Smith, 1849)

L Raw Agama atricollis Smith, 1849: 14 Synonyms Stellio capensis Duméril in Duméril et al., 1851: 106 Type locality: unknown Holotype: MNHP 1866.69 Stellio nigricollis Bocage, 1866: 43 Type locality: Type: MBL, lost in fire Type locality: Durban, South Africa Lectotype: BMNH 1946.8.28.1 Distribution: South Africa, Swaziland, , , and

Acanthocercus atricollis loveridgei (Klausewitz , 1957) Agama atricollis loveridgei Klausewitz, 1957: 163 Holotype: SMF 10138. Type locality: Kakoma, Ost-Afrika (=) Distribution: Tanzania

Acanthocercus branch i Wagner, Greenbaum & Bauer, 2012 Acanthocercus branchi Wagner, Greenbaum & Bauer, 2012: 22 Type locality: Miombo woodland close to the Zambian Wildlife Authority (ZAWA) head office Chipata (ca .13º6053S, 32º6092E), Eastern Province, . Holotype: ZFMK 88682. Distribution: Luangwa and Zambezi valleys, Zambia

Acanthocercus cyanocephalus (Falk, 1925). Agama cyanocephala Falk, 1925: 83 Type locality: in a garden in Ikelenge (−11.241592, 24.273256), northern Mwinilunga District, North-Western Province, Zambia Neotype: ZFMK 88492, Distribution: Angola, Zambia and extreme northern

Acanthocercus cyanogaster (Rüppell, 1835) Stellio cyanogaster Rüppell, 1835: 10 Type locality: Massaua, Eritrea Lectotype : SMF 10145, designated by Mertens 1967 Distribution: Ethiopia, Eritrea, Kenya, Somalia

Acanthocercus gregorii ( Klausewitz, 1957)

bushtucker

Subspecies

Acanthocercus gregorii (Klausewitz, 1957) Agama atricollis gregorii Klausewitz 195: 165 Lectotype: BMNH 1946.8.28.67. Type locality: Mkonumbi, a grassy coast district with salt-swamps, Coast Province, Kenya , coast of Kenya Distribution: Ethiopia, Kenya, Tanzania, Rwanda, Burundi and extreme northern Mozambique

Acanthocercus gregorii loveridgei (Klausewitz , 1957) Agama atricollis loveridgei Klausewitz, 1957: 163 Type locality : Kakoma, Ost-Afrika (=Tanzania) Holotype: SMF 10138. Distribution: coastal Tanzania, Kenya

Acanthocercus guentherpetersi Largen & Sprawl, 2006 Acanthocercus guentherpetersi Largen & Sprawl, 2006: 28 Type locality: ca 40 km E of Harar (09°12’N 42°22’E), on the road toJigjiga, Ethiopia. Holotype: BMNH 1969.1254 Distribution: Ethiopia

Acanthocercus kiwuensis (Klausewitz , 1957) Agama atricollis kiwuensis Klausewitz, 1957:167 Type locality : Kissenji [=Gisenyi] north-east shore of Lake Kivu, Rwanda Holotype: ZMB 17693. Distribution: from Lake Kivu in the south to central and north-eastern DRC in the north: DR Congo, Uganda, Rwanda

Acanthocercus minutus (Klausewitz, 1957) Agama atricollis minuta Klausewitz, 1957: 170 Type locality : Dscheffedenza [=Ciaffedenza], Shoa’, Ethiopia Holotype: ZMB 29089. Distribution: Ethiopia, Kenya

Acanthocercus phillipsii (Boulenger, 1895) Agama phillipsii Boulenger, 1895:167 Synonym Agama trachypleura Peters, 1982:266 Type locality: Eritrea. Holotype: ZMB 37109 Type locality: Sheik Paw, Berbera, Holotype: BMNH 1895.6.14.11 Distribution: Eritrea, Ethiopia, northwestern Somalia

Acanthocercus ugandaensis (Klausewitz, 1957) Agama atricollis ugandaensis Klausewitz, 1957: 169 Type locality : Uganda Holotype: ZMB 11904. Distribution : Uganda, Tanzania, Rwanda

Acanthocercus yemensis (Klausewitz, 1954) Agama cyanogaster yemensis Klausewitz, 1954: 170 Type locality: Sana, Yemen Holotype: SMF 48174 Distribution: north Yemen, southwestern Saudi Arabia

Acanthosaura Gray, 1831 Gray, J.E. 1831. A synopsis of the species of Class Reptilia. In: Griffith, E & E. Pidgeon: The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with additional descriptions of all the species hither named, and of many before noticed [V Whittaker, Treacher and Co., London: 481 + 110 pp. Type species: Gray in Hardwicke, 1827, by original designation

Acanthosaura armata (Gray, 1827) Agama armata Gray in Hardwicke, 1827: 216 Type locality: Singapore Syntypes: BMNH 1946.8.13.88, 1946.8.13.85 Distribution: China: ; Burma; southern Thailand; West Malaysia, incl. Palau Pinang, Pulau Tioman; Indonesia: Sumatra

Acanthosaura bintangensis Wood, Grismer, Grismer, Ahmad, Onn & Bauer, 2009 Acanthosaura bintangensis Wood, Grismer, Grismer, Ahmad, Onn & Bauer, 2009: 30

M Muin Type locality: Bukit Larut (04º51.446'N, 100º48.455'E), , Peninsular Malaysia Holotype: ZRC 2.6804 Distribution: West Malaysia: Perak

Acanthosaura brachypoda Ananjeva, Orlov, Nguyen & Ryabov, 2011 Acanthosaura brachypoda Ananjeva, Orlov, Nguyen & Ryabov, 2011:196 Type locality: Fan Si Pan Mount village of Ban Khoan (22°12’21” N, 103°27’41” E), 12 km NW of Sa Pa town, near O Qui Ho Pass, Lao Cai province, Vietnam. Holotype: ZISP 26273. Distribution: northwestern Vietnam

Acanthosaura capra Günther, 1861 Günther, 1861:188

Nabokov Type locality: Lectotype: BMNH 1946.8.13.90, designated by Orlov 2006 Distribution: Cambodia, south Vietnam

Acanthosaura cardamomensis Wood, Grismer, Grismer, Neang, Chav & Holdnen, 2010 Acanthosaura cardamomensis Wood, Grismer, Grismer, Neang, Chav & Holdnen, 2010: 25 Type locality: Camp 2 (12°01’N, 104°09’E), Phnom Aural Kampong Speu Province, Cambodia. Holotype: LSUHC 7447 Distribution: Cambodia: Cardamom Mountains; Thailand; Vietnam: Phu Quoc Island

Acanthosaura coronata Günther, 1861 Günther, 1861: 187 D Melnikov Type locality: Cambodia Syntypes: BMNH 1946.8.13.81-83. BMNH 1946.8.19.64, 1946.8.13.84 Distribution: south Vietnam: Lam Dong and Dong Nai provinces; Cambodia

Acanthosaura crucigera Boulenger, 1885 Boulenger, 1885: 302 Synonym Acanthosaura horrescens Lönnberg,1916: 5 Type locality: Loi Koon Tan slopes, Bang Hue Pong, Thailand. Syntypes: NRM 3244-45 Type locality: Tavoy, Tenasserim, Burma Syntypes: BMNH 1946.8.19.64, 1946.8.13.84 Distribution: Burma, Thailand, West Malaysia, Cambodia

Acanthosaura lepidogaster (Cuvier, 1829) Calotes lepidogaster Cuvier, 1829: 39 Synonyms Acanthosaura lamnidentata Boulenger, 1885: 302 Type locality: Pegu, Tenasserim, Burma Syntypes: BMNH 1868.4.3.59, 1882.4.14.6 Acanthosaura hainanensis Boulenger, 1900: 957 Type locality: Five Finger Moutains (=Mtzhi), Hainan Island, Hainan Prov, China Syntypes: BMNH 1868.4.3.59, 1882.4.14.6 Acanthosaura fruhstorferi Werner, 1904: 461 Type locality: Tonkin (=northern Vietnam) Lectotype: NMW 11411, designated by Hallermann 2000 Acanthosaura braueri Vogt, 1914: 2 Type locality: Canton, Guangdong, southern China Type: ZMB 24178 Type locality: south Vietnam Syntypes: MHNP 0.5076, 0.6895 Distribution: northern Thailand; Vietnam: Ha Tinh, Lao Cai, Ha Giang, Tuyen Quang, Cao Bang, Bac Can, Lang Son, Son La, Vinh Phu, Ha Tay, Hoa Binh, Quang Ninh, Thai Ngyen, Ha Tay, Ninh Binh, Nghe An, Ha Tinh, Quang Binh, Quang Tri, Thua Thien-Hue, Da Nang, Quang Nam, and Kon Tum Provinces; Burma; ; southern China: Yunnan, Guangxi, Guangdong, Fujian, Hainan

Acanthosaura murphyi Nguyen, Do, Hoang, Orlov, Nguyen, Nguyen, 2018 Acanthosaura murphyi Nguyen, Do, Hoang, Orlov, Nguyen, Nguyen 2018: 262 Type locality : Hon Ba NR (ca. 12°07.308N, 108°57.896E), Khanh Hoa Province, Vietnam Holotype : ITBCZ 3533 Distribution : Vietnam: Khanh Hoa, Phu Yen and Khanh Khoa provinces

Nguyen

Acanthosaura nataliae Orlov, Truong & Sang, 2006 Acanthosaura nataliae Orlov, Truong & Sang, 2006:62

T Ngo Type locality: Tram Lap Village (14°26’39’’N, 108°32’97’’E), 40 km northwest of Kannack Town, Truong Son (Annamite) Mountains, An Khe District, Gia Lai Province, Vietnam. Holotype: ZISP 23430 Distribution: central Vietnam; southern Laos

Acanthosaura phongdienensis Nguyen, Jin, Vo, Nguyen, Zhou, Che, Murphy & Zhang, 2019 Acanthosaura phongdienensis Nguyen, Jin, Vo, Nguyen, Zhou, Che, Murphy & Zhang, 2019: 559 Type locality : Phong Dien Nature Reserve (16°28’ 03.6” N, 107° 16’ 40.8” E), Thua Thien-Hue Province, Vietnam Holotype : KIZ 10695 Distribution: Vietnam: Truong Son Mountains

Acanthosaura phuketensis Pauwels, Sumontha, Kunya, Wood & Grismer, 2015 Acanthosaura phuketensis Pauwels, Sumontha, Kunya, Wood & Grismer, 2015: 475 Type locality: Ban Bangrong (08°02.465N, 98°23.588E), Thalang District, Phuket Island, Phuket Province, southwestern Thailand. Holotype: THNHM 22542 Distribution: Phuket Range: Phuket, Phang-Nga, Surat Thani and Ranong provinces, Thailand

Acanthosaura titiwangsaensis Wood, Grismer, Grismer, Ahmad, Onn & Bauer, 2009 Acanthosaura titiwangsaensis Wood, Grismer, Grismer, Ahmad, Onn & Bauer, 2009: 36 Type locality: Bishop’s Trail, Fraser’s Hill (3º43.230'N; 101º44.887'E), , Peninsular Malaysia Holotype: ZRC 2.6805 Distribution: West Malaysia, Pahang

Acanthosaura tongbiguanensis Liu & Rao, 2019 Acanthosaura tongbiguanensis Liu & Rao, 2019: 111 Type locality : Tongbiguan Township (24°36'51.24"N, 97°35'1.88"E), Yingjiang County, Dehong Autonomous Prefecture, Yunnan, China. Holotype : KIZ L201804 Distribution: China: border region with northern in western Yunnan

Agama Daudin, 1802 Daudin, F.M. 1802. Histoire Naturelle, Générale et Particulière des Reptiles , Vol. 3. F. Dufart, Paris Type species: Lacerta agama Linnaeus, by subsequent designation

Agama aculeata Merrem, 1820

H Hillewaert

Subspecies: aculeata Merrem, 1820 Agama aculeata Merrem, 1820: 53 Synonyms Agama infralineata Peters, 1877:620 Type locality: Otjimbingue, Namibia Syntypes: ZMB 4217, 54518-21 Agama pulchella Bocage 1896: 116 Type locality: Modder River, Orange Free State, South Africa Type: MBL, lost to fire Type locality: Cape of Good Hope, South Africa Holotype: ZMB 750 Distribution: western South Africa, Namibia, Botswana, southern Angola, Zambia

Agama aculeata distanti (Boulenger, 1902) Agama distanti Boulenger 1902: 339 Syntypes: BMNH 1946.8.27.98-99 Type locality: Delagoa Bay, Mozambique Distribution: eastern South Africa, Zimbabwe, Mozambique, Tanzania, Swaziland

Agama africana (Hallowell, 1844) Tropidolepis africanus Hallowell, 1844: 171 Synonyms Oreodeira gracilipes Girard, 1857: 199 Type locality: unknown Holotype: USNM 6307 Agama savattieri Rochebrune, 1884: 89 Type locality: Banjul, Gambia Syntypes: MNHP 1962.112-114 Type locality: Liberia Holotype: ANSP 7361 Distribution: Liberia, Guinea, Ivory Coast, Ghana, Gambia

Agama agama (Linaeus, 1758) Lacerta agama Linnaeus, 1758: 207

M Pickersgill Synonym Agama colonorum Duméril & Bibron, 1837: 489 Type locality: probably Guinea Syntypes: MNHP, not located Type locality: “America” (in error for the coast of West Africa, probably Senegal or Ghana Lectotype: ZMUU 32, designated by Mediannikov et al. 2012 Distribution: Benin, Burkina Faso, Cameroon, Cape Verde Islands, Chad, Gabon, Ghana, Guinea, Guinea Bissau, Liberia, Mali, Mauritania, Nigeria, Senegal, Togo.

Agama anchietae Bocage, 1896 Bocage, 1896: 129 Synonyms Agama mehelyi Tornier, 1902: 672 Type locality: Bipindi , Cameroon Holotype: ZMB 18149 Agama namaquensis Werner, 1910: 320 Type locality: Namibia Syntypes: ZMB 22998, 55937 Agama anchietae methueni Boulenger & Power, 1921: 269 Type locality: Great Karas Mountains, Namibia Syntypes: BMNH 1936.8.1.373-375 (three specimens), MMK (two specimens), SAM (one specimen), TMP 3108, 3112, 3132, 3142 (four specimens). Type locality: Angola. Syntype: BMNH 1946.8.27.97 Distribution: southern Congo; Angola; southern DR Congo; Namibia; South Africa: northwest Cape; Botswana

Agama armata Peters, 1855 Agama armata Peters, 1855: 616

D Robertson Type locality: Rios de Sena, Tete, Mozambique Syntypes: ZMB 4244-48, 9199 Distribution: South Africa, Mozambique, Namibia, Botswana, Zambia, Swaziland, southern DR Congo, southwestern Kenya, central Tanzania, Zimbabwe

Agama atra Daudin, 1802 Daudin, 1802:349

A Shawka Synonyms Agama micropholis Matschie, 1891:607 Type locality: South Africa Lectotype: ZMB 42887, designated by Denzer et al. 1997 Agama microterolepis Boulenger, 1896: 22 Type locality: Rustenburg, Transvaal, South Africa Syntypes: TMP, lost Agama atra rudis Boulenger & Power, 1921: ? Type locality: Port Elizabeth Syntypes: BMHN 1946.9.4.69, 1946.9.4.70-71, 1946.8.27.89 -91 Type locality: South Africa Syntypes: MNHN 0.2604-05 Distribution: South Africa, Swaziland, southern Namibia, southern Botswana

Agama bocourti Rochebrune, 1884 Agama bocourti Rochebrune, 1884: 91 Type locality : Mélacorée River, Casamence, Gambia Type: animal depicted in Rochebrune 1884, plate XI, figs 3, 4, apparently lost Distribution: Senegal, Gambia

Agama boensis Monard, 1940 Agama boensis Monard, 1940:155 Type locality: Madina Boé, Guinea Bissau Syntypes: MHNCF 865, 867-868 Distribution: Guinea-Bissau, Guinea, Mali, Senegal

Agama bottegi Boulenger, 1897 Agama bottegi Boulenger, 1897: 717 Type locality: Lugh ( 03°47’46’’ N, 42°32’32’’ E), Somaliland (= Somalia) Holotype: MSNG 28548 Distribution: Somalia

Agama boueti Chabanaud, 1917 Agama boueti Chabanaud, 1917: 85 Synonym Agama castroviejoi Padial, 2005: 28 Type locality: Dahr Chinguetti, Adrar plateau, road from Atar to Tidjikja, Mauritania Holotype: MNCN 41779 Type locality : Gao, Niger Syntypes: MHNP 1917.44-45 Distribution: Mali, southern Mauritania: Assaba, Hodh Ech Chargui, Hodh El Gharbi, Trarza; northwestern Senegal, Niger, Burkina Faso, southern Morocco, probably in Chad

Agama boulengeri Lataste, 1886 Lataste, 1886: 212

M O’Shea Type locality: Vallon des Singes, à Médine, Haut Sénégal. Holotype: BMNH 1946.8.27.73. Distribution: Mali, Mauritania, Western Sahara

Agama caudospinosa Meek, 1910 Subspecies caudospinosa Meek, 1910 Agama caudospinosa Meek, 1910: 407 Type locality : Lake Elmenteita, Kenya Holotype : FMNH 2312 Distribution: Kenya

Agama caudospinosa spawlsi Wagner, 2010 Agama caudospinosa spawlsi Wagner, 2010: 37 Type locality: Nkunga (00°06’73N, 37°35’77E), north of Meru, Kenya Holotype : ZFMK 83666 Distribution Kenya: Mount Kenya

Agama congica Peters, 1877 Agama colonorum congica Peters, 1877: 612 Type locality : Chinchoxo (05°06’S, 12°06’E), Cabinda, Angola Syntypes: ZMB 9169 Distribution: Angola

Agama cristata Mocquard, 1905 Agama cristata Mocquard, 1905: 288 Type locality: Bomanesco (Sankaran) in Guinea Holotype : MNHP 1901.0394 Distribution: Guinea, Mali

Agama doriae Boulenger, 1885 Subspecies: Agama doriae doriae Boulenger, 1885 Agama doriae Boulenger, 1885: 127 Synonyms Agama sennariensis Werner, 1914: 478 Type locality: Sennaar on the Blue Nile, eastern Syntypes: NMHW 23471:1-2 Agama cordofanensis Werner, 1919: 478 Type locality:. between Ras-el-Fil and Gulfan, Chad Syntypes: NMHW 17038:1 ; NMHW 17038:2 (Sungikai , Sudan); NMHW:3 (Talodi, Sudan); NMHW 17038.4 (Kadugli, Sudan) Type locality: Keren, country of the Bogos, northern Ethiopia Syntypes : BMNH 1946.8.27.80-86 Distribution: Ghana, Togo, Nigeria, northern Cameroon, Central African Republic to Eritrea and Ethiopia, Sudan

Agama doriae benueensis Monard, 1951 benueensis Monard, 1951: 131 Lectotype: MHNCF 961, designated by Grandison 1968. Type locality : Ngnouyanga, upper Benue, Camroon Distribution: Nigeria, Cameroon, Ghana

Agama etoshae McClachlan, 1981 McClachlan, 1981: 223

L Raw Type locality: Onoholongo (18° 28’ S, 15° 43’ E), Kaokoveld, Namibia Holotype: SAM-ZR 16624 Distribution: northern Namibia

Agama finchi Böhme, Wagner, Malonza, Lötters & Köhler, 2005 Subspecies: Agama finchi finchi Böhme, Wagner, Malonza, Lötters & Köhler, 2005 Agama finchi Böhme, Wagner, Malonza, Lötters & Köhler, 2005: 144 Type locality: km 11 on road from Malaba to Busia (00°34 N, 34°11’E), Western Province, Kenya Holotype: NMK L25342 Distribution: western Kenya, Ethiopia

Agama finchi leucerythrolaema Wagner, Freund, Modrý, Schmitz & Böhme, 2011. Agama finchi leucerythrolaema Wagner, Freund, Modrý, Schmitz & Böhme, 2011: 27 Type locality: Murchison Falls, Uganda Holotype: ZFMK 88809 Distribution: western Uganda, DR Congo: Ituri

Agama gracilimembris Chabanaud, 1918 Agama gracilimembris Chabanaud, 1918: 106 Type locality: Dahomey (= Benin) Syntypes: MNHP 1904.114–115 Distribution: Ghana, Benin, Nigeria, Cameroon, Central African Republic, probably in Chad, Mali, Guinea, Burkina Faso

Agama hartmanni Peters, 1869 Agama hartmanni Peters, 1869: 65 Synonym Agama cornii Scortecci, 1928: 318 Type locality: Um Ager, Missione Corni, Calciati, Eritrea Type: MSNM 2258 Type locality: Dongola, Sudan Holotypes: ZMB 4355 Distribution: , Eritrea

Agama hispida (Linnaeus, 1758)

M O’Shea Subspecies hispida (Linnaeus, 1758) Lacerta hispida Linnaeus, 1758: 205 Type locality : ‘South America’, obviously incorrect, probably Cape of Good Hope Holotype: NRM 498 Distribution: South Africa, Zimbabwe, Namibia, southern Angola, Botswana, Mozambique, Malawi

Agama hispida makarikarika Fitzsimons, 1932 Agama hispida makarikarika Fitzsimons, 1932: 36 Holotype: TMP 144591. Type locality: Makarikari Pan, North Bechuanaland (=northeastern Botswana) Distribution: northeastern Botswana

Agama hulbertorum Wagner, 2014 Agama hulbertorum Wagner, 2014: 194 Type locality: Type locality: Elangata Wuas (1°52’18.7’’S, 36°33’46.6’’E), 26 km north of Kajiado, south-central Kenya Holotype: CAS 198880 Distribution: south central Kenya

Agama impalearis Boettger, 1874 Agama colonorum impalearis Boettger, 1874: 132 Synonyms Agama colonorum fritschi Boettger, 1874: 134 Type locality: Marocco Holotype: SMF 10117 Agama bibronii Boulenger, 1885: 357 Type locality:. Marocco Holotype: MNHP 0.6929 Type locality: Marocco Lectotype: SMF 10118, designated by Mertens 1967 Distribution: Morocco, Western Sahara, northwestern Algeria, and probably Mauritania

Agama insularis Chabanaud, 1918 Agama insularis Chabanaud, 1918: 161 Type locality: Rooma Island, Guinea Syntypes: MNHP 1918.41- 42 Distribution: endemic to Rooma Island, Guinea

Agama kaimosae Loveridge, 1935 Agama agama kaimosae Loveridge, 1935: 56 Type locality: Lake Kaimosi, Kenya Holotype: MCZ 40136 Distribution: Kenya

Agama kirkii Boulenger, 1885 Subspecies: Agama kirkii kirkii Boulenger, 1885 Agama kirkii Boulenger, 1885: 354 Type locality: Zambesi River, Holotype: BMNH 1947.3.6.50 Distribution: Malawi, Zambia, Zimbabwe, Mozambique, eastern Botswana

Agama kirkii fitzsimonsi Loveridge, 1950 Agama kirkii fitzsimonsi Loveridge, 1950: 128 Holotype: MCZ 44542. Type locality: Changadzi River, Birchenough Bridge, Southeast Rhodesia (=Zimbabwe) Distribution: Zimbabwe

Agama knobeli Boulenger & Power, 1921 Agama anchietae knobeli 1921: 271 Type locality: Aus, Great Namaqualand, Nambia Syntypes: BMNH 1946.8.27.94-96 Distribution: Namibia, mountains between Lüderitz Bay and Keetmanshoop

Agama lanzai Wagner, Leache, Mazuch & Böhme, 2013 Agama lanzai Wagner, Leache, Mazuch & Böhme, 2013:370 Type locality: Karin (10°58 ′26.4 ′′ N, 49°12 ′49.2 ′′ E; 325 m), Bari region, Puntland State, Somalia Holotype: CAS 227496 Distribution: Somalia

Agama lebretoni Wagner, Barej & Schmitz, 2009 Agama lebretoni Wagner, Barej & Schmitz, 2009: 291 Type locality: north-east of Mamfe (05°53.866’N, 009°33.495’E), , Mukwecha, Amebisu, Cameroon. Holotype: ZFMK 87698 Distribution: Cameroon, Equatorial Guinea, Gabon, Fernando Poo (Bioko Island), Nigeria

Agama lionotus Boulenger, 1896

S Zozaya Subspecies: Agama lionotus lionotus Boulenger, 1896 Agama agama lionotus Boulenger, 1896: 214 Type locality: southeast of Lake Rudolf, Gallaland (= Kenya) Holotype: BMNH 1946.8.28.8 Distribution: Uganda, Kenya, Ethiopia

Agama lionotus dodomae Loveridge, 1923 Agama lionatus dodomae Loveridge, 1923: 944 Synonym Agama lionotus usambarae Barbour & Loveridge, 1928: 150 Type locality: Soni, near Lusotho at the Usambara Mountains, Tanzania. Holotype: MCZ 24129 Syntypes: BMNH 1935.12.1.107-108. Type locality: Gwao, Dodoma, Ugogo, Tanganyika Territory (=Tanzania) Distribution: Tanzania

Agama lionotus elgonis Lönnberg, 1922 Agama elgonis Lönnberg, 1922: 2 Syntypes: MCZ 32798, 41009. Type locality: Mount Elgon, Kenya Distribution: Uganda, Kenya, Tanzania

Agama lionotus ufipae Loveridge, 1932 Agama agama ufipae Loveridge, 1932: 377 Holotype: MCZ 30741. Type locality: Kipili, Ufipa, Tanganyika Territory (=Tanzania) Distribution: western Tanzania, along Lake Tanganyika

Agama lucyae Wagner & Bauer, 2011 Agama lucyae Wagner & Bauer, 2011: 9 Type locality: Oromiya (7.27085ºN, 37.37891ºE), Omo River, 58 km NW of Sodo, Ethiopia Holotype: MCZ R-146689 Distribution: northern Ethiopia

Agama montana Barbour & Loveridge, 1928 montana Barbour & Loveridge, 1928: 147 Type locality: from rocks in a native shamba one mile below Bagilo, Uluguru Mountains, Tanzania Holotype: MCZ 24189 Distribution: Tanzania

Agama mossambica Peters, 1854 Agama mossambica Peters, 1854: 616

S Zozaya Synonyms Agama cariniventris Peters, 1874:159 Type locality: coast of Zanzibar, Tanzania Holotype: ZMB 7916 Type locality: Coast of Mozambique Syntypes: ZMB 4801, 7916, 9197, 9198; ZISP 5306 Distibution: Tanzania, Malawi, Zambia, Mozambique, eastern Zimbabwe

Agama mucosoensis Hellmich, 1957 Agama agama mucosoensis Hellmich, 1957: 44 Type locality: Mucoso, near Dondo, Angola Holotype: ZSM 117/53 Distribution: Angola: Mucoso, Dondo, and Libolo/Luati

Agama mwanzae Loveridge, 1923 Agama lionotus mwanzae Loveridge, 1923: 945 M Pickersgill Type locality: Shanwa, Mwanza, Tanganyika (=Tanzania) Syntypes: MCZ R-30670, BMNH 1923.10.9.27-30, 1946.8.27.67-68 Distribution: Tanzania, Rwanda, Kenya

Agama parafricana Trapé, Mediannikov & Trapé, 2012 Agama parafricana Trapé, Mediannikov & Trapé, 2012: 133 Type locality: Asrama forest (07°00’ N, 01°23’ E), Togo Holotype: MNHP 2010.0630 Distribution: Togo, Ghana, Benin

Agama paragama Grandison, 1968 Agama paragama Grandison, 1968: 75 Type locality: half mile southwest of Ahmadu Bello University, Zaria City, Nigeria Holotype: BMNH 1967.2215 Distribution: northern Nigeria, northern Cameroon, Mali, Central African Republic, Ghana, Gabon, western Chad

Agama persimilis Parker, 1942 Agama persimilis Parker, 1942: 52 Type locality: 45°50’ E, 8° N, Haud, British Somalia (= north-eastern part of Ogaden, of Ethiopia) Holotype: BMNH 1937.12.5.64 Distribution: Somalia, Ethiopia, northeastern Kenya

Agama picticauda Peters, 1877 Agama picticauda Peters, 1877: 612 Type locality: Ada Foah, Accra, Ghana Lectotype: ZMB 403, designated by Mediannikov et al. 2012 Disribution: Senegal, southern Mali, Burkina Faso, southern Niger, southern Chad, Guinea Bissau, Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, Benin, Nigeria, Gabon, northern DR Congo.

Agama planiceps Peters, 1862 Peters, 1862: 15

R Knight Type locality: New Barmen, Otjimbingue, Namibia Syntypes: ZMB 4200-4201, ZISP 1140 Distribution: Namibia: Damaraland, Kaokoveld

Agama robecchii Boulenger, 1891 Agama robecchii Boulenger, 1891: 6 Type locality: route from Obbia to Berbera, Somaliland Holotype: MSNG 31075 Distribution: northern Somalia, eastern Ethiopia

Agama rueppelli Vaillant, 1882 Subspecies: Agama rueppelli rueppelli Vaillant, 1882 Agama ruppelli Vaillant in Revoil, 1882: 6 Type locality: Bender-Meraya, Comalis country, Somalia Holotype: MNHP 0.5897 Distribution: Somalia Agama rueppelli occidentalis Parker, 1932 Agama rueppelli occidentalis Parker, 1932: 225 Type locality: mouth of the Kaliokwell River, Lake Rudolf, Kenya . Syntypes: BMNH 1946.8.27.74-78 Distribution: Ethiopia through northern Kenya Agama rueppelli septentrionalis Parker, 1932 Agama rueppelli septentrionalis Parker, 1932: 225 Type locality: Mount Njero (Njiro), Madago’s Village, Voi and Mbunji, Kenya Syntypes: BMNH 1935.12.3.1, MCZ R-18281 Distribution: eastern Kenya

Agama sankaranica Chabanaud, 1918 Agama sankaranica Chabanaud, 1918:105. Type locality: Moussaia, Sankaran, Mali Holotype: MNHP 1901.395 Distribution: Guinea, Nigeria, Ivory Coast, Ghana, Burkina Faso, Benin, Togo, Mali, Cameroon (?), Senegal, Niger

Agama smithi Boulenger, 1896 Agama smithii Boulenger, 1896: 213 Type locality: between [Webi] Shebeli and Juba Rivers (03°58 ′ N, 41°40 ′ E), Ethiopia. Holotype: BMNH 1946.8.27.79 Distribution: southern Ethiopia; southwestern Somalia

Agama somalica Wagner, Leaché, Mazuch &

Böhme, 2013 Agama somalica Wagner, Leaché, Mazuch & Böhme, 2013: 377 Type locality: Las Geel (09°44.914 ′N; 44°27.393 ′E; 1131 m), Central Somaliland, Somalia Holotype: ZFMK 94230 Distribution: northeastern Somalia

Tomas Mazuch

Agama spinosa Gray, 1831 Agama spinosa Gray, 1831: 57 Type locality: Suakin, Sudan Holotype: BMNH , lost Distribution: Egypt, Sudan, northern Ethiopia, Eritrea, Djibouti, northern Somalia

Agama sylvana MacDonald, 1981 Agama sylvana MacDonald, 1981: 191 Type locality: Krokosua forest (06°29’ N, 02°45’ W). Ghana Holotype: BMNH 1979.389 Distribution: Ghana, Gabon, Benin, Cameroon, Central African Republic

Agama tassiliensis Geniez, Padial & Crochet, 2011 Agama tassiliensis Geniez, Padial & Crochet, 2011: 32 Type locality: Tassili n’Ajjer (84 25.3500°N , 8.3911°E), 5 km south-south-west from Iherir, southeastern Algeria. Holotype: MNHP 2010.0632 Distribution: Mali: Adrar des Ifoghas; Niger: Aïr Mountains; Algeria: Ahaggar Mountains; Libya: Tassili n’Ajjer

Agama turuensis Loveridge, 1932 Agama agama turuensis Loveridge, 1932:376 Type locality: thorn bush country of Unyanganyi, east of Singida, Tanzania Holotype: MCZ 30676 Distribution: Tanzania: probably from the Kenyan Nguruman Escarpment through the Ngorongoro area to mountains in the Dodoma area

Agama wagneri Trapé, Mediannikov & Trapé, 2012 Agama wagneri Trapé, Mediannikov & Trapé, 2012: 137 Type locality: Mokolo (10°44’ N 13°47’ E), Cameroon Holotype: ZFMK 15222 Distribution: Cameroon

Agama weidholzi Wettstein, 1932 Agama weidholzi Wettstein, 1932: 304 Type locality: jungle wetland at Tabadienké, 30 km south of Diallakoto, Senegal Lectotype: NHMW 18318, designated by Wagner & Bauer 2011 Distribution: Senegal, Gambia, western Mali, Guinea-Bissau

Amphibolurus Wagler, 1830 Wagler, J.G. 1830. Natürliches System der Amphibien, mit vorangehender Classification der Säugetiere und Vögel. Ein Beitrag zur vergleichenden Zoologie. 1.0. Cotta, München, Stuttgart, and Tübingen, 354 pp . Type species: Lacerta muricata White, 1790, by original designation

Amphibolurus muricatus M Clancy

Amphibolurus burnsi (Wells and Wellington, 1985) Lophognathus burnsi Wells and Wellington, 1985: 18 Type locality : Collarenebri, New South Wales. Holotype : AMS R116981 Distribution : Australia: southern and central- western Queensland and northern inland New South Wales S Zozaya

Amphibolurus centralis (Loveridge, 1933) Physignathus gilberti centralis Loveridge, 1933 Type locality : Anningie 30 mi W of Teatree Well, Northern Territory. Holotype : MCZ 35207 Distribution : Australia: western Queensland, Northern Territory and Western Australia.

Amphibolurus muricatus (White, 1790) Lacerta muricata White, 1790: 244 Synonym Agama jacksoniensis Cloquet,1816: 74 Type locality: Australia Lectotype: RMNH 3117, designated by Cogger 1983 Type locality: New South Wales Holotype: BMNH 1946.9.4.44 Distribution: Australia: New South Wales, Queensland, South Australia, Victoria, Tasmania

Amphibolurus norrisi Witten & Coventry, 1984 Witten & Coventry, 1984: 155 Type locality: 29 km south of Nhill (36°36'S,141°38'E), Victoria, Australia Holotype: NMV D51499 Distribution: Australia: South Australia, Victoria, Western Australia

Aphaniotis Peters, 1864 Peters, W.C.H. 1864. Über einige neue Säugethiere (Mormops, Macrotus, Vesperus, Molossus, Capromys), Amphibien (Plathydactylus, Otocryptis, Euprepes, Ungalia, Dromicus, Tropidonotus, Xenodon, Hylodes), und Fische (Sillago, Sebastes, Channa, Myctophum, Carassius, Barbus, Capoeta, Poecillia, Saurenchelys, Leptocephalus). Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin , vol. 1864, p. 381-399 Type species: Otocryptis fusca Peters, 1864, by original designation

Aphaniotis acutirostris Modigliani, 1889 Aphaniotis acutirostris Modigliani, 1889: 115 Type locality: Nias Island, Indonesia Lectotype: MSNG 29001, designated by Capocaccia 1961 Distribution: Indonesia: Nimalur, Nias, Pulu Nako, Mentawei, Sumatra,

Aphaniotis fusca (Peters, 1864) Otocryptis fusca Peters, 1864: 385

M Muin Synonym Aphaniotis floweri Werner, 1900:486 Type locality: Sumatra Type: not located Type locality: Forest Hill, West Malaysia Holotype: ZMB 5032 Distribution: southern Thailand: Pattani, Yala; West Malaysia: Pulau Tioman, Johor, Pulau Sibu: Singapore: Indonesia: Simalur, Nias, Sumatra, Singkep, Borneo, Natuna Island

Aphaniotis ornata (Lidth de Jeude, 1893) Japalura ornata Lidth de Jeude, 1893: 251 Synonym Japalura nasuta De Jong, 1930: 115 Type locality: Sabah, East Malaysia Holotype : MZB, lost Type locality: Sandakan Bay, north Borneo, Indonesia Holotype: RMNH 4344 Distribution: Indonesia: Kalimantan; East Malaysia: Sabah, Sarawak

Brachysaura Blyth, 1856 Blyth, E. 1856. Proceedings of the Society. Report of the Curator. Journal of the Asiatic Society of Benga l 25: 448-449 Type species: Brachysaura ornata Blyth, 1856, by original designation

Brachysaura minor (Hardwicke & Gray, 1827) Agama minor Hardwicke & Gray, 1827: 218

G Bhardwa Synonym Brachysaura ornata Blyth,1856: 448 Type locality: Sangor, central India Holotype: Icontype: Hardwicke NHM archives No. 82 Type locality: Chittagong, southeastern Type: Iconotype: based on Hardwicke’s sketch, Brit. Mus. Coll, no. 82 Distribution: Pakistan: Sind, Punjab; India: Madhya Pradesh, Uttar Pradesh, Rajasthan, Gujarat, Orissa; Bangladesh

Bronchocela Kaup, 1827 Kuhl,H. 1820. Beiträge zur Zoologie und vergleichenden Anatomie. Hermannsche Buchhandlung, Frankfurt, 152 pp Type species: Agama cristatella Kuhl, 1820, by original designation

Bronchocela celebensis Gray, 1845 Bronchocela celebensis Gray, 1845: 241 Type locality: Celebes (= ), Indonesia Holotype: BMNH 1946.8.11.48 Distribution: Indonesia: Sulawesi

Bronchocela cristatella (Kuhl, 1820) Agama cristatella Kuhl, 1820: 108 L Kee Synonyms Agama gutturosa Merrem, 1820:51 Type locality: unknown Type: iconotype: based on Seba I, pl. 89, figs 1&2 Agama moluccana Lesson, 1830: pl. i Type locality: Buru, Maluku, Indonesia Syntypes: MNHP 1999.8206, 0.6886 Bronchocela burmana Blanford, 1878: 127 Type locality: Dawei, Taninthayi Division, southern Burma Holotype: ZSI 5337 Bronchocela intermedia Peters & Doria, 1878: 375 Type locality: Kandari, Sulawesi, Indonesia Holotype: MSNG 29109 Pseudocalotes archiducissae Steindachner, 1867: 27 Type locality: Nicobar Islands Type: lost Calotes octospinosus Baumann,1913: 265 Type locality: Sumatra, Indonesia Syntypes: NMBE 1023528, NMBE1018463-66 Type locality: not mentioned; probably Penang, Malaysia Syntypes: BMNH XXIV.21.a-b Distribution: West Malaysia: Pulau Tioman, Pulau Langkawi, Pulau Singa Besar, Johor: Pulau Sibu, Borneo), East Malaysia; southern Burma: Tenasserim; southern Thailand; Singapore; India: Andaman and Nicobar Islands; Philippines: Palawan, Calamian Islands, Panay, Luzon, Mindanao, Sulu, Cebu, Bohol, Leyte, Negros, Samar, Mindoro, Masbate; Indonesia: Kalimantan, Halmahera, New Guinea

Bronchocela danieli (TiwariI & Biswas, 1973) Calotes danieli TiwariI & Biswas, 1973: 57 S Harikishnan Type locality: Campbell Bay, Great Nicobar Island, India. Holotype: ZSI 22455 Distribution: India: Nicobar Islands

Bronchocela hayeki (Müller, 1928) Calotes hayeki Müller, 1928: 67 Type locality: near Berastagi (3°11’ N, 98° 31’ E), at the base of Sibayak, Sumatera Utara, Indonesia Neotype: ZMB 55931 Distribution: Indonesia: North Sumatra

Bronchocela jubata Duméril & Bibron, 1837 Duméril & Bibron, 1837: 397 Synonym Calotes intermedius Berthold, 1842: 28 Type locality: Java, Indonesia Syntypes: ZFMK 27099–101 Type locality: Java, Indonesia Syntypes: MHNP 0.2541, 2542, 2543 Distribution: Indonesia: Java, , Sulawesi, Karakelang, Salibabu; Nias Island, Singkap Island, Borneo (Kalimantan); Philippines (Mindanao)

Bronchocela marmorata Gray, 1845 Bronchocela marmorata Gray, 1845: 242 Synonyms Lophyrus spinosus Duméril, 1851: 91 Type locality: Zamboanga, Philippines Holotype: MNHP 0.6896 Calotes philippinus Peters, 1867: 16 Type locality: Daraga, Luzon, Philippines Syntypes: MSNG 29108; ZMB 5433, 5642 Bronchocela marmorata sanchezi Taylor, 1922: 138 Type locality: Polillo Island, Philippines Holotype: BSP 827, destroyed in WWII Type locality: Philippine Islands Holotype: BMNH 1946.8. 11.16 Distribution: Philippines: Luzon, Polillo, Mindoro, Catanduares, Sibuan

Bronchocela orlovi Hallermann, 2004 Bronchocela orlovi Hallermann, 2004: 30 Type locality: at Buoenloy (14°20’ N, 108°36’ E), Ankhe District, Gia Lai Province, southern Vietnam. Holotype: ZISP 22827 Distribution: south Vietnam: northern part of Tai Nguyen Plateau in the Annam Mountains Last seen 1983

Bronchocela rayaensis Grismer, Wood Jr, Quah, Anuar, Ngadi, 2015 Bronchocela rayaensis Grismer, Wood Jr, Quah, Anuar, Ngadi, 2015: 8 Type locality: Gunung Raya (06°22.096 N, 99°49.004 E), Pulau Langkawi, Kedah, West Malaysia. Holotype: LSUHC 7535 Distribution: Langkawi Island, Kedah, West Malaysia; Thailand: Phuket Island and Phang-nga Province

Bronchocela rubrigularis Hallermann, 2009 Bronchocela rubrigularis Hallermann, 2009: 280

S Harikishnan Type locality: Trinkat Island, central Nicobar Islands Holotype: ZMH R09271 Distribution: India: Nicobar Islands

Bronchocela shenlong Grismer, Wood Jr, Quah, Anuar, Ngadi, 2015 Bronchocela shenlong Grismer, Wood Jr, Quah, Anuar, Ngadi, 2015: 15 Type locality: Bukit Larut (04°51.715 N, 100°47.933 E), Perak, West Malaysia. Holotype: LSUHC 9017 Distribution: Bintang Mountain Range, Perak, and , Pahang in the Titiwangsa Mountain Range

Bronchocela smaragdina Günther, 1864 Bronchocela smaragdina Günther, 1864: 138 Type locality: Cambodia Syntypes: BMNH 1946.8.11.35-36 Distribution: Cambodia: Mondolkiri Province; central Vietnam; eastern Thailand

Bronchocela vietnamensis Hallermann & Orlov, 2005 Bronchocela vietnamensis Hallermann & Orlov, 2005: 176 Type locality: near Tram Lap village (14° 20’ N, 1`08° 36’ E), 40 km NW from Kannack, Annam mountains, Ankhe District, Gia Lai Province, Vietnam Holotype: FMNH 252295 Distribution: south Vietnam: Gia Lai Province

Bufoniceps Arnold, 1992 Arnold,E.N. 1992. The Rajasthan Toad-headed , Phrynocephalus laungwalaensis (Reptilia: Agamidae), represents a new genus. Journal of Herpetology 26 (4): 467-472 Type species: Phrynocephalus laungwalaensis Sharma, 1978, by original designation

Bufoniceps laungwalaensis (Sharma, 1978) Phrynocephalus laungwalaensis Sharma, 1978: 291

I Das Type locality: Laungwala ( 26°46’N, 70°13’E’) , Jaisalmer District, Rajasthan, India Holotype: ZSI 23452 Distribution: India: Rajasthan Desert, Jaisalmer District; Pakistan: Sindh

Calotes Cuvier, 1817 Cuvier, G. 1817. Le règne animal distribué d’après son organisation, pour servir de base a l’historire naturelle des animaux et d’introduction a l’anatomie comparée . Vol. 2. Les reptiles, les poissons, les mollusques et les annélides. Déterville, Paris Type species: Lacerta calotes Linnaeus, 1758, by tautology

Calotes bachae Hartmann, Geisserler, Poyarkov, Ihlow, Galoyan, Rödder & Böhme, 2013 Calotes bachae Hartmann, Geisserler, Poyarkov, Ihlow, Galoyan, Rödder & Böhme, 2013: 252 Type locality: Cat Tien National Park (11°38'04'' N 107°27'24'' E), Dong Nai Province, Vietnam Holotype: ZFMK 88935 Distribution: south Vietnam: Dong Nai, Binh Phuoc

Calotes bhutanensis Biswas, 1975 Calotes bhutanensis Biswas, 1975: 775 Type locality: Janjurmane, central Holotype: ZSI 22480 Distribution: central Bhutan

Calotes calotes (Linnaeus, 1758) Lacerta calotes Linnaeus 1758: 207 M Pickersgill Synonyms Agama ophiomachus Merrem, 1820: 51 Type locality: unknown Holotype: UUZM 33 Agama lineata Kuhl, 1820 : 108 Type locality: unknown type: unknown Type locality: Ceylon (=Sri Lanka) Holotype: NRM 106 Distribution: southern India: Travancore, Shevaroy Hills, ; Sri Lanka

Calotes ceylonensis Müller, 1887 Calotes mystaceus ceylonensis Müller, 1887: 292

I Das Synonyms Calotes kelaarti Nevill, 1887 :134 Type locality: northwest Sri Lanka Type: unknown Calotes saleoides Werner, 1896: 7 Type locality: Sri Lanka Holotype: NMHW 23356 Type locality: Kumbukan-aar, southeastern Ceylon Holotype: NHMB 3340 Distribution: Sri Lanka

Calotes chincollium Vindum, 2003 Calotes chincollium Vindum, 2003:3 Type locality: 21°23 ′11.2 ″N, 93°58 ′15.9 ″E, Min Dat Township, Min Dat District, Chin State, Burma Holotype: CAS 220009 Distribution: Burma

Calotes desilvai Bahir & Maduwage, 2005 Calotes desilvai Bahir & Maduwage, 2005: 382 Type locality: Morningside Forest (06°24’N, 80°36’E), Eastern Sinharaja, Sri Lanka Holotype: WHT 1832 Distribution: Sri Lanka

Calotes emma Gray, 1845

S Harikishnan Type locality: Afghanistan, in error Holotype: BMNH 1946.8.11.26 Subspecies: Calotes emma emma Gray, 1845 Calotes emma Gray, 1845: 244 Type locality: Afghanistan, in error Holotype: BMNH 1946.8.11.26 Distribution: peninsular Thailand south to Perak in Malaysia

Calotes emma alticristatus Schmidt, 1925 Calotes emma alticristatus Schmidt, 1925:482 Type locality: Yunnanfu, Yunnan, China Holotype: AMNH 17395 Distribution: southern China: Guangdong, Yunnan; India: Assam; Burma; Thailand; India: Tripura; Laos; Vietnam; Cambodia

Calotes grandisquamis Günther, 1875 Calotes grandisquamis Güunther, 1875: 226 J Jose Type locality: at the foot of the Canoot Ghat, near Manantoddy, Bramagherry Hills, India Syntypes: BMNH 1946.8.11.44-47 Distribution: India: Anaimalai, Bramagherry Hills, Travancore; southern Western Ghats

Calotes htunwini Zug & Vindum, 2006 Calotes htunwini Zug & Vindum, 2006: 50 Type locality: San Myaung Camp (23°34 ′27.6 ″N, 95°44 ′15.6 ″E), Chatthin Wildlife Sanctuary, approx. 2 km WNWof Chatthin, Burma Holotype: USNM 524044 Distribution: Burma

Calotes irawadi Zug, Brown, Schulte & Vindum, 2006 Calotes irawadi Zug, Brown, Schulte & Vindum, 2006: 54 Type locality: San Myaung Camp (23°34 ′27.6 ″N, 95°44 ′15.6 ″E), Chatthin Wildlife Sanctuary, approx. 2 km WNWof Chatthin, Burma Holotype: USNM 520543 Distribution: Burma

Calotes jerdoni Günther, 1870 Calotes jerdoni Günther, 1870: 778 S Harikishnan Synonyms Calotes platyceps Jerdon, 1870:77 Type locality: Khasi Hills, India Type: not found Calotes yunnanensis Annandale, 1905: 87 Type locality: Teng-yueh, (=Momien, Tengchong County, Yunnan Province, southern China). Holotype: ZSI 6571 Type locality: Khasi Hills, northeastern India Syntypes: BMNH 1946.8.11.49-58; ZMB 8448 Distribution: India: Khasi Hills in Assam & Shillong; Burma; Bhutan, China: Yunnan

Calotes liocephalus Günther, 1872 Calotes liocephalus Günther, 1872:86 Type locality: Peradeniya District, Ceylon (=Sri Lanka) Holotype: BMNH 1946.8.11.33 Distribution: Sri Lanka

Calotes liolepis Boulenger, 1885 Calotes liolepis Boulenger, 1885: 326

M O’Shea Type locality: Sri Lanka Syntypes: BMNH 1854.2.10.12, 1869.7.24.2 Distribution: Sri Lanka

Calotes manamendrai Amarasinghe, Karunarathna and Fujinuma, 2014 Calotes manamendrai Amarasinghe, Karunarathna and Fujinuma, 2014: 3 Type locality: Riverstone, Knuckles, Sri Lanka Holotype: NMSL 5976 Distribution: Knuckles massif, Sri Lanka

Calotes maria Gray, 1845 Calotes maria Gray, 1845: 243 Type locality: Kassi Hill (=Khasi Hills), Assam, India Holotype: BMNH 1946.8.11.24 Distribution: India: Khasi Hills in Assam

Calotes medogensis Zhao & Li, 1984 Calotes medogensis Zhao & Li, 1984: 77 Type locality: Yarang, Medog Co., Xizang, China Holotype: CIB 8370177 Distribution: Tibet

Calotes mystaceus Duméril & Bibron, 1837 Calotes mystaceus Duméril & Bibron, 1837: 408

M Pickersgill Type locality: Birmans, Burma Holotype: MNHP 0.2557 Distribution: China: Yunnan; Burma: Tenasserim to Mandalay; Thailand: north of the Istmus of Kra; Cambodia; south Vietnam; Laos

Calotes nemoricola Jerdon, 1853 Calotes nemoricola Jerdon, 1853: 471 S Harikishnan Type locality: foot of Coonoor Ghat, Nilgiri Hills, India Holotype: ZSI 6560 Distribution: southern India: Nilgiri Hills, southern Western Ghats

Calotes nigrilabris Peters, 1860 Calotes nigrilabris Peters, 1860: 183

I Das Type locality: Nuwara Eliya, Sri Lanka Holotype: NHMW 23355 Distribution: Sri Lanka

Calotes nigriplicatus Hallermann, 2000 Calotes nigriplicatus Hallermann, 2000: 156 Type locality: Ambon Island, Moluccas, Indonesia Holotype: ZFMK 26379 Distribution: Indonesia: Ambon Island Last seen 1885

Calotes paulus (Smith, 1935) Oriocalotes paulus Smith, 1935: 166 Type locality : Khasi Hills, India Lecotype: BMNH 1946.8.11.27 Distribution : India: Meghalaya State Khasi Hills

Calotes pethiyagodai Amarasinghe, Karunarathna & Hallermann, 2014 Calotes pethiyagodai . Amarasinghe, Karunarathna & Hallermann, 2014: 62 Type locality: near Midland Estate (7°31’N, 80°44’ E), Knuckles, Sri Lanka Holotype: WHT 6211 Distibution: Sri Lanka: Knuckles massif

Calotes versicolor (Daudin, 1802) I Das Subspecies: Calotes versicolor versicolor (Daudin, 1802) Agama versicolor Daudin, 1802:395 Synonyms Agama tiedemanni Kuhl, 1820: 109 Type locality: Pondicherry, India Type: not located Agama vultuosa Harlan, 1825: 296 Type locality: Calcutta Holotype: ANSP 7296 Agama indica Gray, 1827: 217 Type locality: Dum-Dum, Calcutta Type: Iconotype: Illustrations of Indian Zoology, chiefly selected from the collection of Major - General Hardwicke , t. 74 Calotes cristatus Jaquemont, 1844 pl. xi Type locality: India Holotype: MNHP 0.2554 Calotes viridis Gray, 1846: 429 Type locality: Madras Type, lost Calotes gigas Blyth, 1853: 648 Type locality: unknown Holotype: ZSI , not found Type locality: Pondicherry University Campus (12.029097° N, 79.850343°E), Kalapet, Pondicherry, India Neotype: NCBS AT102 Distribution: southeastern Iran; Afghanistan; Pakistan; Nepal; Bhutan; India: Tamil Nadu, Assam, Gujarat, Maharashtra, Himachal Pradesh, Andaman Island; Sri Lanka; Bangladesh; Burma; Thailand; West Malaysia; Vietnam; Cambodia; southern China: Yunnan, Guangdong, Guangxi, Hongkong, Hainan Island; Maldives; Indonesia: Sumatra; Singapore

Calotes versicolor farooqi Auffenberg & Rehmann, 1993 Calotes versicolor farooqi Auffenberg & Rehmann, 1993: 27 Type locality: rocky hillside, Shargal (34.3° N, 73.4° E), 20 km S Balakot, Manshera Dist., Northwest Frontier Province, Pakistan Holotype: FMNH 79470. Distribution: alpine Punjab, Pakistan

Calotes zolaiking Giri, Chaitanya, Mahony, Lalroungas, Lalrinchhana,2019 Calotes zolaiking Giri, Chaitanya, Mahony, Lalroungas, Lalrinchhana,2019: 466 Type locality : Synod Hospital Compound (23°46 ´19.06 ´´ N, 92°43 ´56.37 ´´ E), Durtlang, Aizawl District, Mizoram state, Northeast India Holotype : NCBS-AU152 Distribution : India: Mizoram state: Hmuifang forest

Ceratophora Gray, 1835 Gray. J. E. 1835. Illustrations of Indian Zoology, chiefly selected from the collection of Major - General Hardwicke . Vol. 2. London (1833-1834): 263 pp., 95 plates Type species: Ceratophora stoddartii Gray, 1834, by original designation

Ceratophora aspera Günther, 1864 Ceratophora aspera Günther, 1864:131 Type locality: Sri Lanka Syntypes: BMNH 1946.8.30.51-52; ZMB 5120-21 Distribution: Sri Lanka

Ceratophora erdeleni Pethiyagoda & Manamendra-Arachchi, 1998 Ceratophora erdeleni Pethiyagoda & Manamendra-Arachchi, 1998: 11 Type locality: mountains in east Sinharaja Forest (06°24’N, 80°38’E), Morningside Forest Reserve, near Rakwana, Sri Lanka Holotype: BMNH 1996.448 Distribution: Sri Lanka

Ceratophora karu Pethiyagoda & Manamendra-Arachchi, 1998 Ceratophora karu Pethiyagoda & Manamendra-Arachchi, 1998:18 Type locality: Morningside Forest Reserve (06°24’N, 80°38’E), near Rakwana, Sri Lanka Holotype: BMNH 1996.445 Distribution: Sri Lanka Last seen 1996

Ceratophora stoddartii Gray, 1834 Ceratophora stoddartii Gray, 1834: 68 Type locality: Sri Lanka Holotype: BMNH 1946.8.27.37 Distribution: Sri Lanka

I Das

Ceratophora tennentii Günther, 1861 Ceratophora tennentii Günther, 1861: 281

I Das Type locality: Sri Lanka Holotype: BMNH 1946.8.27.32 Distibution: Sri Lanka

Chelosania Gray, 1845 Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum . Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. Type species: Chelosania brunnea Gray, 1845, by original designation

Chelosania brunnea Gray, 1845 Chelosania brunnea Gray, 1845: 245 S Macdonald Type locality: Western Australia Holotype: BMNH 1946.8.11.43 Distribution: Australia: North Territory, Queensland, Western Australia

Chlamydosaurus Gray, 1825 Gray, J.E. 1825. A synopsis of the genera of reptiles and Amphibia, with a description of some new species. Annals of Philosophy , 10:193—217 Type species: Chlamydosaurus kingii Gray, 1825, by original designation

Chlamydosaurus kingii Gray, 1825 Chlamydosaurus kingii Gray, 1825: 197

S Zozaya Type locality: Port Nelson, northwest coast of Australia Holotype: presumed lost Distribution: Australia: North Territory, Queensland, Western Australia; southern Western Province, Papua New Guinea; southeastern southeastern Papua Province, Indonesia

Complicitus Manthey & Grossmann, 1997 Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens . Natur und Tier Verlag (Münster), 512 pp Type species: Calotes nigrigularis Ota & Hikida, 1991, by by original designation

Complicitus nigrigularis (Ota & Hikida, 1991) Calotes nigrigularis Ota & Hikida, 1991: 185 M Islam Type locality: Kinabalu Park (06°00-15’N; 116°15-45’E), Sabah, East Malaysia Holotype: KUZ 13223 Distribution: East Malaysia: Sabah (=Borneo)

Cophotis Peters, 1861 Peters, W.C. H. 1861. On a new genus of lizards from Ceylon. Annals and magazine of natural history 1861: 419-420 Type species: Cophotis ceylanica Peters, 1861, by original designation

Cophotis ceylanica Peters, 1861 Cophotis ceylanica Peters, 1861: 1103

I Das Type locality: Sri Lanka Lectotype: ZMB 4240, designated by Marx 1958 Distribution: Sri Lanka

Cophotis dumbara Samarawickrama, Ranawana, Rajapaksha, Ananjeva, Orlov, Ranasinghe & Samarawickrama, 2006 Cophotis dumbara Samarawickrama, Ranawana, Rajapaksha, Ananjeva, Orlov, Ranasinghe & Samarawickrama, 2006: 2 Type locality: Cardamon cultivated area, Dothalugala Man and Biosphere Reserve (07°20’31.4”N, 80°50’25.6”E), The Knuckles Forest Range, Central Province, Sri Lanka Holotype: NMSL 2006850

Coryphophylax Blyth, 1860 Blyth, E. 1860. Of reptiles ... [In: Report of the curator, Zoological department, pp. 87-115] Journal of the Asiatic Society of Bengal 29: 107-111 Type species: Coryphophylax maximiliani Fitzinger, 1861, by subsequent designation

Coryphophylax brevicaudus Harikrishnan, Vasudevan, Chandramouli, Choudhuery, Dutta & Das, 2012 Coryphophylax brevicaudus Harikrishnan, Vasudevan, Chandramouli, Choudhuery, Dutta & Das, 2012: 35

S Harikishnan Type locality: Mount Harriet National Park (92.735944°N, 11.710579°E) South Andaman Island, Bay of Bengal, India Holotype: ZSI 25962 Distribution: India: Andaman Islands

Coryphophylax subcristatus (Blyth, 1860) Tiaris subcristata Blyth, 1860:109 S Harikishnan Synonyms Coryphophylax maximiliani Fitzinger, 1861 :30 Type locality: Nicobar Islands Holotype: NMHW, not found Tiaris humei Stoliczka, 1873:167 Type locality: Tillanchong Island (08º 30'N; 93º 37'E), Nicobar Islands, Bay of Bengal, India Holotype: ZSI 50412 Type locality: Port Blair" (11º36'N; 92º45'E), in South Andaman Islands, Bay of Bengal, India Syntypes: ZSI 5191–92, ZSI 5369–70, ZSI 5199–202 Distribution: India: Andaman Island, Nicobar Island

Cristidorsa Wang, Deepak, Datta-Roy, Lin, Jiang, Che & Siler, 2018 Wang, K., J. Che, S. Lin, V. Deepak, D. Aniruddha, K. Jiang, J. Jin, H. Chen and C.D. Siler. 2018. Multilocus Phylogeny and Revised Classification for Mountain Dragons of the Genus Japalura s.l. (Reptilia: Agamidae: Draconinae) from Asia. Zoological Journal of the Linnean Society 20: 13 Type species: Cristidorsa otai (Mahony, 2009).

Cristidorsa otai (Mahony, 2009) Japalura otai Mahony, 2009: 43 Rohit Naniwadekar Type locality : Sairep (22°49'0 N, 92°49'0 E), Lunglei District, central Mizoram, northeast India). Holotype: ZSI 25217 Distribution : northeastern India: Mizoram state Last seen 1995

Cristidorsa planidorsata (Jerdon, 1870) Japalura planidorsata Jerdon, 1870: 76 Type locality : Khasi Hills, India Syntypes : BMNH 1946.8.30.86–87 Distribution: Burma: Chin state; India: Assam, Meghalaya

Cryptagama Witten, 1984 Witten, G.J 1984. Relationships of Tympanocryptis aurita . Records of the Western Australian Museum 11 (4): 399-401 Type species: Tympanocryptis aurita Storr, 1981, by original designation

Cryptagama aurita (Storr, 1981) Tympanocryptis aurita Storr, 1981: 599

Stephen Mahony Type locality: 27 km SSE of Halls Creek (18°27'S, 127°45'E), Western Australia Holotype WAM R66296 Distribution: Australia: Northern Territory, Western Australia

Ctenophorus Fitzinger, 1843 Fitzinger, L.J. 1843. Systema Reptilium . Vienna : Braümüller u. Seidel vi 106 pp. Type species: Grammatophora decresii Duméril & Bibron, 1837, by original designation

Ctenophorus adelaidensis (Gray, 1841) Grammatophora muricata adelaidensis Gray, 1841: 88 Type locality: Swan River, Western Australia Syntypes : BMNH 1846.9.4.45–57 Distribution: Australia: coastal southwestern Western Australia

Ctenophorus butlerorum (Storr, 1977) Amphibolurus parviceps butleri Storr: 77 Type locality: near east shore of Useless Inlet (26°18'S 113°21'E), 29 km NW of Carrarang, Western Australia Holotype: WAM R54728 Distribution: Australia: Edel Land, Western Australia

Ctenophorus caudicinctus (Günther, 1875) Subspecies: caudicinctus (Günther, 1875) Grammatophora caudicincta Günther, 1875: 19

S Zozaya Synonym Amphibolurus imbricatus Peters, 1876: 529 Type locality: Mermaid Strait, Western Australia Syntypes: ZMB 8933 Type locality: Nickol Bay, Western Australia Holotype: BMNH 1946.9.4.43 Distribution: Australia: Pilbara region and offshore islands, West Australia

Ctenophorus caudicinctus graafi (Storr, 1967) Amphibolurus caudicinctus graafi Storr, 1967: 51 Type locality: Mt. Eveline, in 26° 10’ S, 127° 06’ E, Western Australia Holotype: WAM R25914 Distribution: Australia: far eastern interior of Western Australia

Ctenophorus caudicinctus infans (Storr, 1967) Amphibolurus caudicinctus infans Storr, 1967: 51 Holotype: WAM R25945 Type locality: Deeba Rock-Hole (28° 22’ S, 122° 35’E), 25 mi NE of Laverton, Western Australia Distribution: Australia: Mt. Margaret Goldfield, Western Australia

Ctenophorus caudicinctus macropus (Storr, 1967) Amphibolurus caudicinctus macropus Storr, 1967: 53 Holotype: SAMA R3229 Type locality: 4. 5 mi SSE of Oenpelli, Northern Territory, Australia Distribution: Australia: Arnhem Land, Northern Territory

Ctenophorus caudicinctus mensarum (Storr, 1967) Amphibolurus caudicinctus mensarum Storr, 1967: 51 Holotype: WAM R19486 Type locality: 5 mi S of Meekatharra, in 26° 40’ S, 118° 27’ E, Western Australia Distribution: Australia: Murchison and East Murchison Goldfields, Western Australia

Ctenophorus caudicinctus slateri (Storr, 1967) Amphibolurus caudicinctus slateri Storr, 1967: 52 Holotype: WAM R26793 Type locality: Hermannsburg, in 23'58’ S, 132° 46’ E, Northern Territory, Australia Distribution: Australia: highlands of southern Northern Territory

Ctenophorus chapmani (Storr, 1977) Amphibolurus adelaidensis chapmani Storr, 1977: 80 Type locality: 11 km SSE of Cocklebiddy (32°08'S, 126°08'E), Western Australia Holotype: WAM R24657 Distribution: Australia: coastal Western Australia and coastal South Australia

Ctenophorus clayi (Storr, 1967) Amphibolurus clayi Storr, 1967: 24 Type locality: Maryvale (24°40'S 134°04'E), Northern Territory, Australia Holotype: WAM R14462 Distribution: Australia: Northern Territory, South Australia, Western Australia

Ctenophorus cristatus (Gray, 1841) Grammatophora cristata Gray, 1841: 87 Type locality: Western Australia Holotype: BMNH 1946.9.4.41 Distribution: Australia: South Australia, Western Australia

Ctenophorus decresii (Duméril & Bibron, 1837) Grammatophora decresii Duméril & Bibron, 1837: 472

M Clancy Synonym Amphibolurus modestus Ahl, E. 1926: 187 Type locality: Australia Syntypes: ZMB 30092 Type locality: Kangaroo Island, South Australia Syntypes: MHNP 0.6545 Distribution: Australia: New South Wales, South Australia

Ctenophorus femoralis (Storr, 1965) Amphibolurus femoralis Storr, 1965: 49 S Zozaya Type locality: 23 mi NE of Giralia (22°29'S 114°33'E), Western Australia Holotype: WAM R9008 Distribution: Australia: Western Australia: Exmouth Gulf

Ctenophorus fionni (Procter, 1923) Amphibolurus fionni Procter, 1923:1075 Type locality: Port Lincoln, South Australia Holotype: BMNH 1946.9.4.67 Distribution: Australia: coast of South Australia

Ctenophorus fordi (Storr, 1965) Amphibolurus fordi Storr, 1965: 48

S Rohrlach Type locality: 12 mi NW of Coolgardie (30°49'S 121°02'E), Western Australia Holotype: WAM R19151 Distribution: Australia: New South Wales, Queensland, South Australia, Victoria, West Australia: from Comet Vale and Coolgardie eastwards to the southern rim of the Victoria desert

Ctenophorus gibba (Houston, 1974) Amphibolurus gibba Houston, 1974: 209 Type locality: 5.5 km NNW of Alberrie Creek Railway Siding (29°35'S 137°31'E), South Australia Holotype: SAMA R13954 Distribution: Australia: South Australia

Ctenophorus isolepis (Fischer, 1881) Subspecies: isolepis (Fischer, 1881) Grammatophora isolepis Fischer, 1881: 232 S Zozaya Type locality: Nickol Bay, Western Australia Syntypes: SMNS 2051, ZMB 10072 Distribution: Australia: from North West Cape, southern Kimberley, northern interior Northern Territory, western Queensland

Ctenophorus isolepis gularis (Sternfeld, 1925) Amphibolurus maculatus gularis Sternfeld, 1925: 231 Lectotype: SMF 10294, designated by Mertens 1967 Type locality: Hermannsburg Mission, upper Finke River, Northern Territory, Australia Distribution: Australia: interior of Western Australia to western Queensland

Ctenophorus isolepis citrinus (Storr, 1965) Amphibolurus isolepis citrinus Storr, 1965: 51 Holotype: WAM R21706 Type locality: 2 mi W of Boorabbin (31°12’S, 120°17’E), Western Australia Distribution: Australia: southern interior of Western Australia

Ctenophorus maculatus (Gray, 1831) Subspecies: maculatus (Gray, 1831) Uromastyx maculatus Gray, 1831: 62 Synonym Grammatophora gaimardii Duméril & Bibron, 1837: 470 Type locality:. vicinity of Shark Bay, Western Australia. Holotype: MNHP 0.1259 Type locality: Shark Bay, Western Australia Holotype: MHNP 0. 1259 Distribution: Australia: coast of Western Australia, north to Shark Bay

Ctenophorus maculatus badius (Storr, 1965) Amphibolurus maculatus badius Storr, 1965: 46 Holotype: WAM R18624 Type locality: 15 mi SE of Cardabia ( 23°15’S, 113°55’E), Western Australia Distribution: Australia: northwest coast of Western Australia

Ctenophorus maculatus dualis (Storr, 1965) Amphibolurus maculatus dualis Storr, 1965: 47 Holotype: WAM R23637 Type locality: 7 mi SSE of Cocklebiddy (32°08’S, 126°08’E), Western Australia Distribution: Australia: southern edge of Nullarbor Plain, Western Australia

Ctenophorus maculatus griseus (Storr, 1965) Amphibolurus maculatus griseus Storr, 1965: 47 Holotype: WAM R21708 Type locality: 14 mi west of Boorabbin (31°15'S 120°05'E), Western Australia Distribution: Australia: southern interior and south coast, Western Australia

Ctenophorus maculosus (Mitchell, 1948) Tympanocryptis maculosa Mitchell, 1948:78 Type locality: Lake Eyre North, South Australia Holotype: SAMA R2220a Distribution: Australia: salt lakes of interior South Australia

Ctenophorus mckenziei (Storr, 1981) Amphibolurus mckenziei Storr, 1981: 605 Type locality: 8 km SW of Ponier Rock (32°59'S 123°26'E), Western Australia Holotype: WAM R59754 Distribution: Australia: southeastern interior of Western Australia; southwestern South Australia

Ctenophorus mirrityana McLean, Moussalli, Sass, Stuart-Fox, 2013 Ctenophorus mirrityana McLean, Moussalli, Sass, Stuart-Fox, 2013: 54 Type locality: Mootwingee (31°17'S, 142°18'E), [Historic Site, Mutawintji National Park], New South Wales, Australia Holotype: AMS R47295 Distribution: Australia: arid northwest New South Wales

Ctenophorus nguyarna Doughty, Maryan, Melville & Austin, 2007 Ctenophorus nguyarna Doughty, Maryan, Melville & Austin, 2007: 75 Type locality: Savory Creek mouth (23°21'08"S 122°40'03"E), Lake Disappointment, Western Australia. Holotype: WAM R157979 Distribution: Australia: Little Sandy Desert, Lake Disappointment, Western Australia

Ctenophorus nuchalis (De Vis, 1884) Macrops nuchalis De Vis, 1884: 97

S Zozaya Synonyms Amphibolurus reticulatus major Sternfeld, 1919:78 Type locality: Hermmansburg Mission, upper Finke River, NT Lectotype: SMF 10316, designated by Mertens 1967 Grammatophora inermis De Vis, 1888: 812 Type locality: Central Queensland Type: QM, not located Type locality: Delta Station, Bogatungan, Queensland, Australia Syntypes: QM J1405–1410 Distribution: Australia: interior of New South Wales, Northern Territory, Queensland, South Australia, Western Australia

Ctenophorus ornatus (Gray, 1845) Grammatophora ornata Gray, 1845: 253 S Zozaya Synonym Amphibolurus tibialis Ahl, E. 1926:188 Type locality: Australia Holotype: ZMB 30100 Type locality: Western Australia Holotype: BMNH 1946.4.42 Distribution: Australia: arid southwestern Western Australia

Ctenophorus parviceps (Storr, 1964) Tympanocryptis parviceps Storr, 1964:46 Type locality: 11 mi SE of Ningaloo (22°48'S 113°49'E), Western Australia Holotype: WAM R16984 Distribution: Australia: Western Australia: between Exmouth Gulf and Shark Bay

Ctenophorus pictus (Peters, 1866) Amphibolurus pictus Peters, 1866: 88

S Zozaya Type locality: Adelaide, South Australia Syntypes: ZMB 4738-4739 Distribution: Australia: semiarid to arid regions of New South Wales, Northern Territory, Queensland, South Australia, Victoria, Western Australia

Ctenophorus reticulatus (Gray, 1845) Grammatophora reticulata Gray, 1845: 252 D Robertson Synonyms Grammatophora laevis Günther, 1867:52 Type locality: Champion Bay, Western Australia Syntypes: BMNH 1946.9.4.37–40 Amphibolurus darlingtoni Loveridge, 1932:33 Type locality: Mullewa, Western Australia Holotype: MCZ 32958 Type locality: Champion Bay, Western Australia Syntypes: BMNH 1946.9.4.35–36 Distribution: Australia: arid to semiarid regions of South Australia and Western Australia

Ctenophorus rubens (Storr, 1965) Amphibolurus isolepis rubens Storr, 1965:50

S Zozaya Type locality: Yanrey (22°30'S 114°48'E), Western Australia Holotype: WAM R18740 Distribution: Australia: Exmouth Gulf and arid interior, Western Australia

Ctenophorus rufescens (Stirling & Zietz, 1893) Amphibolurus rufescens Stirling & Zietz,1893:164 Type locality: Mt. Sir Thomas, Birksgate Range, South Australia Lectotype: SAMA R1423, designated by Houston 1976 Distribution: Australia: nortwestern South Australia, southwestern Northern Territory, eastern Western Australia

Ctenophorus salinarum (Storr, 1966) Amphibolurus pictus salinarum Storr, 1966: 18 Type locality: Norseman (32°10'S, 121°46'E), Western Australia Holotype: WAM R17649 Distribution: Australia: arid Western Australia; South Australia: western plateau

Ctenophorus scutulatus (Stirling & Zietz, 1893) Amphibolurus scutulatus Stirling & Zietz, 1893: 165 Synonyms Amphibolurus websteri Boulenger, 1904: 414 Type locality: Coolgardie District, 400 mi inland of Perth, Western Australia Syntypes: BMNH 1946.9.4.31–32 Amphibolurus holsti Rosén, 1905:141 Type locality: Western Australia Holotype: ZMLU 284 Type locality: between Fraser Range and Queen Victoria Springs, Western Australia Syntypes: SAMA R1459, R3024, R4814 Distribution: Australia: arid and semiarid South Australia, Western Australia

Ctenophorus spinodomus Sadlier, Colgan, Beatson, Cogger, 2019 Ctenophorus spinodomus Sadlier, Colgan, Beatson, Cogger, 2019: 207 Type locality : Yathong Nature Reserve (32.5742°S, 145.3828°E) New South Wales, Australia Holotype : AMS R.156615 Distribution : Australia: Mallee dunefields of central-west and southwest NSW and adjacent areas of inland southeast SA that lie to the north and west of the main Murray River channel, within the IBRA bioregions of the Murray-Darling Depression (with the exception of the Darling Depression subregion) and Cobar Peneplain bioregions.

Ctenophorus tjantjalka Johnston, 1992 Ctenophorus tjantjalka Johnston, 1992: 51 Type locality: just north of Mabel Range (26°45'S,113°48'E), South Australia Holotype: SAMA R17934a Distribution: Australia: from Painted Hill to Everard Range, South Australia

Ctenophorus vadnappa (Houston, 1974) Amphibolurus vadnappa Houston, 1974:55 Type locality: Aroona Waters (138°21'E 30°35'S), Flinders Ranges, South Australia Holotype: SAMA R3416B Distribution: Australia: northern Flinders Range to Lake Torrens, South Australia

Ctenophorus yinnietharra (Storr, 1981) Amphibolurus yinnietharra Storr, 1981: 602 Type locality: 5 km E of Yinnietharra (24°41'S,116°13'E), Western Australia Holotype: WAM R51675 Distribution: Australia: arid central western Western Australia

Dendragama Doria, 1888 Doria, G. 1888. Note Erpetologiche - Alcuni nuovi Sauri raccolti in Sumatra dal Dr. O. Beccari. Annali del Museo civico di storia naturale Giacomo Doria 2 (ser. 6): 646-652 Type species: Calotes boulengeri Doria, 1888, by original designation

Dendragama australis Harvey, Shavey, Sidik, Kurniawan & Smith, 2017

Dendragama australis Harvey, Shavey, Sidik, Kurniawan & Smith, 2017: 77 Type locality: trail up Gunung Dempo above Kampung Empat (4.037448°S, 103.145268°E) , Kabupaten Pagaralam, Provinsi Sumatera Selatan, Indonesia Holotype: MZB 13786 Distribution: Indonesia: Sumatra: Barisan Range

EN Smith

Dendragama boulengeri Doria, 1888 Dendragama boulengeri Doria, 1888: 649

J Holden Synonyms Acanthosaura schneideri Ahl, 1926: 186 Type locality: Simbolon, Battaker Plateau, Sumatra. Holotype: ZMB 15664 Salea rosaceum Thominot, 1889: 24 Type locality: Singapore Holotype: MNHP 0.6806 Type locality: Mount Singalang, West Sumatra, Indonesia Lectotype: MSNG 29936, designated by Capocaccia 1961 Distribution: Indonesia: Sumatra

Dendragama dioidema Harvey, Shavey, Sidik, Kurniawan & Smith, 2017 EN Smith Dendragama dioidema Harvey, Shavey, Sidik, Kurniawan & Smith, 2017: 87 Type locality: Bukit Sama (4.665128°N, 96.809378°E), Kampung Telege Atu, Kebayakan, Kabupaten Aceh Tengah, Provinsi Aceh, Indonesia Holotype: MZB 13814 Distribution: Indonesia: Sumatra: Pusat Gaya Range

Diploderma Hallowell, 1861 Hallowell, E. 1861. Report upon the Reptilia of the North Pacific Exploring Expedition, under command of Capt. John Rogers, U. S. N. Proceedings of the Academy of Natural Sciences of Philadelphia 12 [1860]: 490 Type species: Diploderma polygonatum Hallowell, 1861.

Diploderma polygonatum I Das

Diploderma batangensense (Li, Deng, Wu & Wand, 2001) Japalura batangensis Li, Deng, Wu & Wand, 2001: 330 Type locality : Wangjiao (29°17.6 ′N, 99°5.2 ′E), Batang Country, Sichuan, China Holotype : STC Ba-98012 Distribution: China: western Sichuan

Diploderma brevicauda (Manthey, Denzer, Hou & Wang, 2012) Japalura brevicauda Manthey, Denzer, Hou & Wang, 2012: 29 Type locality : Snow Mt. Village (ca. 27°05’N, 100°11’E), 19 km north of Lijiang, northwestern Yunnan, China Holotype: AMNH 19879 Distribution: China: northwestern Yunnan

Diploderma brevipes (Gressitt, 1936) Japalura brevipes Gressitt, 1936: 117 Type locality : Bukai, near Horisha, central Taiwan Holotype: CAS 71998 Distribution: central Taiwan

Diploderma chapaensemse (Bourret, 1937) Japalura chapaensis Bourret, 1937: 62 Type locality : Chapa (22°20'N, 103°55' E), Vietnam Holotype: MHNP 1948.45 Distribution: northern Vietnam; China: Yunnan Province

A Nguyen

Diploderma drukdaypo Wang, Ren, Jiang, Zou, Wu, Che, Siler, 2019 Diploderma drukdaypo Wang, Ren, Jiang, Zou, Wu, Che, Siler, 2019: 508 Type locality : Chaya County (30.7294°N, 97.3808°E), Chamdo, Tibet, China Holotype: KIZ 027616 Distribution: China: valleys near the headwaters of the Mekong River in Karuo and Chaya Counties of Chamdo, Tibet

Diploderma dymondi (Boulenger, 1906) Acanthosaura dy mondi Boulenger, 1906:567 Type locality : Tongchuan-fu, Yunnan, China Holotype: BMNH 1905.5.30.1 Distribution: China: western Yunnan, southwestern Sichuan

Diploderma fasciatum (Mertens, 1926) Japalura fasciata Mertens, 1926: 146 Synonym Japalura szechwanensis Hu & Zhao, 1966:158 Type locality: Peng Hsien (= Peng Xien) and Tianquan Xien (= Tien Tsuan Hsien), Sichuan (= Szechwan), China. Holotype: CIB 613047 Type locality : Tonkin, north Vietnam Holotype: SMF 9871 Distribution : northern Vietnam; China: Sichuan, Guizhou, Guangxi, Hunan

Diploderma flaviceps (Barbour & Dunn, 1919) Japalura flaviceps Barbour & Dunn, 1919: 16 Type locality : Tung River (ca. 30.6006° 101.4621°), western Szechuan, China Holotype: MCZ R-12469 Distribution : China: western Sichuan, eastern Xizang, Qinghai, Gansu, Shenxi, Shanxi, northern Yunnan

Diploderma grahami (Stejneger, 1924) Phoxophrys grahami Stejneger, 1924: 120 Type locality : Sui-fu, Szechuan, China Holotype : USNM 65500 Distribution: China: Sichuan

Diploderma hamptoni (Smith, 1935) Japalura hamptoni Smith, 1935: 175 Type locality : Mogok, northern Burma Holotype: BMNH 1946.8.14.1 Distribution : northern Burma

Diploderma iadinum (Wang, Jiang, Siler, and Che, 2016) Japalura iadina Wang, Jiang, Siler, and Che, 2016: 51 Type locality : Lancang Valley at Ninong, Deqin, northwest Yunnan, China Holotype : KIZ 019321 Distribution: Lancang Valley, Deqin County, Yunnan Province, China

K Wang

Diploderma laeviventris (Wang, Jiang, Siler, and Che, 2016) Japalura laeviventris Wang, Jiang, Siler, and Che, 2016: 42 Type locality: near the Nujiang Bridge in the upper Nujiang Valley at Baxoi (=Basu), Qamdo (=Changdu), eastern Tibet (=Xizang) Holotype: KIZ 014038 Distribution: Nujiang Valley, Changdu, Sichuan Province, China

Diploderma luei (Ota, Chen & Shang, 1998) Japalura luei Ota, Chen & Shang, 1998: 649 Type locality : Mt. Taipingshan (24º31’N, 121º30’E), Nanao Hsiang, Ilan Pref., Taiwan Holotype : NTNUB 2235 Distribution: northern Taiwan

Diploderma makii (Ota, 1989) Japalura makii Ota, 1989: 570 Type locality : Chitou (23º42’N, 120º47’E), Nantou Pref, Taiwan Holotype : OMNH R2565 Distribution: central Taiwan

Diploderma micangshanense (Song, 1987) Japalura micangshanensis Song, 1987:59 Type locality : Qingmuchuan, Ningqiang Co., Shaanxi Prov., China Holotype: SIZ 82158 Distribution: China: Shaanxi, Gansu, Sichuan, Henan

Diploderma ngoclinense (Ananjeva, Orlov & Nguyen, 2017) Japalura ngoclinensis Ananjeva, Orlov & Nguyen, 2017: 15 Type locality : above Mang Xang Village, Ngoc Linh Mountain (15°05'N 107°57'E), Dac Glei District, Kon Tum Province, Vietnam Holotype: ZISP 26230 Distribution: Vietnam: Kon Tum Province: Ngoc Linh Mountain

Diploderma polygonatum Hallowell, 1861 Subspecies: Diploderma polygonatum polygonatum (Hallowell, 1861) Diploderma polygonatum Hallowell, 1861: 490 Type locality: Kerama shima, Ryukyu Islands, Japan Holotype : USNM 12203 Distribution: Kerama shima, Ryukyu Islands, Japan

Diploderma polygonatum ishigakiense (Van Denburgh, 1912) Japalura polygonata ishigakiensis Van Denburgh, 1912: 211 Holotype: CAS 21354 Type locality : Ishigaki, Loo Choo Islands, Japan Distribution: Japan: Yaeyama Islands

Diploderma polygonatum miyakense (Van Denburgh, 1912) Japalura polygonata miyakensis Van Denburgh, 1912: 211 Holotype: CAS 21353 Type locality : Miyako, Loo Choo Islands, Japan Distribution: Japan: Miyako Islands, Ryukyu Islands

Diploderma polygonatum xanthostoma (Ota, 1991) Japalura polygonata xanthostoma Ota, 1991:288 Holotype: KUZ R9855 Type locality : Waishuangchi ,Taipei Prefecture, Taiwan Distribution : northernTaiwan

Diploderma polygonatum donan (Ota, 2003) Japalura polygonata donan Ota, 2003: 63 Holotype: KUZ R234 Type locality: Mt. Urabudake (24°27'N, 123°00'E), Yonagunijima Island of the Yaeyama Group, Ryukyu Archipelago, Japan Distribution : Japan: Yonagunijima Island, Ryukyu Archipelago

Diploderma slowinskii (Rao, Vindum, Ma, Fu and Wilkinson, 2017) Japalura slowinskii Rao, Vindum, Ma, Fu and Wilkinson, 2017: 88 Type locality : small village S of Fugong (County), along Nu River (26º48'44.0"N, 98º53'10.7"E), Nujiang Prefecture, Yunnan Province, China Holotype: KIZ 2000R0608 Distribution: Nu River Valley, Liushu and Gongshan Counties, Hengduan Mountains, Yunnan Province, China

Diploderma splendidum (Barbour & Dunn, 1919) Japalura splendida Barbour & Dunn, 1919: 18 Type locality : from the gorge of the Yangtze River near Tchang, Hupeh, central China Holotype : USNM 35522 Distribution: southeastern Tibet; southwestern China: Szechuan, Guizhou, Yunnan A Liman

Diploderma swild Wang, Wu, Jiang, Chen, Miao,Siler,Che, 2019 Diploderma swild Wang, Wu, Jiang, Chen, Miao,Siler,Che, 2019: 459 Type locality: Hongbao Village, Yanbian County, Panzhihua District, Sichuan Province, China Holotype: KIZ 034912 Distribution: China: Sichuan Province, Yanbian, Xichang, Huili Countys

Diploderma swinhonis (Günther, 1864) Japalura swinhonis Günther, 1864: 133 Synonyms Japalura mitsukurii Stejneger, 1898:218 Type locality: Botel Tobago (=Lanyu Island), Taiwan Holotype: USNM 36505 Japalura yunnanensis popei Wettstein, 1938: 176 Type locality: Cambodja, in error for Taiwan Holotype: NMHW 23358:1 Japalura swinhonis formosensis Liang & Wang, 1976: 162 Type locality: Puli, Nan-tou Hsien (=Nantou Co.) , central Taiwan Holotype: NTUM R0045 Type locality : Tamsui, Taiwan JP Fischer Holotype : BMNH 1946.8.14 Distribution: Taiwan, incl. Lanyu

Diploderma varcoae (Boulenger, 1918) Acanthosaura varcoae Boulenger, 1918: 162 Type locality : Jizi Mountain, Dali City, Yunnan, China Syntypes: BMNH 1946.8.11.15, 1946.8.14.6 Distribution: southwestern China: western Yunnan, Guizhou

Diploderma vela (Wang, Jiang, Pan & Hou, 2015) Japalura vela Wang, Jiang, Pan & Hou, 2015: 160 Type locality: Quzika of Markam (29°5' N, 98°36' E), eastern Tibet, China Holotype: KIZ 013801 Distribution: Upper Mekong (Lancang) River valley, eastern Tibet, China

Diploderma yulongense (Manthey, Denzer, Hou & Wang, 2012) Japalura yulongensis Manthey, Denzer, Hou & Wang, 2012: 34 Type locality : Ulukay Village, eastern slopes of the Yulong Snow Mountains, near Lijiang (ca. 26°53’N, 100°11’E), northwestern Yunnan, China Holotype: ZMB 76395 Distribution: China: northwestern Yunnan Last seen 1914

Diploderma yunnanense (Anderson, 1878) Japalura yunnanensis Anderson, 1878: 803 Type locality : Longling, western Yunnan, China Lectotype : BMNH 1932.12.1.1, designated by Mahony Distribution : China: Yunnan; Burma: Kachin

Diploderma vela (Wang, Jiang, Pan & Hou, 2015) Japalura vela Wang, Jiang, Pan & Hou, 2015: 160 Type locality: Quzika of Markam (29°5' N, 98°36' E), eastern Tibet, China Holotype: KIZ 013801 Distribution: Upper Mekong (Lancang) River valley , eastern Tibet, China

Diploderma zhaoermii (Goa & Hou, 2002) Japalura zhaoermii Goa & Hou, 2002: 3 Type locality : Mianchi Town (31.35°N, 103.57°E), Wenchuan County, Sichuan Province, China Holotype : CIB2001-GZF0001 Distribution: China: western Sichuan

Diporiphora Gray, 1842 Gray, J.E. 1842. Description of some hitherto unrecorded species of Australian reptiles and batrachians. pp. 51-57 in Gray, J.E. (ed.). The Zoological Miscellany . London : Treuttel, Würz & Co Type species: Gray, 1842 by monotypy

Diporiphora adductus Doughty, Kealley & Melville, 2012 Diporiphora adductus Doughty, Kealley & Melville, 2012: 17

S Zozaya Type locality: 2 km west of Bullara Homestead (22°40 ′23 ″S, 114°0 ′58 ″E), Western Australia Holotype: WAM R141589 Distribution: Australia: Western Australia: northern Carnarvon Basin

Diporiphora albilabris Storr, 1974 albilabris Storr, 1974: 133 Type locality: Mitchell Plateau (14°48'S 125°50'E), Western Australia Holotype: WAM R43517 Distribution: Australia: central and northern Kimberley, Western Australia

Diporiphora ameliae Emmott, Couper, Melville & Chapple, 2012 Diporiphora ameliae Emmott, Couper, Melville & Chapple, 2012: 44 S Zozaya Type locality: SW corner (24°14'06''S, 143°05'49.9''E) of Valetta Stn, Queensland, Australia Holotype: QM J91277 Distribution: Australia: central Queensland

Diporiphora amphiboluroides Lucas & Frost, 1902 Diporiphora amphiboluroides Lucas & Frost, 1902: 76 Type locality: Western Australia Holotype: NMV D8813 Distribution: Australia: Western Australia: northwest coast, western plateau

Diporiphora arnhemica Storr, 1974 arnhemica Storr, 1974:137 Type locality: near upper Katherine River (14°13'S, 132°36'E), Northern Territory, Australia Holotype: ANWC R740 Distribution: Australia: Northern Territory: southern Kimberleys to Arnhem Land

Diporiphora australis (Steindachner, 1867) Calotella australis Steindachner, 1867: 29 S Zozaya Synonyms Diporiphora nuchalis De Vis, 1884:98 Type locality: central and south coast district, Queensland. Syntypes: not located Diporiphora ornata De Vis, 1884:99 Type locality: unknown Holotype: not located Physignathus nigricollis Lönnberg & Andersson,1915:4 Type locality: Cooktown, Queensland Holotype: NRM 3209 Type locality: Cape York, Queensland, Australia Syntypes: NHMW 19821:1-2 Distribution: Australia: northeastern New South Wales to southern Cape York, Queensland

Diporiphora bennettii (Gray, 1845) Gindalia bennettii Gray, 1845:247 Type locality: northwest coast of Australia Holotype: BMNH 1946.8.12.77 Distribution: Australia: Western Australia: northern Kimberleys to adjacent Northern Territory

Diporiphora bilineata Gray, 1842 Diporiphora bilineata Gray, 1842: 54

S Zozaya Synonyms Grammatophora macrolepis Günther, 1867:51 Type localiy: Australia Holotype: BMNH 1946.8.12.74 Diporiphora brevicauda De Vis, 1884:99 Type locality: Cape York, Queensland Holotype, not located Diporiphora pentalineata De Vis, 1884:99 Type locality:. Cape York, Queensland Holotype, not located Type locality: Port Essington, Northern Territory, Australia Syntypes: BMNH 1946.8.12.75–76 Distribution: Australia: coastal Northern Territory

Diporiphora carpentariensis Melville, Date, Horner, Doughty, 2019 Diporiphora carpentariensis Melville, Date, Horner, Doughty, 2019: 50

Type locality : Littleton National Park (18.2243°S, 142.7489 E), northern Queensland

E. Vanderduys Holotype: QM J88197 Distribution : Australia: Queensland: north-east Gulf of Carpentaria region of the western extent of Cape York Peninsula .

Diporiphora convergens Storr, 1974 Diporiphora convergens Storr, 1974:133 Type locality: Crystal Creek (14°30'S, 125°47'E), Western Australia Holotype: WAM R42931 Distribution: Known only from Crystal Creek, on Admiralty Gulf in northwestern Kimberely, Western Australia Last seen 1972

Diporiphora gracilis Melville, Date, Horner, Doughty, 2019 Diporiphora gracilis Melville, Date, Horner, Doughty, 2019: 46 Type locality : Fairfield-Leopold Downs Road(17° 29' 37.0" S, 125° 2' 17.7" E), south of Gibb River Road, Western Australia Holotype: WAM R177291 Distribution: Australia: Western Australia: south-western Kimberley region

Diporiphora granulifera Melville, Date, Horner, Doughty, 2019 Diporiphora granulifera Melville, Date, Horner, Doughty, 2019: 48 Type locality : Downs Road (20.3714°S, 139.1529° E), 2 km from Barkly Highway, Queensland Holotype : QM J96362 Distribution: Australia: Queensland, north-western Gulf of Carpentaria region

Diporiphora jugularis (Macleay, 1877)

S. Wilson

Grammatophora jugularis Macleay, 1877:104 Type locality : Cape Grenville, Cape York Peninsula, Queensland. Syntypes: AMS R40672–74 Distribution: Australia: Queensland: eastern Cape York Peninsula

Diporiphora lalliae Storr, 1974 Storr, 1974: 138 Type locality: Langey Crossing (17°39'S, 123°34'E), Western Australia Holotype: WAM R23030 Distribution: Australia: western deserts: from the southern Kimberley in Western Australia, central Northern Territory to the far west of Queensland

Diporiphora linga Houston, 1977 Houston, 1977: 203 Type locality: 23 km N of Koonibba Mission (31°42'S, 133°26'E), South Australia Holotype: SAMA R15020F Distribution: Australia: southwestern South Australia

Diporiphora magna Storr, 1974 Storr, 1974:137 Type locality: Old Lissadell (now submerged by Lake Argyle) (16°30'S, 128°41'E), Western Australia Holotype: WAM R42786 Distribution: Australia: tropical savannah region: northern Western Australia, central Northern Territory, western Queensland

Diporiphora margaretae Storr, 1974 Diporiphora bilineata margaretae Storr,1974: 143 Type locality: Kalumburu (14°18'S, 126°30'E), Western Australia Holotype: WAM R27648 Distribution: Australia: Western Australia: north Kimberley region, Anjo Peninsula and Sir Graham Moore Island in the extreme north, Kalumburu, Mitchell Plateau, Prince Regent River and Drysdale River National Parks, and on Mary Island.

Diporiphora nobbi (Witten, 1972) Amphibolurus nobbi nobbi Witten, 1972: 191

S Zozaya Synonyms Amphibolurus nobbi coggeri Witten, 1972:192 Type locality: Warrumbungle Mts, New South Wales Holotype: AMS R17660 Wittenagama parnabyi Wells & Wellington,1985:20 Type locality: 88 km W Alpha, Queensland Holotype: AMS R65949 Type locality: 24 mi ENE of Guyra, New South Wales, Australia Holotype: AMS R33440 Distribution: Australia: New South Wales, eastern Queensland, far eastern South Australia, northwestern Victoria Diporiphora pallida Melville, Date, Horner, Doughty, 2019 Diporiphora pallida Melville, Date, Horner, Doughty, 2019: 52 Type locality : Mitchell Plateau (14° 49.45' S, 125° 42.12' E), Western Australia Holotype : WAM R177292 Distribution : Australia: Western Australia, Mitchell Plateau

Diporiphora paraconvergens Doughty, Kealley & Melville, 2012 Diporiphora paraconvergens Doughty, Kealley & Melville, 2012: 20 Type locality: 23 km north-east of Warrawagine Homestead (20º41 ′54 ″S, 120º51 ′23 ″E), Pilbara Biodiversity Survey quadrat PHYE01, Western Australia Holotype: WAM R163948 Distribution: Australia: Western Australia: Great Sandy Desert; northwestern South Australia, southwestern Northern Territory

Diporiphora perplexa Melville, Date, Horner, Doughty, 2019 Diporiphora perplexa Melville, Date, Horner, Doughty, 2019: 37 Type locality : Gibb River Road (15° 57.31' S,126° 52.9' E), 20 km west of Ellenbrae Station, WA Holotype : WAM R177290 Distribution: Australia: Western Australia: Yampi Peninsula in the south-western Kimberley, extending eastwards to the Kununurra area; western Northern Territory, Jasper Gorge area.

Diporiphora phaeospinosa Edwards & Melville, 2011 Diporiphora phaeospinosa Edwards & Melville, 2011: 536

S Zozaya Type locality: Bauhinia Station (25.17°S 149.20°E), Queensland, Australia Holotype: NMV D 74128 Distribution: Australia: central western Queensland

Diporiphora pindan Storr, 1980 Storr, 1980: 256 Type locality: 5 km N of Coulomb Point (17°18'S, 122°10'E), Western Australia Holotype: WAM R58402 Distribution: Australia: Western Australia: southwestern Kimberely, Great Sandy Desert

Diporiphora reginae Glauert, 1959 Glauert, 1959: 10 Type locality: Karin Rock, 14 mi from Cunderlee Mission in the Queen Victoria Springs area, Western Australia. Holotype: WAM R12961 Distribution: Australia: southern interior of Western Australia

Diporiphora sobria Storr, 1974 Diporiphora albilabris sobria Storr, 1974: 135 Type locality : Pine Creek (14°04'S, 131°58'E), Northern Territory, Australia Holotype : WAM R23180 Distribution: Australia: Western Australia, southern Kimberley region; northern Northern Territory; western Queensland

Diporiphora superba Storr, 1974 Diporiphora superba Storr, 1974: 144

M O’Shea Type locality: Mitchell River (14°25'S, 125°50'E), Western Australia Holotype: WAM R43178 Distribution: Australia: Kimberley, Western Australia

Diporiphora valens Storr, 1980 Diporiphora valens Storr, 1980:. 260

S Zozaya Type locality: near Tom Price ( c. 22°45'S, 117°45'E), Western Australia Holotype: WAM R31009 Distribution: Australia: West Australia: interior Pilbara region

Diporiphora vescus Doughty, Kealley & Melville, 2012 Diporiphora vescus Doughty, Kealley & Melville, 2012: 12

S Zozaya Type locality: 86 km south of Port Hedland (21°03 ′36 ″S,118°45 ′00 ″E), Western Australia Holotype: WAM R145536 Distibution: Australia: Western Australia: northern Pilbara region

Diporiphora winneckei Lucas & Frost, 1896 Lucas & Frost, 1896: 132

L Young Type locality: Charlotte Waters, Northern Territory, Australia Lectotype : NMV D10156, designated by Coventry 1970 Distribution: Australia: from the Simpson Desert to northwest coast: southeastern Northern Territory, southeastern Queensland, northern South Australia, all central Western Australia

Draco Linnaeus, 1758 Linnaeus, C. 1758. Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis . Tomus I. Editio decima, reformata. Laurentii Salvii, Holmiæ. 10th Edition: 824 pp. Type species: Draco volans Linnaeus, 1758, by original designation

Draco abbreviatus Hardwicke & Gray, 1827 Draco abbreviatus Hardwicke & Gray, 1827: 219 Type locality: Singapore Holotype: BMNH, lost Distribution: Singapore; West Malaysia

Draco beccarii Peters & Doria, 1878 Draco beccarii Peters & Doria, 1878: 373 Type locality: Kandari, Sulawesi, Indonesia Lectotype: MSNG 9523, designated by Capocaccia 1961 Distribution: Indonesia: eastern Sulawesi, Buton Is., Kabaena Is., Muna Is

Draco biaro Lazell, 1987 Draco biaro Lazell, 1987: 3 Type locality: Pulau Biaro, Kepulauan Sangihe, Indonesia Holotype: MCZ 170898 Distribution Indonesia: Sangihe Archipelago Last seen 1998

Draco bimaculatus Günther, 1864 Draco bimaculatus Günther, 1864: 127 Type locality: Philippine Islands Holotype: BMNH XXIV.1.g Distribution: southern Philippines: Basilan, Bohol, Dinagat, Jolo, Lapac, Leyte, Lugus, Mindanao, Ponson, Samar, Sanga Sanga, Siargao, Tawitawi, Zamboanga, Siasi, Agusan

Draco blanfordii Blanford, 1878 Draco blanfordii Blanford, 1878: 267

M Muin Synonyms Draco cyanolaemus Boulenger, 1908: 64 Type locality: Telcom, Perak/Pahang boundary, West Malaysia Holotype: BMHN 1946.8.26.35 Type locality: forest of Tavoy , Tenasserim province, Burma Holotype: BMNH 1946.8.26.76 Distribution: China: southwestern Yunnan; eastern Thailand; West Malaysia; Burma: Vietnam; India; Bangladesh

Draco boschmai Hennig, 1936 Draco volans boschmai Hennig, 1936: 180 Type locality: Maumeri, Flores, Indonesia Holotype : ZMA 11025 Distribution: Indonesia: Lombok, Sumbawa, Komodo, Rintja, Sumba, Adunara, Flores

Draco bourouniensis Lesson, 1834 Draco bourouniensis Lesson, 1834: pl. 37 Synonym Draco toxopei De Jong, 1926: 88 Type locality: Buru Island, Maluku, Indonesia Lectotype: ZMA 10934, designated by Musters 1983 Type locality: Buru Island, Maluku, Indonesia Syntypes: ZMA 10931-33 Distribution: Indonesia: Buru Island

Draco caerulhians Lazell, 1992 Draco caerulhians Lazell, 1992: 482 Type locality: Manganitu, Sangihe Besar Island, Indonesia Holotype: MCZ 173321 Distribution: Indonesia: Sangir Island, Sangihe Archipelago, North Sulawesi

Draco cornutus Günther, 1864 Draco cornutus Günther, 1864:125

S Harikishnan Synonym Draco gracilis Barbour. 1903 : 59 Type locality : Sarawak, Borneo, East Malaysia Holotype : MCZ 6713 Type locality: Borneo Syntypes: BMNH 1946.8.27.23-24 Distribution : Indonesia: Kalimantan (Borneo); East Malaysia: Sabah, Sarawak

Draco cristatellus Günther, 1872 Draco cristatellus Günther, 1872: 592 Type locality: Sarawak, Borneo, East Malaysia Holotype: BMNH 1872.2.19.4 Distribution: Indonesia: Kalimantan (Borneo); East Malaysia :Sarawak; Brunei

Draco cyanopterus Peters, 1867 Draco reticulatus cyanopterus Peters, 1867: 15 Synonym Draco everetti Boulenger, 1885: 258 Type locality: Placer, northeastern Mindanao, Philippines Lectotype: BMNH 1877.12.13.16, designated by Musters 1983 Type locality: Mindanao, Philippines Lectotype: ZMB 725, designated by Denzer et al. 1997 Distribution: Philippines: Camiguin Sur, Dinagat, Mindanao

Draco dussumieri Duméril & Bibron, 1837 Draco dussumieri Duméril & Bibron, 1837: 456 S Harikishnan Type locality: Malabar, southern India Holotype: MNHP 0.2590 Distribution: southern India: Goa, Madras, Malabar coast, Nilghiri Hills, Travancore, Western Ghats & Eastern Ghats, Gujarat

Draco fimbriatus Kuhl, 1820 Draco fimbriatus Kuhl, 1820: 101

M Muin Synonyms Draco grandis, Bartlett, 1895: 83. Type locality : Sarawak, Matang, East Malaysia Holotype : SMK, cannot be found Draco fimbriatus hennigi Musters, 1983:27 Type locality: Soekamandi, Jawa Barat, Indonesia Holotype: RMNH 19973 Type locality: Java, Indonesia Holotype: ZMB 713 Distribution: Indonesia: Java, Bali, Kalimantan (Borneo), Sumatra, Natuna; East Malaysia: Sarawak, Sabah

Draco formosus Boulenger, 1900 Draco formosus Boulenger, 1900: 190 Type locality: Larut Hills, Perak, West Malaysia Holotype: BMNH 1946.8.13.43 Distribution: southern Thailand; West Malaysia; Indonesia: Natuna Island

Draco guentheri Boulenger, 1885 Draco guentheri Boulenger, 1885: 257 Synonym Draco rizali Wandolleck, 1900: 15 Type locality: Dapitan, Mindanao,Philippines Holotype: MTKD D 1168; destroyed in the bombing of Dresden in WWII Type locality: Philippine Islands Holotype: BMNH 1879.4.16.4 Distribution: Philippines: Basilan, Bongao, Jolo, Mindanao, Sanga Sanga, Siasi, Siminul

Draco haematopogon Gray, 1831 Draco haematopogon Gray, 1831: 59

I Das Synonym Draco microlepis Boulenger, 1893: 532 Type locality: Merabah, Borneo, Sabah, East Malaysia Syntypes : BMNH 1946.8.26.81-82 Type locality: Parang Island off Java (05° 45’S; 110° 15’E), Greater Sundas, Indonesia Syntypes: MNHP 0.2577, 0.2577A, 0.230 Distribution: Indonesia: Sumatra, Java, Kalimantan (Borneo) ; West Malaysia, East Malaysia : Sarawak, Sabah ; southern Thailand

Draco indochinensis Smith, 1928 Draco indochinensis Smith, 1928: 248 Type locality: Kamchay Mountains, Cambodia Holotype: BMNH 1946.8.26.77 Distribution: southern Vietnam, Cambodia

Draco iskandari McGuire & Brown, Mumpuni, Riyanto & Andayani, 2007 Draco iskandari McGuire & Brown, Mumpuni, Riyanto & Andayani, 2007L 183 Type locality: Desa Haasi (2.32012° N, 125.41974° E), Tahulandang Island, Sangihe Islands, Provine Sulawesi Utara, Indonesia. Holotype: MZB.Lace.6336 Distribution: Indonesia: Tahulandang Island, off North Sulawesi Last seen 1998

Draco jareckii Lazell, 1992 Draco jareckii Lazell, 1992: 488

M O’Shea Type locality: Basco, Batan Island, Batanes Province, Philippines Holotype: PNM 1797 Distribution: Philippines: Batan Island

Draco lineatus Daudin, 1802 Subspecies: Draco lineatus lineatus Daudin, 1802 Draco lineatus Daudin, 1802: 298 Synonyms Draco amboinensis Lesson, 1834: pl 38 Type locality: Ambon Island, Indonesia Type: MNHP, not located Draco reinwardtii Boie 1827: ms Type locality: Java, Indonesia Syntypes: RMNH 2965 Type locality: Java (in error, probably Maluku), Indonesia Holotype: MHNP, lost Distribution: Indonesia: Ambon, Ceram, Misol

Draco lineatus ochropterus Werner, 1910 Draco ochropterus Werner, 1910: 11 Syntypes: ZMH, lost Type locality: Kei Islands, Indonesia Distribution: Indonesia: Kei Islands

Draco maculatus (Gray, 1845) Subspecies: Draco maculatus maculatus (Gray, 1845) Dracunculus maculatus Gray, 1845: 236 Type locality: Penang, West Malaysia Syntypes: BMNH 1946.8.27.5-6, 1946.8.27.22 Distribution: southern China: Guangxi, Yunnan, southeastern Xizang (= Tibet); India: eastern Himalayas to Assam - Naga hills and Abor country, Mizoram; Burma; Laos; southern Thailand; West Malaysia

Draco maculatus divergens Taylor, 1934 Draco divergens Taylor, 1934: 291 Holotype: ANSP 19938 Type locality: Doi Suthep Mountain, northern Thailand Distribution: northwestern Thailand

Jonathon Hakim

Draco maculatus haasei Boettger, 1893 Draco haasei Boettger, 1893: 429 Holotype: SMF 9700 Type locality: Chantabuni, Thailand Distribution: easternThailand, Cambodia, southern Vietnam

Draco maculatus whiteheadi Boulenger, 1900 Draco whiteheadi Boulenger, 1900: 956 Holotype: BMNH 1946.8.26.79 Type locality: Five-finger Mountains, interior of Hainan, China Distribution: northern Vietnam; China: Hainan

Draco maximus Boulenger, 1893 Draco maximus Boulenger, 1893: 522 Synonyms Draco cryptotis Despax, 1912 : 202 Type locality : Java, Greater Sundas, Indonesia Holotype : MNHP 1912.38 Draco intermedius Werner. 1910 :14 Type locality : Bandar Kwala, northwestern Sumatra, Indonesia Holotype : ZMH, not located Type locality: Mount Dulit, Sarawak, Borneo, East Malaysia Holotype: BMNH 1946.8.13.41 Distribution: West Malaysia; Indonesia: Sumatra, Natuna Island, Kalimantan (Borneo); East Malaysia: Sarawak

Draco melanopogon Boulenger, 1887 Subspecies: Draco melanopogon melanopogon Boulenger, 1887 Draco melanopogon Boulenger, 1887: 492 M Muin Type locality: Melaka; ca. 02° 12’N; 102° 15’E; West Malaysia Syntypes: BMNH 1946.8.26.87-90 Distribution: southern Thailand (south of Isthmus of Kra); West Malaysia; Indonesia: Natuna Island; Singapore

Draco melanopogon nigriappendiculatus Bartlett, 1894 Draco nigriappendiculatus Bartlett, 1894: 92 Syntypes: SMK, not found Type locality: Kuching (01° 28’N; 110° 19’E), Sarawak, Borneo, East Malaysia Distribution : Indonesia: Kalimantan, Sumatra; East Malaysia: Sarawak

Draco mindanensis Stejneger, 1908 Draco mindanensis Stejneger, 1908: 677 Type locality: Datu Anib’s place, near Catagan, northwest Mindanao, at base of Malindang Mountain, Philippines Holotype: USNM 37388 Distribution: Philippines: Mindanao, Dinagat, Leyte, Samar

Draco modiglianii Vinciguerra, 1892 Draco modiglianii Vinciguerra, 1892: 253 Type locality: BuaBua, Enggano Island, southwest of Sumatra Lectotype: MSNG 10047, designated by Capocaccia 1961 Distribution: Indonesia: endemic to Enggano Island

Draco norvillii Alcock, 1895 Draco norvillii Alcock, 1895: 14 Type locality: Assam, India Holotype: ZSI 13767 Distribution: northeastern India: Assam: Naga Hills; Arunachal Pradesh

Draco obscurus Boulenger, 1887 Subspecies: Draco obscurus obscurus Boulenger, 1887 Draco obscurus Boulenger, 1887: 95 Type locality: Larut Hills (04°47’N; 100°45’E), Perak State, West Malaysia Holotype: BMNH 1946.8.13.43 Distribution: Indonesia: Sumatra, Kalimantan (Borneo), Natuna Islands; East Malaysia: Sabah, Sarawak; West Malaysia; southern Thailand

Draco obscurus laetepictus Henning, 1936 Draco formosus laetepictus Henning, 1936: 217 Holotype: MTKD, destroyed in WWII Type locality: Siberut Island, Mentawei Islands, Indonesia Distribution: Indonesia: Mentawei Islands

Draco ornatus (Gray, 1845) Dracunculus ornatus Gray, 1845: 235 Type locality: Philippines Lectotype : BMNH 1946.8.27.30, designated by Musters 1983 Distribution: Philippines: Bohol, Dinagat, Leyte, Mindanao, Samar

Draco palawanensis McGuire & Alcala, 2000 Draco palawanensis McGuire & Alcala, 2000: 85 Type locality: Barangay Alfonzo 13, Quezon, Palawan Island, Palawan Province, Republic of the Philippines Holotype: PNM 5770 Distribution: Philippines: Palawan

Draco punctatus Boulenger, 1900 Draco punctatus Boulenger, 1900: 189 Holotype: BMNH 1946.8.26.93 Type locality: peninsular Malaysia: (04°47’N; 100°45’E); Perak State, West Malaysia Distribution: West Malaysia; southern Thailand

Draco quadrasi Boettger, 1893 Draco quadrasi Boettger, 1893: 41 Type locality: Sibuyan Island, Philippines Holotype: ZMB 728 Distribution: Philippines: Mindoro, Romblon, Semirara, Sibuyan

Draco quinquefasciatus Hardwicke & Gray, 1827 Draco quinquefasciatus Hardwicke & Gray, 1827: 219

I Das Synonyms Draco quinquefasciatus longibarba, Hennig. 1941 :193 Type locality : Baram, Borneo, Sarawak, East Malaysia Holotype : ZRC 2.659 Type locality: Penang, West Malaysia Holotype: BMNH 1946.8.27.4 Distribution: southern Thailand; West Malaysia; Singapore; Indonesia: Sumatra, Singkep, Belitung, Kalimantan (Borneo); Brunei; East Malaysia: Sarawak, Sabah .

Draco reticulatus Günther, 1864 Draco reticulatus Günther, 1864:125 Type locality: Philippine Islands Holotype: BMNH 1946.8.27.28 Distribution: Philippines: Bohol, Lapinin, Chico, Leyte, Samar

Draco rhytisma Musters, 1983 Draco lineatus rhytisma Musters, 1983:47 Type locality: Lipulalongo, Labobo Island, Banggai Islands, Indonesia Holotype: RMNH 20993 Distribution: Indonesia: Banggai Archipelago, off eastern Sulawesi

Draco spilonotus Günther, 1872 Draco spilonotus Günther, 1872: pl. 35 Type locality: Manado, North Sulawesi, Indonesia Lectotype : BMNH 1946.8.27.26, designated by Musters, 1983 Distribution: Indonesia: North Sulawesi

Draco spilopterus (Wiegmann, 1834) Dracunculus spilopterus Wiegmann, 1834:216

M O’Shea Synonym Draco rostratus Günther, 1864: 127 Type locality: Borneo, probably in error Holotype: BMNH 1946.8.27.25 Type locality: Manila, Luzon, Philippines Syntypes: BMNH 1946.8.27.29-31 Distribution: Philippines: Bantayan, Boracay, Carabao, Catunduanes, Cebu, Guimaras, Inampulugan, Kalotkot, Lubang, Luzon, Marinduque, Masbate, Negros, Panay, Polillo, Siquijor

Draco sumatranus Schlegel, 1844 Draco viridis sumatrana Schlegel, 1844:: 91 I Das Type locality: Sumatra, Indonesia Lectotype: RMNH 2933F, designated by Müller Distribution: southern Thailand; West Malaysia; Indonesia: Sumatra, Simeulue, Sipora, Nias, Enggano, Riau Archipelago, Bangka, Belitung, Natuna Islands, Kalimantan (Borneo)

Draco supriatnai McGuire & Brown, Mumpuni, Riyanto & Andayani, 2007 Draco supriatnai McGuire & Brown, Mumpuni, Riyanto & Andayani, 2007:186 Type locality: Desa Wakai (0.44479°S, 121.86502°E), Batudaka Island, , Kecamatan Una Una, Kabupaten , Province Sulawesi Tengah, Indonesia Holotype: MZB.Lace 6443 Distribution: Indonesia: Togian Islands

Draco taeniopterus Günther, 1861 Draco taeniopterus Günther, 1861: 187

P Museum Type locality: Chartaboum, coast of Thailand Holotype: BMNH 1946.8.26.75 Distribution: southern Burma; Thailand; Cambodia; southern Vietnam; extreme northwestern West Malaysia

Draco timorensis Kuhl, 1820 Draco timorensis Kuhl, 1820: 103 M O’Shea Type locality: Timor Holotype : BMNH 1844.2.22.89 Distribution: Indonesia: Timor, Roti, Alos, Semau, Wetar; Timor-Leste

Draco volans Linnaeus, 1758 Draco volans Linnaeus, 1758:109

C Sharp Synonyms Draco praepos Linnaeus, 1758:358 Type locality: unknown Type: Iconotype: Seba I.t.102.f.2 Draco major Laurenti, 1768:50 Type locality: unknown Type: Iconotype: Seba I.86.3 Draco minor Laurenti, 1768:51 Type locality: unknown Type: Incontype: Seba II.102.2 Draco viridis Daudin, 1802:301 Type locality: Java, Indonesia Type: MNHP, not found Draco fuscus Daudin, 1802: 307 Type locality: unknown Type: MNHP, not found Draco daudinii Duméril & Bibron, 1837: 451 Type locality: Java, Indonesia Syntypes: MNHP 0.195, 0.193 Draco viridis javanica Schlegel, 1844:91 Type locality: Java, Indonesia Lectotype: RMNH 2934 A , designated by Musters 1983 Draco viridis samaoensis Schlegel, 1844: 92 Type locality: Semau Island (= Pasar Pusmau),Lesser Sunda Islands, Indonesia Holotype: RMNH 2903 Type locality: Java. Indonesia Syntypes: NRM 104-105 Distribution: Indonesia: Kalimantan (Borneo), Sumatra, Mentawei and Rhiao Archipelagos, Java, Natuna Islands, Sulawesi, Timor; West Malaysia, East Malaysia: Sarawak; Thailand; Singapore; Vietnam.

Draco walkeri Boulenger, 1891 Draco walkeri Boulenger, 1891:279 Type locality: ‘‘Koepang, Timor’’, in error Lectotype: BMNH 1946.8.26.94, designated by Musters, 1983 Distribution: Indonesia: Sulawesi

Gonocephalus Kaup, 1825 Kaup, J.J. 1825. "Einige Bemerkungen zu Merrems Handbuch ." Isis von Oken 16: 589-593. Type species: Lophyrus tigrinus Duméril & Bibron, 1837, by monotypy

Gonocephalus abbotti Cochran, 1922 Gonocephalus abbotti Cochran, 1922: 1 Type locality: Trang, southern Thailand Holotype: USNM 24028 Distribution: West Malaysia: Pahang; southern Thailand: Pattani, Trang, Yala; southwestern Cambodia

Gonocephalus bellii (Duméril & Bibron, 1837) Lophyrus bellii Duméril & Bibron, 1837: 416

I Das Type locality: “ Bengale” Holotype: MNHP 0.6897 Distribution: Thailand: between Surat Thani and Nakhon Si Thammarat; West Malaysia: Malacca, Perak, Pahang , Selangor

Gonocephalus beyschlagi Boettger, 1892 Gonocephalus beyschlagi Boettger, 1892: 104 Type locality: Langkat in Deli, northeastrtn Sumatra Holotype: SMF 9760 Distribution: Indonesia: North Sumatra

Gonocephalus bornensis (Schlegel, 1848) Lophyrus bornensis Schlegel, 1848: 6

I Das Synonyms Tiaris miotympanum Günther, 1872:592 Type locality: Pulau Labuan (05°19’N; 115°13’E), a Federal Territory off eastern Sarawak, East Malaysia Syntypes : BMNH 1946.8.19.82 and 1946.8.14.64 Gonocephalus denzeri Manthey, 1991: 153 Type locality: Kapit. 3. Division (02°00’N; 112°54’E), Sarawak, Borneo, East Malaysia Holotypes : ZFMK 50527 Type locality: Borneo, Indonesia Syntypes: RMNH 3043-44 Distribution: Indonesia: Kalimantan (Borneo); East Malaysia: Sabah, Sarawak

Gonocephalus chamaeleontinus (Laurenti, 1768) Iguana chamaeleontina Laurenti, 1768: 47

I I Das Synonyms Agama gigantea Kuhl, 1820: 106 Type locality: ‘India’ type: Iconotype: Seba Agama tigrina Merrem, 1820: 590 Type locality: Indonesia Type: UUZM, not located Galeotes lophyrus Schlegel, 1844 :73 Type locality: Ambon Island, Indonesia Type: RMNH, not located Lophyrus sumatranus Schlegel, 1848: 5 Type locality: Palembang, South Sumatra Type: RMNH, not located Type locality: Asia (“America” in error) Holotype: MNHP 0.2564 Distribution: Indonesia: Mentewei Island, Sumatra, Bangka, Natuna Island, Java, Nias, Sipora, Siberut, Bangkaru, Tuangku, Banyak Islands, Tanahmasa; West Malaysia: Pahang

Gonocephalus doriae (Peters, 1871) Gonyocephalus doriae Peters, 1871: 570

I Das Type locality: Sarawak, Borneo, East Malaysia Syntypes: ZMB 7147, MSNG 29152 Distribution: Indonesia: Kalimantan (Borneo); East Malaysia: Sarawak, Sabah

Gonocephalus grandis (Gray, 1845) Dilophyrus grandis Gray, 1845:239

I Das Type locality: Rangoon, Burma Syntypes: BMNH 1946.8.14.83,1946.9.4.26 Distribution: southern Thailand; Vietnam; West Malaysia: Pulau Tioman, Pulau Pinang; Indonesia: Sumatra, Mentawei Archipelago : Nias, Nako, Sipora; Kalimantan; East Malaysia: Sarawak

Gonocephalus interruptus (Boulenger, 1885) Gonyocephalus interruptus Boulenger, 1885: 290 Type locality: Mindanao, Philippines Holotype: BMNH 1884.12.31.1 Distribution: Philippines: Mindanao, Mindoro, Danguan, Aninuan

Gonocephalus klossi (Boulenger, 1920) Gonyocephalus klossi Boulenger, 1920: 287 Type locality: Siolak Daras, Korinchi Valley, Sumatra, Indonesia Holotype: BMNH 1946.8.14.65 Distribution: Indonesia: western Sumatra

Gonocephalus kuhlii (Schlegel, 1848) Lophyrus kuhlii Schlegel, 1848: 4 Type locality: Java, Sumatra Syntypes: RMNH, not found Distribution: Indonesia: Sumatra, Java

Gonocephalus lacunosus Manthey & Denzer, 1991 Gonocephalus lacunosus Manthey & Denzer, 1991: 3 Type locality: stream next to Sikulikap Waterfall at ca. 1200 m elevation, near Berastagi, North Sumatra, Indonesia Holotype: ZMB 48616 Distribution: Indonesia: North Sumatra Last seen 1990

Gonocephalus liogaster (Günther, 1872) Tiaris liogaster Günther, 1872: 592 Synonym Goniocephalus herveyi Boulenger, 1887:493 Type locality: Laut Island, Natuna Islands, Indonesia Holotype: BMNH 1946.8.14.80 Type locality: Sarawak, Borneo, East Malaysia Syntypes: BMNH 1946.8.14.82, 1946.8.27.16-17, 1946.8.13.86, 1946.8.14.79 Distribution: Indonesia: Sumatra, Kalimantan (Borneo), Natuna Island); East Malaysia: Sarawak, Sabah

Gonocephalus megalepis (Bleeker, 1860) Lophyrus megalepis Bleeker, 1860: 327 Synonym Tiaris tuberculatus Günther, 1872:538 Type locality: Sumatra Holotype: BMNH 1946.8.27.21 Type locality: Agam, Sumatra, Indonesia Holotype: RMNH 3954 Distribution: Indonesia: Sumatra

Gonocephalus mjobergi Smith, 1925 Gonocephalus mjobergi Smith, 1925: 7 I Das Type locality: Mount Murud (03°55’N; 115°32’E), Sarawak, Borneo, East Malaysia Holotype: BMNH 1946.8.13.87 Distribution: Indonesia: Kalimantan (Borneo); East Malaysia: Sarawak

Gonocephalus semperi (Peters, 1867) Lophyrus semperi Peters, 1867: 16 Type locality: Mindanao, Philippines Syntypes: NMHW 23352:1, 14955:1-2; ZMB 5577, ZMB 54501 Distribution: Philippines: Mindanao, Mindoro, Bohol

Gonocephalus sophiae (Gray, 1845) Tiaris sophiae Gray, 1845: 240 Type locality: Philippine Islands Syntypes: BMNH 1946.8.27.10 -15 Distribution: Philippines: Negros, Mindanao, Panay, Masbate, Luzon

Gowidon Wells, R.W. & C.R. Wellington. 1984 . Wells, R.W. & C.R. Wellington. 1984. A synopsis of the class Reptilia in Australia. Australian Journal of Herpetology 1(3-4): 73-129 [1983 on title page]. Type species: Lophognathus longirostris Boulenger, 1883 by original designation

Gowidon longirostris (Boulenger, 1883) Lophognathus longirostris Boulenger, 1883: 225

S Zozaya Synonyms Physignathus eraduensis Werner, 1909: 275 Type locality: Eradu,Western Australia. Holotype: not found Physignathus longirostris quattuorfasciatus Sternfeld,1924:236 Type locality: Hermannsburg Misson, Northern Territory. Lectotype: SMF 10366, designated by Mertens 1967 Type locality: Champion Bay and Nickol Bay, Western Australia Syntypes: BMNH 1946.8.12.64–65 (Nickol Bay); BMNH 1946.8.28.73 (Champion Bay) Distribution: Australia: central and southern North Territory, west-central Queensland, northern South Australia, West Australia north to Fitzroy River and south to the rim of the Great victoria Desert and the Murchison River

Harpesaurus Boulenger, 1885 Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae . London: 450 pp Type species: Arpephorus tricinctus Duméil, in Duméril & Duméril, 1851, by monotypy

Harpesaurus beccarii Doria, 1888 Harpesaurus beccarii Doria, 1888: 646

unknown Type locality: Lubu Selassi near Padang, Sumatra, Indonesia Holotype: MSNG 28489 Distribution: Indonesia: Sumatra

Harpesaurus borneensis (Mertens, 1924) Hylagama borneensis Mertens, 1924: 157 Synonym Harpesaurus thescelorhinos King, 1978: 205 Type locality: Kuala Tangap, below Kooboo Niah and 1/2 mile below (03° 52’N; 113° 43’E), Sarawak, Borneo, East Malaysia Holotype : FMNH 131540 Type locality: central area of Kapuas River (00°25’S; 109°40’E), Kalimantan, Indonesia Holotype: SMNS 4596 Distribution : Indonesia: Kalimantan (Borneo); East Malaysia: Sarawak,

Harpesaurus ensicauda Werner, 1913 Harpesaurus ensicauda Werner, 1913:8 Type locality: Pulo, Nias Island, Indonesia Holotype : ZMH, lost Distribution: Indonesia: Nias Last seen 1912

Harpesaurus modiglianii Vinciguerra, 1933 Harpesaurus modiglianii Vinciguerra, 1933: 356 Type locality: Si-Rambé Forest, North Sumatra, Indonesia Holotype: MSNG 32191 Distribution: Indonesia: Sumatra Last seen 1933

Harpesaurus tricinctus (Duméril, 1851) Arpephorus tricinctus Duméril in Duméril & Duméril, 1851: 93 Type locality: Java, Indonesia Holotype: MNHP 0.623 Distribution: Indonesia: Java

Hydrosaurus Kaup, 1828 Kaup, J.J. 1828. Über Hyaena, Uromastix, Basiliscus, Corythaeolus, Acontias. Isis 21, columns 1144–1150. Type species : Lacerta amboinensis Schlosser, 1768, by original designation

Hydrosaurus amboinensis (Schlosser, 1768) Lacerta amboinensis Schlosser, 1768: 19

M O’Shea Synonyms Lacerta lophura Shaw, 1802: 218 Type locality: unknown Holotype: BMNH 1946.8.11.61 Istiurus microlophus Bleeker, 1860:85 Type locality: Makassar, Sulawesi, Indonesia Holotype: RMNH 5037 Lophura celebensis Peters, 1872: 581 Type locality: Poso River, Tomini Bay, Sulawesi, Indonesia Holotype: ZMB 7393 Type locality: Ambon Island, Indonesia Holotype: NRM 1047 Distribution: Indonesia: Ambon, Halmahera, Sulawesi, Togian Islands, Buton, Ceram, Batjan, Ternate, Waigeo, Papua; Papua New Guinea

Hydrosaurus pustulatus (Eschscholtz, 1829) Istiturus pustulatus Eschscholtz, 1829: 2 M O’Shea Type locality: Luzon Island, Philippines Syntype: ZMB 66245 Distribution: Philippines: Luzon, Polillo, Mindoro, Negros, Dinagat, Mindanao, Panay, Bohol

Hydrosaurus weberi Barbour, 1911 Hydrosaurus weberi Barbour, 1911:20 Type locality: Weeda, Halmahera Island, Maluku, Indonesia Holotype: MCZ 7486 Distribution: Indonesia: Halmahera, Ternate

Hypsicalotes Manthey & Denzer, 2000 Manthey, U. & W. Denzer 2000. Description of a new genus, Hypsicalotes gen. nov. (Sauria: Agamidae) from Mt. Kinabalu, North Borneo, with remarks on the generic identity of Gonocephalus schultzewestrumi Urban, 1999. Hamadryad 25 (1): 13-20 Type species: Calotes kinabaluensis De Grijs, 1937, by original designation

Hypsicalotes kinabaluensis (De Grijs, 1937) Calotes kinabaluensis De Grijs, 1937: 136

B Dupont Type locality: Kinabalu Park, Sabah, Borneo, Malaysia Neotype : KUZ 31291 Distribution: East Malaysia: Sabah

Hypsilurus Peters, 1867 Peters, 1867 : Über Flederthiere (Pteropus gouldii, Rhinolophus deckenii, Vespertilio lobipes, Vesperugo temminckii) und Amphibien (Hypsilurus godeffroyi, Lygosoma scutatum, Stenostoma narirostre, Onychocephalus unguirostris, Ahaetulla poylepis, Pseudechis scutellatus, Hoplobatrachus reinhardtii, Hyla coriacea) . Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, vol. 1867, p. 703-712 Type species: Lophura godeffroyi Peters, 1867 by monotypy

Hypsilurus auritus (Meyer, 1874) Gonyocephalus auritus Meyer, 1874: 130 Type locality: Ansus , Yapen Island (1º45’S.36º15E), West Papua Province, Indonesia Holotype: ZMB 8782 Distribution: Indonesia: Yapen, Misol, Salawati, Batanta Islands as well as mainly coastal areas of the main island, West Papua Province, New Guinea; Papua New Guinea: Bogadjim in Astrolabe Bay, Madang Province; Eastern and Western Highlands, Chimbu Province, West Sepik Province

Hypsilurus binotatus (Meyer, 1874) Gonyocephalus binotatus Meyer, 1874: 130 Type locality: Ansus , Yapen Island (1º45’S.36º15E), West Papua Province, Indonesia Holotype: MTKD, lost in WWII Distribution: Indonesia: Aru Islands, Kei Islands, Yapen Island; Papua Province: Vogelkopf, Wondowoi mountains, Wandammen Peninsula, Tami River; Papua New Guinea: Gulf, West and East Sepik, Western Provinces

Hypsilurus boydii (Macleay, 1884) Tiaris boydii Macleay, 1884: 432

S Zozaya Type locality: Herbert River, Ripple Creek district, Queensland, Australia Syntypes: AMS R31884–85 Distribution: Australia: northeastern Queensland

Hypsilurus bruijnii (Peters & Doria, 1878) Gonyocephalus bruijnii Peters & Doria, 1878: 379 Type locality: Arfak mountains, Doberai (Vogelkop) Peninsula, West Papua Province , Indonesia Holotype : MSNG 9562 Distribution: Indonesia: West Papua province: Arfak mountains

Hypsilurus capreolatus Kraus & Myers, 2012 Hypsilurus capreolatus Kraus & Myers, 2012 : 396 Type locality: Parkop (3.42457ºS, 142.51866ºE), Torricelli Mountains, West Sepik Province, Papua New Guinea Holotype: BPBM 23413 Distribution: Papua New Guinea: West Sepik province

Hypsilurus dilophus (Duméril & Bibron, 1837) Lophyrus dilophus Duméril & Bibron, 1837: 419

M O’Shea Synonym Tiaris megapogon Gray, 1845: 239 Type locality: New Guinea Syntypes: MNHP, not found; RMNH, not found Type locality: New Guinea Holotype: MNHP 0.2561 Distribution: Indonesia: Papua provinces, Aru Islands, Kei Islands, Batanta, Salawati, Numfoor, Yapen; Papua New Guinea: all provinces, Ferguson Island, Normanby Island.

Hypsilurus geelvinkianus (Peters & Doria, 1878) Gonyocephalus geelvinkianus Peters & Doria, 1878: 381 Type locality: Numfor Island (1º03’S, 134º54’E), and Biak Island (1ºS, 136ºE), Cenderawasih Bay, West Papua Province , Indonesia Syntypes: ZMB 9729-30; MSNG 29144; NMHW 21050 Distribution: Indonesia: West Papua Province: Windiwoi Mts; Numfoor Island, Biak Island; Papua New Guinea: East Sepik Province

Hypsilurus godeffroyi (Peters, 1867) Lophura godeffroyi Peters, 1867: 707 Type locality: islands off the northern New Guinea coast Lectotype: ZMB 5891, designated by Marx 1958 Distribution: Papua New Guinea: Bismarck Archipelago, Admirality Islands: Palau Island

Hypsilurus hikidanus Manthey & Denzer, 2006 Hypsilurus hikidanus Manthey & Denzer, 2006: 28 Type locality: Enarotali, Wisselmeren, West Papua Province, Indonesia Holotype: RMNH 28905 Distribution: Indonesia: West Papua

Hypsilurus longi (Macleay, 1877) Tiaris longii Macleay, 1877: 103 Type locality: Northern Queensland, Australia Holotype: AMS R 31935 Distribution: Australia: northern Cape York, Queensland; Papua New Guinea: New Britain, Bougainville, New Ireland; Solomon Islands: Shortland Islands

Hypsilurus macrolepis Peters, 1872 Hypsilurus macrolepis Peters, 1872: 775 Type locality: southern Solomon Islands Holotype: ZMB 7510 Distribution: southern Solomon Islands: Isabel, Makira and adjacent smaller islands, Malaupaina, Olu Malau Ids., Ugi, Santa Ana and Uki Ni Masi

Hypsilurus magnus Manthey & Denzer, 2006 Hypsilurus magnus Manthey & Denzer, 2006:26 Type locality: Gariau near Lake Jamur (3°42’S 134°56’E), New Guinea, West Papua Province, Indonesia Holotype: RMNH 29093 Distribution: Indonesia: Papua Province; Papua New Guinea: Prince Alexander Mts., Madang, West Sepik province

Hypsilurus modestus (Meyer, 1874) Gonyocephalus modestus Meyer, 1874: 130 Synonyms Gonyocephalus inornatus Doria, 1874: 345 Type locality: Wokam, Aru Islands, Indonesia Syntype: ZMB 7941 Diptychodera lobata Boettger, 1893: 45 Type locality: New Britain, Bismark Archipelago, PNG Holotype: SMF 9778 Gonyocephalus modestus carinatus Sternfeld, 1918: 393 Type locality: Anir Island, New Ireland Province, PNG Lectotype: SMF 9770, designatd by Mertens 1967 Type locality : Yapen Island, New Guinea, Papua Province, Indonesia Holotype: ZMB 8783 Distribution: Indonesia: Aru Island, Yapen Island; Papua New Guinea: Bismarck Archipelago, Admirality Islands

Hypsilurus nigrigularis (Meyer, 1874) Gonyocephalus nigrigularis Meyer, 1874: 129 Type locality: Rubi (~ 3° 23’S; 135° 15’E), south of Cenderawasih Bay, West Papua Province, Indonesia. Holotype: MTKD 2967 Distribution: Indonesia: West Papua Province, New Guinea Last seen 1874

Hypsilurus ornatus Manthey & Denzer, 2006 Hypsilurus ornatus Manthey & Denzer, 2006: 30 Type locality: Didessa Village, Mt. Bosavi (6°35’S, 142°50’E), Southern Highlands Province, Papua New Guinea Holotype: ZSM 122/1979 Distribution: Papua New Guinea: Southern Highlands Province Last seen 1989

Hypsilurus papuensis (Macleay, 1877)

M O’Shea Subspecies: Hypsilurus papuensis papuensis (Macleay, 1877) Tiaris papuensis Macleay, 1877: 101 Synonym Gonyocephalus albertisii Peters & Doria, 1878:377 Type locality: Nikura, opposite Yule Island, Hall Sound (8°48’S, 146°37’E), Central Province, PNG Syntypes: ZMB 9722, 9724-25 Type locality: Hall Sound (8°49’S 146°36’E), Central Province, Papua New Guinea Holotype: AMS R 31883 Distribution: Papua New Guinea: coastal Central Province

Hypsilurus papuensis longicauda Manthey & Denzer, 2006 Hypsilurus papuensis longicauda Manthey & Denzer, 2006: 21 Holotype: ZMB 21367 Type locality: Sattelberg (6º29S, 147º47E), Huon Peninsula, Morobe Province, Papua New Guinea Distribution: Papua New Guinea: coastal Madang and Morobe provinces

Hypsilurus schoedei (Vogt, 1932) Gonyocephalus schoedei Vogt, 1932: 282 Type locality: Lambussu Island (=Rambutyo) (2º18’S, 147º47’E), Admirality Islands, Papua New Guinea Syntypes: ZMB 32205, ZMB 54474–500 Distribution: Papua New Guinea: Admirality Islands

Hypsilurus schultzewestrumi (Urban, 1999) Gonocephalus schultzewestrumi Urban, 1999: 185 Type locality: near Nondugl( 5°52’S, 144°46’E), Papua New Guinea Holotype: ZSM 1965/82 Distribution: Papua New Guinea Central Range; Indonesia: West Papua Province, central ranges

Hypsilurus spinipes (Duméril & Bibron, 1851) Lophyrus spinipes Duméril & Bibron, 1851: 90

S Zozaya Type locality: Australia Holotype : MNHP 0.2560 Distribution: Australia: northeastern New South Wales, southeastern Queensland

Hypsilurus tenuicephalus Manthey & Denzer, 2006 Hypsilurus tenuicephalus Manthey & Denzer, 2006: 29 Type locality: highlands of Western Papua Province, Indonesia Holotype: MTKD D 30608 Distribution: Indonesia: highlands of Western Papua

Intellagama Wells & Wellington, 1985 Wells, R.W. & Wellington, C.R. 1985. A classification of the Amphibia and Reptilia of Australia. Australian Journal of Herpetology Supplementary Series 1: 1-61 Type species: Lophura lesueurii Gray, 1831 by original designation

Intellagama lesueurii (Gray, 1831)

S Macdonald Subspecies: Intellagama lesueurii lesueurii (Gray, 1831) Lophura lesueurii Gray, 1831: 60 Synonyms Amphibolurus maculiferus Girard, 1857: 199 Type locality: About fences at Patrick's Plains, New South Wales Type: ANSP, not located Amphibolurus heterurus Peters, 1866: 86 Type locality: Clarence River, NSW Syntypes: ZMB 5321–22 Amphibolurus branchialis De Vis, 1884:55 Type locality: neighbourhood of Brisbane, QLD Type: QM, lost Type locality: Paramattan, New South Wales, Australia Holotype: MNHP 0.392 Distribution: Australia: New South Wales, Queensland

Intellagama lesueurii howittii (McCoy, 1884) Physignathus lesueuri howitti McCoy, 1884: 7 Lectotype : NMV D1822 Type locality: Upper Buchan River, Victoria, Australia Distribution: Australia: from Gippsland, Victoria, to Kangaroo Valley, New South Wales

Japalura Gray. 1853 Gray, J. E. 1853. Descriptions of some undescribed species of reptiles collected by Dr. Joseph Hooker in the Khassia Mountains, East Bengal, and Sikkim Himalaya. Annals and magazine of natural history (2) 12: 386 – 392 Type species: Japalura variegata Gray, 1853, by original designation

Japalura andersoniana Annandale, 1905 Japalura andersoniana Annandale, 1905: 85 Type locality: Dafla Hills, Assam-Bhutan border Holotype: ZSI 4104 Distribution: India: Assam, Dafla Hills, Abor County, Arunachal Pradesh: Bhutan; China: Xizang

Japalura dasi (Shah & Kästle, 2002) Oriotiaris dasi Shah & Kästle in Schleich & Kästle, 2002: 608 Type locality: near Agaupani (29°26’54’’N, 81°36’11”E), Bajhura District, far western Nepal. Holotype: ZSM 855/2012 Distribution: western Nepal Last seen 1997

Japalura flaviceps Barbour & Dunn, 1919 Japalura flaviceps Barbour & Dunn, 1919: 16 Type locality: Tung River (ca. 30.6006° 101.4621°), western Szechuan, China Holotype: MCZ R-12469 Distribution: China: western Sichuan, eastern Xizang, Qinghai, Gansu, Shenxi, Shanxi, northern Yunnan

Japalura kumaonensis (Annandale, 1907) Acanthosaura kumaonensis Annandale, 190: 152

Hudong Type locality: Naini Tal , Burma Holotype: ZSI 15755 Distribution: northern India: Himachal Predesh, Uttaraanchai; Pakistan: NWFP; Nepal; China: Xizang (= Tibet)

Japalura major (Jerdon, 1870) Oreocalotes major Jerdon, 1870: 77

jgdhakim Type locality: valley of the Sutlej, near Kotgarh, western Himalaya, India Holotype: BMNH 1946.8.14.7 Distribution : northern India: Himachal Pradesh; Nepal

Japalura planidorsata Jerdon, 1870 Japalura planidorsata Jerdon, 1870: 76 Type locality: Khasi Hills, India Syntypes: BMNH 1946.8.30.86–87 Distribution: Burma: Chin state; India: Assam, Meghalaya

Japalura sagittifera Smith, 1940 Japalura sagittifera Smith, 1940: 477 Type locality: Pangnamdim, Dadung, northern Burma Lectotype: BMNH 1946.8.13.97, designated by Mahony 2009 Distribution: Burma; India: Arunachal Pradesh

Japalura tricarinata (Blyth, 1853) Calotes tricarinatus Blyth, 1853: 650 Synonym Tiaris elliotti Günther, 1860: 151 Type locality: Sikkim Holotype: BMNH 1946.8.28.9 Type locality: Darjeeling (27º 02'N; 88º 16'E), West Bengal State, eastern India Holotype: ZSI 5300 Distribution: northern India: Sikkim, West Bengal; eastern Nepal; China: Tibet

Japalura variegata Gray, 1853 Japalura variegata Gray, 1853: 388 Synonyms Biancia niger Gray,1853:388 Type locality: Sikkim Holotype: BMNH 1853.8.12.5 Japalura microlepis Jerdon, 1870: 76 Type locality: Sikkim Holotype: BMNH 1946.8.14.10 Japalura bengalensis Anandale, 1912: 57 Type locality: Buxa, Jalpaiguri district, at the border of West Bengal State, eastern India and Bhutan Holotype: ZSI 12564 Type locality: Sikkim, in the Himalaya mountains, northern India Syntypes: BMNH 1946.8.14.4-5 Distribution: northern India: Sikkim, West Bengal; Nepal; Bhutan

Laudakia Gray, 1845 Gray, J.E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum . Trustees of the British Museum/Edward Newman, London: xxvii + 289 pp. Type species : Agama tuberculata Gray, 1827, by montypy.

Laudakia agrorensis (Stoliczka, 1872) Stellio agrorensis Stoliczka, 1872: 128 Type locality: Sussel Pass, at the entrance into Agror Valley, Hazara district, northwestern Punjab, Pakistan Syntypes: ZSI 4206, NMHW 16754, BMNH 1880.11.10.9 Distribution: eastern Afghanistan; northwester India: Punjab, Himachal Pradesh, Kashmir; northern Pakistan

Laudakia badakhshana (Anderson & Leviton, 1969) Agama badakhshana Anderson & Leviton 1969: 33 Type locality: Mazar-i-Sharif (36° 34’ N, 67° 05’ E), northern Afghanistan Holotype: FMNH 161108 Distribution: northeastern Afghanistan; northern Pakistan; northern India; China: Xinjiang; southeasternTurkmenistan, eastward through Tajikistan to western Kyrgyzstan

Laudakia bochariensis (Nikolsky, 1897) Stellio bochariensis Nikolsky, 1897: 159 Synonyms Agama isozona Werner, 1899:479 Type locality: Margelan, Turkestan Holotype: NMHW 23343 Agama chernovi Ananjeva, Peters & Rzepakovsky, 1981: 23 Type locality: vicinity of the Dashti-Khonako in the Surkhku mountain ridge near Nurek, Tajikistan Holotype. ZISP 19385 Type locality: Sarym-Sakhly, Sughd province, Tajikistan Syntypes: ZISP 7332, ZISP 8897 Distribution: southwestern Tajikistan; Uzbekistan; southeastern Turkmenistan; southern Kyrgyzstan

Laudakia caucasia (Eichwald, 1831) Stellio caucasicus Eichwald, 1831: 187

P Weigell Synonyms Stellio persicus Anderson,1872: 382 Type locality: Teheran, Iran Type: lost Agama reticulata Nikolsky, 1912: 272 Type locality: Chubek, Khatlon province, Tajikistan Holotype: ZISP 10718 Type locality: Tbilisi, Georgia, and Baku, Azerbaijan Syntypes: not located Distribution: southern Russia: Caucasus, Dagestan; eastern and southern Georgia; Armenia; Azerbaijan; southwestern Turkmenistan; Tajikistan; eastern Turkey, northern Iraq; northern Iran; Afghanistan; northwestern Pakistan: Waziristan, Baluchistan

Laudakia dayana (Stoliczka, 1871) Stellio dayanus Stoliczka, 1871: 194 Type locality: Hardwar, Uttarakhand, India Holotype: ZSI 6780 Distribution: northwest India: Uttranchal, Kashmir

Laudakia erythrogaster (Nikolsky, 1896) Stellio erythrogaster Nikolsky, 1896: 370 Synonyms Stellio erythrogaster pallida Nikolsky, 1897: 319 Type locality: Mashhad, Razavi Khorasan province, Iran Holotype. ZISP 8761 Agama caucasica mucronata Guibé, 1957: 137 Type locality:. Langarak, 60 km east of Mashad, on the route of the Sarakhs, Iran Syntypes: MNHP 1957.24-26 Stellio erythrogaster nurgeldievi Tuniyev, Atayev & Shammakov, 1991: 50 Type locality: eastern Kopet-Dagh, Turkmenistan. Holotype: ZMMU R -248 Type locality: Fariman, Khorasan Province, Iran Lectotype: ZISP 8760, designated by Rastegar-Pouyani & Nilson 2002 Distribution: northeastern Iran; northern Afghanistan; southeastern Turkmenistan

Laudakia himalayana (Steindachner, 1867) Stellio himalayanus Steindachner, 1867: 22 D Ash Type locality: Leh and Kargill, Ladakh Province, Kashmir, India Syntypes : NMHW 16752:1-12 Distribution: northeastern Afghanistan; northern Pakistan; northern India: Kashmir; Nepal; China: Xinjiang, Tibet; Turkmenistan; western Tajikistan; Kirgizstan

Laudakia lehmanni (Nikolsky, 1896) Stellio lehmanni Nikolsky, 1896:14

D Melnikov Synonym Agama borstschewskyi Elpatjewsky & Sabanrjew, 1907: 250 Type locality: Buchara region, Bukhara Province, Uzbekistan Holotype: ZMMU R -248 Type localities: [see below] Syntypes: ZISP 2367 (Oalyk mountain, Samarqand province, Uzbekistan); ZISP 8664 (Zerbent river, Nuratau mountains, Jizzakh province, Uzbekistan); ZISP 8665 (Temur-Kauk well, Nuratau mountains, Jizzakh province,Uzbekistan); ZISP 8666.1, 8666.2, 8689.1, 8689.2 (Kum-Kuduk well, southern part of Kyzylkum desert, Uzbekistan Distribution: eastern Turkmenistan, eastern Uzbekistan, western Tajikistan, southern Kyrgyzstan, northern Afghanistan

Laudakia melanura Blyth, 1854 Subspecies: Laudakia melanura melanura Blyth, 1854 Laudakia melanura Blyth, 1854: 738 Synonym Stellio liratus Blanford, 1874: 453 Type locality: Saman, Dasht province, Baluchistan, Pakistan Holotype: ZSI 3410 Type locality: Salt Range, Punjab, Pakistan Type: BMNH, lost Distribution: Pakistan: Salt Range in northern Punjab, Waziristan in NW Frontier province, western Sind and Baluchistan provinces; southeastern Iran; southeastern Afghanistan

Laudakia melanura nasiri Baig, 1999 Laudakia melanura nasiri Baig, 1999: 81 Holotype: PMNH 735 Type locality: Tanishpa (31º12’N, 68º28’E), Dist. Killa Saifullah, Toba Kakar Range, Balochistan, Pakistan Distribution: southern Pakistan: Toba Kakar Range, Balochistan

Laudakia microlepis (Blanford, 1874) Subspecies: Laudakia microlepis microlepis (Blanford, 1874) Stellio microlepis Blanford, 1874: 453 Synonym Stellio caucasica triannulatus Ananjeva & Atajev, 1984: 4 Type locality: Meshed sand area near Madau village, southwestern Turkmenistan Holotype: ZISP 19640 Type locality: Khaneh-Sorkh Pass, between Sirjan and Kerman, Iran Lectotype: BMNH 1946.8.28.74, designated by Rastegar-Pouyani & Nilson 2002 Distribution: southern Iran; Afghanistan; western Pakistan; southern Turkmenistan

Laudakia microlepis taftanica Sanchooli, Rahimian, Rastergar-Pouyani & Rastergar-Pouyani, 2016 Laudakia microlepis taftanica Sanchooli, Rahimian, Rastergar-Pouyani & Rastergar-Pouyani, 2016: 200 Type locality: 25 km north of Khash (28°36.0'25.1"N, 61° 04' 47.3"E) , Sistan and Baluchistan province, southeastern Iran Holotype: ZUTC rep1438 Distribution: Taftan Mountain, in Sistan and Baluchistan province, Iran .

Laudakia nupta (De Filippi, 1843) Subspecies: Laudakia nupta nupta (De Filippi, 1843) Agama nupta De Filippi, 1843: 407 Synonyms Stellio carinatus Duméril, 1851: 107 Type locality: Iran MNHP, not found Agama flavicauda Werner, 1897: 264 Type locality: unknown Syntypes: ZMB 13723, ZMB 54608-09 Type locality: Persepolis, Fars province, Iran Syntypes: MSNTO R322; ZSI 6798; ZMB 6078 Distribution: eastern Iraq; Iran; Afghanistan; Pakistan

Laudakia nupta fusca (Blanford, 1876) Stellio nuptus fuscus Blanford, 1876: 319 Type locality: near Jalk, Sistan-Baluchistan Province, Iran Lectotype: BMNH 1874.11.32.11, designated by Rastegar-Pouyani & Nilson 2002 Distribution: southern Pakistan; southern Iran

Laudakia nuristanica (Anderson & Leviton, 1969) Agama nuristanica Anderson & Leviton, 1969: 39 Type locality: Kamdesh, eastern Afghanistan Holotype: FMNH 161136 Distribution: eastern Afghanistan; northwestern Pakistan

Laudakia pakistanica (Baig, 1989) Subspecies: Laudakia pakistanica pakistanica (Baig, 1989) Agama pakistanica Baig, 1989: 117 Type locality: Jaglotgah (36°20'N, 74°S0'E), Gilgit; FANA Holotype: PMNH 0134 Distribution: Gilgit–Baltistan, northern Pakistan

Laudakia pakistanica auffenbergi Baig & Böhme, 1996 Laudakia pakistanica auffenbergi Baig & Böhme, 1996: 3 Type locality: Besham (35°55’N, 72°55’E), Distr. Swat, Northwest Frontier Province, Pakistan Holotype: ZFMK 54471 Distribution: Pakistan: Mansehra Besham and Kohistan districts, Northwestern Frontier Province

Laudakia pakistanica khani Baig & Böhme, 1996 Laudakia pakistanica khani Baig & Böhme, 1996: 1 Type locality: Hadar (Chilas) (32°25’N, 74°E), Federal Administered Northern Area, Pakistan Holotype: ZFMK 54469 Distribution: Pakistan: Chilas, Federal Administered Northern Areas

Laudakia papenfussi Zhao, 1998 Laudakia papenfussi Zhao, 1998: 401 Type locality: Mayang River Valley between Mayang Village and Diya Village, Zanda Co., Xizang Autonomous Region, China Holotype: CIB 775001 Distribution: China: Xizang = Tibet

Laudakia sacra (Smith, 1935) Agama himalayana sacra Smith, 1935: 214 Type locality: near Lhasa, Tibet Lectotype: BMNH 1946.8.28.57, designated by Ananjeva et al. 1990 Distribution: China: Xizang = Tibet

Laudakia stellio (Linnaeus, 1758)

M Pickersgill Subspecies: Laudakia stellio stellio (Linnaeus, 1758) Lacerta stellio Linnaeus, 1758:202 Synonyms Iguana cordylina Laurenti, 1768: 47 Type locality: Delos or Mykonos, Greece Icontype: Seba I, 107.2 Agama stellio mykonensis Xyda, 1983: 115 Type locality: Delos Syntypes: lost Type locality: Delos Island, Cyclades, Greece Neotype: ZFMK 2063, designated by Crochet et al. 2006 Distribution: Greece : Delos Island, Cyclades

Laudakia stellio brachydactyla (Haas, 1951) Agama stellio brachydactyla Haas, 1951: 1052 Type locality: Jebel Lussan on the border between Israel and Sinai, south-southwest of Beer-Sheba, Egypt Holotype: HUJ-R 1801 Distribution: northern Saudi Arabia; southern Israel; Egypt: Sinai; Jordan

Laudakia stellio cypriaca (Daan, 1967) Agama stellio cypriaca Daan, 1967: 127 Type locality: Limasol, Cyprus Holotype: BMNH 1930.10.5.6 Distribution: Cyprus

Laudakia stellio daani (Beutler & Frör, 1980) Agama stellio daani Beutler & Frör, 1980: 270 Type locality: between Agh. Kirikos and Evdilos, Ikaria, Samos Region, Greece Holotype: ZSM 201/1978 Distribution: Greece: islands in the Aegian Sea (Antiparos, Despotiko, Naxos, Paros, Samos, Thimena, Syrni, Chalki, Kalymnos, Kastelorizo, Kos, Leros, Nisyros, Rhodos, Symi, Telendos, Tilos, Lesbos, Chios, Fourni, Ikaria); western and southwestern Turkey; Macedonia

Laudakia stellio picea (Parker, 1935) Agama stellio picea Parker, 1935: 137 Type locality: Black Lava Desert of Transjordania (32° 10’ N × 36° 40’ E), Jordan Syntypes: BMNH 1946.8.28.53-56 Distribution: southwestern Syria; southern Lebanon; northern Israel; northwestern Jordan

Laudakia stellio salehi (Werner in Lachmann et al., 2006) stellio salehi Werner in Lachmann et al., 2006: 1271 Type locality: 3 km from Sheikh Harun towards Watiya Pass (28° 35’ 30’’ N, 33° 59’ ), Sinai, Egypt Holotype: HUJ-R 10732 Distribution: Egypt: Sinai

Laudakia stellio vulgaris (Sonnini & Latreille, 1802) Stellio vulgaris Sonnini & Latreille, 1802: 22 Type locality: El Amiria, Alexandria Gov., Lower Egypt Holotype: MNHP 0.2481 Distribution: northern Egypt

Laudakia stoliczkana (Blanford, 1875)

PL Bogomolov Subspecies: Laudakia stoliczkana stoliczkana (Blanford, 1875) Stellio stoliczkanus Blanford, 1875: 191 Synonym Agama tarimensis Zugmayer, 1909: 493 Type locality: Hotan, Xinjiang Uygur AR, China Syntypes:, ZSM 314/1910/1 – 9. Type locality : plains of eastern Turkestan Syntypes: ZSI 3813–15, ZSI 4205, Distribution: Mongolia; China: Kashgar, Tien-Shan and southern Gobi

Laudakia stoliczkana altaica (Munkhbayar & Shagdarsuren, 1970) Agama himalayana altaicus Munkhbayar & Shagdarsuren, 1970: 86 Holotype: ZMB 42359 Type locality: granite mountains on the northern side of Uljastajn-gol Valley (46° 14’ N, 91° 35 E), Mongolian Altai, Mongolia Distribution: Mongolia: Gobi Desert

Laudakia tuberculata (Gray, 1827) Agama tuberculata Gray, 1827: 218

S Harikishnan Synonym Stellio indicus Blyth, 1853: 646 Type locality: upper Hindustan (northern India) Syntypes: ZSI 6826, 6831 Type locality: Bengal, India Syntypes: BMNH 1946.8.28.15-17, 1946.8.28.60 Distribution: eastern Afghanistan; northwestern Pakistan; India: Kashmir, Uttar Pradesh; southwestern Nepal.

Laudakia wui Zhao, 1998 Laudakia wui Zhao, 1998: 440 Type locality: Yi'ong, Bomi Co., Xizang Autonomous Region (=Tibet) Holotype: CIB 7315012 Distibution: China: Xizang = Tibet Last seen 1973

Leiolepis Cuvier, 1829 Cuvier, G.J.L.N.F.D. 1829. Le Regne Animal Distribué, d'apres son Organisation, pur servir de base à l'Histoire naturelle des Animaux et d'introduction à l'Anatomie Comparé . Nouvelle Edition [second edition]. Vol. 2. Les Reptiles. Déterville, Paris, i-xvi, 1-406 Type species: Leiolepis guttatus Cuvier, 1829, by original designation

Leiolepis belliana (Hardwicke & Gray, 1827)

C Smith Subspecies: Leiolepis belliana belliana (Hardwicke & Gray, 1827) Uromastyx belliana Hardwicke & Gray, 1827: 220 Synonyms Leiolepis bellii Gray, 1845: 263 Type locality: Penang, West Malaysia Holotype: BMNH XXIV.77.a Type locality: Penang, West Malaysia Type: Iconotype: Based on a drawing by Hardwicke Distribution: Thailand; Burma; Laos; Cambodia; Vietnam; Indonesia: Sumatra, Pulau Bangka; West Malaysia

Leiolepis belliana ocellata Peters, 1971 Leiolepis belliana ocellata Peters, 1971: 116 Type locality: Kokariet, Burma Holotype: ZMB 11604 Distribution: Burma; northern Thailand: Chinag Mai, Lampang, Nan, Phayao, Tak

Leiolepis boehmei Darevsky & Kupriyanova, 1993 Leiolepis boehmei Darevsky & Kupriyanova, 1993: 4 Type locality: Songkhla, southern Thailand Holotype: ZFMK 24651 Distribution: southern Thailand

Leiolepis guentherpetersi Darevsky & Kupriyanova, 1993 Leiolepis guentherpetersi Darevsky & Kupriyanova, 1993: 9

T Ngo Type locality: Thuy Phu, 17 km SE of Hue, Binhtrithien Province, central Vietnam Holotype: ZISP 20326 Distribution: central Vietnam: Binhtrithien & Quang Nam Provinces

Leiolepis guttata Cuvier, 1829 Leiolepis guttata Cuvier, 1829: 37

P Rojanadilok Synonyms Leiolepis belliana annamensis Smith, 1921: 429 Type locality: Cham Island, central Vietnam Syntypes: BMNH 1946.8.14.57-58, 1946.8.14.60-63 Type locality: Cochinchina, Vietnam Syntypes: MNHP 0. 2587-88 Distribution: coastal central Vietnam

Leiolepis ngovantrii Grismer & Grismer, 2010 Leiolepis ngovantrii Grismer & Grismer, 2010: 52 Type locality: Binh Chau – Phuoc Buu Nature Reserve (10°32.418N 107°32.177E), Xuyen Moc district, Ba Ria Vung Tau Province, Vietnam Holotype: LSUHC 9234 Distribution: Vietnam: Ba Ria Vung Tau Province

Leiolepis peguensis Peters, 1971 Leiolepis peguensis Peters, 1971: 119 Type locality: Pegu, Burma Holotype: NMHW 18890:3 Distribution: southern Burma

Leiolepis reveesii (Gray, 1831) Uromastyx reveesii Gray, 1831: 62 Type locality: China Holotype: BMNH 1946.8.14.59 Distribution: China: Macao, Guangdong, Hainan, Guangxi: Vietnam; Cambodia

Leiolepis rubritaeniata Mertens, 1961 Leiolepis rubritaeniata Mertens, 1961: 508 Type locality: Pakchong, 160 km northeast of Bangkok, on the road between Bangkok and Korat, Thailand. Holotype: SMF 57459 Distribution: eastern and central Thailand; Laos; Cambodia; west-central Vietnam .

Leiolepis triploida Peters, 1971 Leiolepis triploida Peters, 1971:123 Type ocality: Malaysia-Thailand borderlands of the Malaya Peninsula Holotype : MCZ 104381 Distribution: extreme southwestern Thailand; northwestern West Malaysia

Lophocalotes Günther, 1872 Günther, A. 1872. On the reptiles and amphibians of Borneo. Proceedings of the Zoological Society of London, 1872: 586-600. Type species: Lophocalotes interruptus Günther, 1872, by monotypy

Lophocalotes achlios Harvey, Scrivani, Shaney, Hamidy, Kurniawan, Smith, 2018 Lophocalotes achlios Harvey, Scrivani, Shaney, Hamidy, Kurniawan, Smith, 2018: 79 Type locality: along the main trail up Gunung Kaba (3.50757°S, 102.62927°E), Kabupaten Rejang Lebong, Provinsi Bengkulu, Sumatera, Indonesia Holotype: MZB 14038 Distribution: Indonesia: Sumatra Island: southern Bukit Barisan Range

Lophocalotes ludekingi (Bleeker, 1860) Calotes ludekingi Bleeker, 1860:326

H Kurniati Synonym: Lophocalotes interruptus Günther, 1872 Type locality: Agam, Sumatra, Indonesia Holotype: BMNH 1946.8.27.88 Distribution: Indonesia: west central Sumatra

Lophognathus Gray. 1842 Gray, J.E. 1842. Description of some hitherto unrecorded species of Australian reptiles and batrachians. pp. 51-57 in Gray, J.E. (ed.). The Zoological Miscellany . London : Treuttel, Würz & Co . Type species: Redtenbacheria fasciata Steindachner, 1867 by monotypy

Lophognathus gilberti Gray, 1842 Lophognathus gilberti Gray, 1842: 53

S Zozaya Synonyms Redtenbacheria fasciata Steindachner,1867:31 Type locality: Australia Holotype: not found Physignathus incognitus Ahl,1926:190 Type locality: Australia Holotype ZMB 30086 Type locality: Port Essington, Northern Territory Holotype: BMNH 1946.8.28.69 Distribution: Australia: North Territory, western-central Queensland, northern & northwestern Western Australia

Lophognathus horneri Melville, Ritchie, Chapple, Glor & Schulte, 2018 Lophognathus horneri Melville, Ritchie, Chapple, Glor & Schulte, 2018: 54 Type locality : Sambo Bore(18°52'48"S, 130°40'12"E), Wave Hill Station, Northern Territory. Holotype: NTM R16472 Distribution : Australia: central and western portions of Northern Australia .

Lyriocephalus Merrem, 1820 Merrem, B. 1820. Versuch eines Systems der Amphibien I (Tentamen Systematis Amphibiorum). J. C. Krieger, Marburg, 191 pp. Type species: Lyriocephalus margaritaceus Merrem, 1820, by monotypy

Lyriocephalus scutatus (Linnaeus, 1758) Lacerta scutata Linnaeus, 1758: 201

I Das Synonyms Iguana clamosa Laurenti, 1768:49 Type locality: not found Iconotype: Seba, l.c.s. Lyriocephalus margaritaceus Merrem, 1820: 49 Type locality: Ceylon Iconotype: Seba, l.c.s. Lyriocephalus macgregorii Gray, 1835: pl.lxviii, fig. 1 Type locality: not found Iconotype: collection of Maj. General Hardwicke Type locality: Asia (=Sri Lanka) Type: Iconotype: Seba Mus. vol. I. pl. 109. fig. 3. 71 Distribution: Sinharaja lowlands, Sri Lanka

Malayodracon Denzer, Manthey, Mahlow & Böhme, 2015. The systematic status of Gonocephalus robinsonii Boulenger, 1908 (: Agamidae: Draconinae) Zootaxa 4039 (1): 129–144. Type species: Gonyocephalus robinsonii Boulenger, 1908 Atheris

Malayodracon robinsonii (Boulenger, 1908 ) Gonyocephalus robinsonii Boulenger, 1908: 65 Type locality : summit of Gunung Tahan, Selangor, West Malaysia Holotype: BMNH 1946.8.14.81 Distribution: West Malaysia: Selangor, Pahang

Mantheyus Ananjeva and Stuart, 2001 Ananjeva, N.B. & B.L. Stuart. 2001. The agamid lizard Ptyctolaemus phuwuanensis Manthey & Nabhitabhata, 1991 from Thailand and Laos represents a new genus. Russian Journal of Herpetology . 8 (3): 165-170 Type species. Ptyctolaemus phuwuanensis Manthey and Nabhitabhata, 1991, by original designation

Mantheyus phuwuanensis (Manthey & Nabhitabhata, 1991) Ptyctolaemus phuwuanensis Manthey & Nabhitabhata, 1991: 3

Thomas Calame Type locality: Phu Wua Wildlife Sanctuary (18°15' N, 103°58' E) Nong Dem Subdistrict , Bung Khan District, Nong Khai Province, Thailand Holotype: ZMB 49040 Distribution: northeastern Thailand; west-central Laos

Microauris Pal, Vijayakumar, Shanker, Jayarajan & Deepak, 2018. Saunak Pal, S.P. Vijayakumar, Kartik Shanker , Aditi Jayarajan and V. Deepak. 2018. A Systematic Revision of Calotes Cuvier, 1817 (Squamata: Agamidae) from the Western Ghats adds Two Genera and Reveals Two New Species. Zootaxa . 4482(3); 401–450. Type species: Calotes aurantolabium Krishnan, 2008

Microauris aurantolabium (Krishnan, 2008) Calotes aurantolabium Krishnan, 2008: 531 Type locality : Kalakad Mundanthurai Tiger Reserve, Tamil Nadu, India Holotype: BNHM 1436 Distribution: India: Western Ghats

Microauris aurantolabium Manjo P

Moloch Gray, 1841 Gray, J.E. 1841. Description of some new species and four new genera of reptiles from Western Australia, discovered by John Gould, Esq. Annals and Magazine of Natural History 7 42 : 86-91 Type species: Moloch horridus Gray, 1841, by monotypy

Moloch horridus Gray, 1841 Moloch horridus Gray, 1841: 89

S Zozaya Synonym Acanthosaura gibbosus Berthold, 1846: 12 Type locality: Western Australia Holotype: ZUUG, not located Type locality: Western Australia Syntype: BMNH 1860.10.25.144.a Distribution: central Australia: Northern Territory, Queensland, South Australia, Western Australia

Monilesaurus Pal, Vijayakumar, Shanker, Jayarajan & Deepak, 2018 Saunak Pal, S.P. Vijayakumar, Kartik Shanker , Aditi Jayarajan and V. Deepak. 2018. A Systematic Revision of Calotes Cuvier, 1817 (Squamata: Agamidae) from the Western Ghats adds Two Genera and Reveals Two New Species. Zootaxa . 4482(3); 401–450. Type species: Calotes rouxii (Duméril & Bibron, 1837)

Monilesaurus rouxii M O’Shea

Monilesaurus acanthocephalus Pal, Vijayakumar, Shanker, Jayarajan & Deepak, 2018

Monilesaurus acanthocephalus Pal, Vijayakumar, Shanker, Jayarajan & Deepak, 2018: 432 Type locality : evergreen forest—tea garden edge in Upper Manalar (934'35.81"°N, 720'11.43"E°), Periyar tiger reserve, Megamalai Holotype : BNHS 2409 Distribution: India: Megamalai hills of southern Western Ghats

Monilesaurus acanthocephalus Pal

Monilesaurus ellioti (Günther, 1864) Subspecies: Monilesaurus ellioti ellioti (Günther, 1864) Calotes ellioti Günther,1864:142 Synonym Bronchocela indica Theobald, 1876: 105 Type locality: southern India Syntypes: ZSI 4325, 4328 Type locality : Chembra reserve forest, Kerala, India Neotype: CESL 045 Distribution: India: Western Ghats Monilesaurus ellioti Harikishnan

Monilesaurus ellioti amarambalamensis (Murthy, 1978) Calotes ellioti amarambalamensis Murthy, 1978: 79 Type locality : Mennmutty, New Amarambalam Reserve Forest, Kerala, India Holotype : ZSI-Madras 159 Distribution : southern India

Monilesaurus montanus Pal, Vijayakumar, Shanker, Jayarajan & Deepak, 2018

Monilesaurus montanus Pal, Vijayakumar, Shanker, Jayarajan & Deepak, 2018: 436 Type locality : Kudremukh National Park (1307'54"°N, 07516'39"°E, Karnataka Holotype: BNHS 2411 Distribution : India: Kudremukh, Brahmagiri, Nilgiri and Elivalmalai Ranges

Monilesaurus montanus Pal

Monilesaurus rouxii (Duméril & Bibron, 1837) Calotes rouxii Duméril & Bibron, 1837: 407 Type locality : India Syntypes: MNHP 0.6894, BMNH 1994.1857 Distribution: India: Gujarat, Goa, Maharashtra, Karnataka

Otocryptis Wagler, 1830 . Wagler, Jean G. 1830. Natürliches System der Amphibien, mit vorangehender Classification der Säugetiere und Vögel. Ein Beitrag zur vergleichenden Zoologie . 1.0. Cotta, München, Stuttgart, and Tübingen, 354 pp. Type species: Otocryptis wiegmanni Wagler, 1830, by monotypy

Otocryptis beddomei Boulenger, 1885 Otocryptis beddomei Boulenger, 1885: 272

D Momaya Type locality: Sivagiri Ghat, southern India Syntypes: BMNH 1882.5.22.101-104, ZSI 15733 Distribution: southern India: Kerala, Tamil Nadu

Otocryptis nigristigma Bahir & Silva, 2005 Otocryptis nigristigma Bahir & Silva, 2005: 394 G Jayaweera Type locality: Ritigala (08°07’N, 80°40’E), Sri Lanka Holotype: WHT 6176 Distribution: Sri Lanka

Otocryptis wiegmanni Wagler, 1830 Otocryptis wiegmanni Wagler, 1830: 150

I Das Synonym Otocryptis bivittata Wiegmann, 1831: 291 Type locality: Ceylon Holotype: ZMB 708 Type locality: “America”, in error for Sri Lanka Holotype: ZMB 708 Distribution: Sri Lanka

Phoxophrys Hubrecht, 1881 A. A. W. Hubrecht. 1881 . Phoxophrys . Notes from the Royal Zoological Museum of the Netherlands, Leyden 3: 51

Type species: Phoxophrys tuberculata Hubrecht, 1881, by monotypy and designation

Phoxophrys borneensis Inger, 1960 Phoxophrys borneensis Inger, 1960: 222

I Das Type locality: Bundu Tuhan, Mount Kina Balu, Borneo, Sabah State, East Malaysia Holotype: USNM 130228 Distribution: Indonesia: Kalimantan (north Borneo); East Malaysia: Sabah, Sarawak

Phoxophrys cephalum (Mocquard, 1890) Pelturagonia cephalum Mocquard, 1890:130

T Charlton Type locality: Mount Kinabalu (06° 00-15’N, 116° 15-45’E), Sabah, East Malaysia Syntypes: MNHN 1889.166-172; BMNH 1946.8.14.8 Distribution: Indonesia: Kalimantan; East Malaysia: Sabah, Sarawak

Phoxophrys nigrilabris (Peters, 1864) Otocryptis nigrilabris Peters, 1864: 385 I Das Type locality: Pulau Matjan, Kalimantan, Indonesia Holotype: ZMB 5023 Distribution: Indonesia: Kalimantan (Borneo); East Malaysia: Sarawak

Phoxophrys spiniceps Smith, 1925 Phoxophrys spiniceps Smith, 1925: 7

I Das Type locality: Mt. Murud (03° 55’N, 115° 32’E), Borneo, Sarawak, East Malaysia Holotype: BMNH 1946.8.27.45 Distribution: Indonesia: Kalimantan; East Malaysia: Sabah, Sarawak

Phoxophrys tuberculata Hubrecht, 1881 Phoxophrys tuberculata Hubrecht, 1881: 51 Synonym Japalura robinsoni Boulenger, 1920: 287 Type locality: Sungei Kumbang, Korintji Peak, Sumatra, Indonesia Holotype: ZRC, not located Type locality: Batang, Singalang, western Sumatra, Indonesia Holotype: BMNH 1946.8.13.92 Distribution: Indonesia: western Sumatra

Phrynocephalus Kaup, 1825 Kaup, J.J. 1825. Einige Bemerkungen zu Merrems Handbuch. Isis von Oken 16: 589-593. Type species: Lacerta caudivolvula Pallas, 1814, by subsequent designation

Phrynocephalus ahvazicus Melnikov, Melnikova, Nazarov, Rajabizadeh, Al-Johany & Amr, 2014 Phrynocephalus ahvazicus Melnikov, Melnikova, Nazarov, Rajabizadeh, Al-Johany & Amr, 2014:155

D Melnikov Type locality Ahvaz, Khuzestan province, Iran Holotype: ZISP 27131 Distribution: Ahvaz plains, Khuzestan province, southwestern Iran.

Phrynocephalus albolineatus Zhao, 1979 Phrynocephalus albolineatus Zhao, 1979: 113 Type locality: 45 km south of Techeng Co., Xinjiang Uygur Autonomous Region, China Holotype: NMU 77181 Distribution: China: northern Xinjiang

Phrynocephalus ananjevae Melnikov, Melnikova, Nazarov & Rajabizadeh, 2013 Phrynocephalus ananjevae Melnikov, Melnikova, Nazarov & Rajabizadeh, 2013: 40 Type locality: Qahferokh (ca. 32°16’ N, 50°58’ E), vicinity of Farokhshahr, Zagros mountains, southern Iran Holotype: ZISP 10256.1 Distribution: southern Iran: Zagros mountains

Phrynocephalus arabicus Anderson, 1894 Phrynocephalus arabicus Anderson, 1894: 377

M O’Shea Type locality: plateau of Hadramut, southeastern Arabia Syntypes: BMNH 1946.8.28.33-34 Distribution: eastern Saudi Arabia; United Arab Emirates; Oman; southern Iraq; Jordan

Phrynocephalus arcellazzii Bedriaga, 1909 Phrynocephalus arcellazzii Bedriaga, 1909: 309 Type locality: Qitai (44°00’N, 89°50’E), Junggar basin, Xinjiang Uygur Autonomous Region, China Holotype: ZISP 8166 Distribution: China: Xinjiang Uygur Autonomous Region

Phrynocephalus axillaris Blanford, 1875 Phrynocephalus axillaris Blanford, 1875:192

Y Qi Synonyms Phrynocephalus axillaris idae Bedriaga, 1907: 159 Type locality: Nan Shan, Xinjiang Uygur Autonomous Region, China Lectotype : ZISP 10062.1, designated by Barabanov and Ananjeva, 2007 Phrynocephalus axillaris murielis Bedriaga, 1907: 170 Type locality:. Lukqun (ca. 42°43’N, 89°41’E), Xinjiang Uygur Autonomous Region, China Lectotype: ZISP 8164, designated by Barabanov and Ananjeva, 2007 Phrynocephalus axillaris ericae Bedriaga, 1907: 168 Type locality: Lukqun (ca. 42°43’ N, 89°41’ E), Xinjiang Uygur Autonomous Region, China Lectotype : ZISP 10060.1, designated by Barabanov and Ananjeva, 2007 Phrynocephalus kozlovi Bedriaga, 1906: 162 Type locality : Lukqun (ca. 42°43’N, 89°41’E), Xinjiang Uygur Autonomous Region, China. Holotype . ZISP 10059 Phrynocephalus axillaris klemenzi Bedriaga, 1907: 163 Type locality:. Turpan county, Xinjiang Uygur Autonomous Region, China Lectotype: ZISP 10050.1, designated by Barabanov and Ananjeva, 2007 Phrynocephalus axillaris maculata Bedriaga, 1907: Table III, Fig. 3 Type locality: probably Qiemo Xian, Qarqan Xian (ca. 38°08’ N, 85°32’E), Xinjiang Uygur Autonomous Region, China Holotype: ZISP 10663 Phrynocephalus ludovici Moquard, 1910: 146 Type locality: black desert near Oasis of Kouchar, eastern Shantung, Xinjiang Uygur Autonomous Region, China Lectotype: MNHP 1909.39, designated by Peters, 1984 Phrynocephalus nasatus Golubev & Dunayev, 1995: 5 Type locality: Topa-Bell, Karaduvan Ridge (41°11’N, 80°48’E), and the town of Aksu (41°11’N, 80°14 E), Kashgaria, western China Holotype, ZMMU Re-7614. Type locality: plains of eastern Turkestan Syntypes: ZSI 3612-18, 3620, 3622-23, 3625-38, 3832-34, 4839-40 Distribution: eastern Turkestan; Mongolia; China: Tibet, Xinjiang, Gansu

Phrynocephalus bannikovi Darevsky, Rustamov & Shammakov, 1976 Phrynocephalus reticulatus bannikovi Darevsky, Rustamov & Shammakov, 1976: 114 Type locality: Tuarkyr mountains, Balkan province, Turkmenistan Holotype: ZISP 18505 Distribution: northwestern Turkmenistan: Krasnovodsky Peninsula

Phrynocephalus bedriagai Tzarevsky, 1926 Phrynocephalus bedriagai Tzarevsky, 1926: 214 Type locality: Zaissan Lake region, eastern Kazakhstan Lectotype: ZISP 10606.1, designated by Peters, 1984 Distribution: Kazakhstan

Phrynocephalus clarkorum Anderson & Leviton, 1967 Phrynocephalus clarkorum Anderson & Leviton, 1967: 228 Type locality: 20 miles southeast of Kandahar (31º20' N, 65º50' E), Afghanistan Holotype: CAS 97989 Distribution: Afghanistan; Pakistan

Phrynocephalus erythrurus Zugmayer, 1909 Phrynocephalus erythrurus Zugmayer, 1909: 504 Type locality: Sagüs Kul (a lake), northwestern Tibet Lectotype: ZSM 317/1910, designated by Peters 1984 Distribution: China: northwestern Tibet: Keria mountains

Phrynocephalus euptilopus Alcock & Finn, 1897 Phrynocephalus euptilopus Alcock & Finn, 1897: 556 Type locality: Darband, northern Baluchistan, Pakistan Syntypes: BMNH 1946.8.28.39; ZSI 14070, 14072; MCZ R-7227 Distribution: southwestern Pakistan: western Baluchistan; southwestern Afghanistan

Phrynocephalus forsythii Anderson, 1872 Phrynocephalus forsythii Anderson, 1872: 390

D Melinkov Type locality: Shache (38º25' N, 77º14' E), southern Xinjiang Uygur Autonomous Region, western China Syntypes: BMNH 1946.8.28.40; NMHW 24809; ZSI 5167, 5168, 5181 Distribution: China: Xinjiang

Phrynocephalus frontalis Strauch, 1876 Phrynocephalus frontalis Strauch, 1876: 15 Type locality: Ordos Desert, Inner Mongolia AR, China Lectotype: ZISP 3920, designated by Peters, 1984 Distribution: China: (Gansu, Ningxia, Shaanxi, Nei Mongol, Hebei

Phrynocephalus golubewii Shenbrot & Semyonov, 1990 Phrynocephalus golubewii Shenbrot & Semyonov, 1990:155 Type locality: vicinity of the Bami rail-road station, Bami, Turkmenistan Holotype: ZMMU Re-6411 Distribution: Turkmenistan

Phrynocephalus grumgrzimailoi Bedriaga, 1907 Phrynocephalus Grum-Grzimailoi Bedriaga, 1907: 145 Type locality: “Gutschen” [= Qitai County, Xinjiang Uygur Autonomous Region, China]. Lectotype: ZISP 8178, designated by Barabanov and Ananjeva, 2007 Distribution: China: northern Xinjiang Province

Phrynocephalus guinanensis Xiang, Wang & Wang, 2009 Phrynocephalus guinanensis Xiang, Wang & Wang, 2009: 62 Type locality: Senduo (35°34’N, 101°05’E0, Guinan County, Qinghai Province, China. Holotype: NNU P 2004.006 Distribution: China: Qinghai Province

Phrynocephalus guttatus (Gmelin, 1789)

S Sklyarenko Subspecies: Phrynocephalus guttatus guttatus (Gmelin, 1789) Lacerta guttata Gmelin in Linnaeus, 1789: 1078 Synonyms Lacerta caudivolvula Pallas, 1814: 27 Type locality: vicinity of Atyrau (Gurjev) (ca. 47°06’N, 51°54’E), Atyrau province, northwestern Kazakhstan Neotype: ZISP 4910.1, designated by Barabanov and Ananjeva, 2007 Phrynocephalus suschkini Bedriaga in Nikolsky, 1905: 472 Type locality: vicinity of the Chalkar Teniz lake, Turgai depression, Kazakhstan Holotype: ZISP 9180 Phrynocephalus caudivolvulus moltschanowi Nikolsky, 1913: 34 Type locality: Beltau mt., Karakalpakstan, Uzbekistan Lectotype: ZISP 19585, designated by Semenov and Shenbrot, 1982 Phrynocephalus guttatus kalmykus Badmayeva & Szczerbak, 1983: 34 Type locality: Khulkhuta village, approximately 46º19' N, 46º22' E, Chernozemelskiy District, Kalmykia. Russia Holotype: IZANU Re 15. Type locality: vicinity of Atyrau (Gurjev), approximately 47° 06' N, 51° 54' E, Atyrau Province, north- western Kazakhstan Neotype: ZISP 4910.1, designated by Barabanov & Ananjeva 2007 Distribution: Russia: from northwest coast of Caspian Sea eastward to eastern Kazakhstan; Turkmenistan

Phrynocephalus guttatus alpherakii Bedriaga, 1906 Phrynocephalus alpherakii Bedriaga, 1907: 11 Synonyms: Phrynocephalus acutirostris Bedriaga, 1907: 339 Type locality: Yining Shi (ca. 43°54’N, 81°18’E), western Xinjiang Uygur Autonomous Region, China. Lectotype: ZISP 5548.1, designated by Peters, 1984 Phrynocephalus versicolor paraskiwi Semenov, Brushko, Shenbrot and Kubykin, 1987: 106. Type locality: Ili river hollow, urochishe Ajakkalkan, Taldy-Kurgan Region, Kazakhstan Holotype: IZK 178. Lectotype ZISP 10038.3, designated by Peters, 1984 Type locality: Korgas river, tributary of the upper Ily river, western Xinjiang Uygur Autonomous Region, China Distribution: Kazakhstan; China: Xinjiang, Inner Mongolia

Phrynocephalus guttatus melanurus Eichwald, 1831 Phrynocephalus melanurus Eichwald, 1831:186 Synonyms: Phrynocephalus caudivolovulus incerta Bedriaga in Nikolsky, 1905: 475 Type locality: vicinity of Almaty (ca. 43°17’N 76°55’E), Kazakhstan Lectotype: ZISP 6751.1, designated by Peters, 1984 Phrynocephalus kuschakewitschi Bedriaga in Nikolsky, 1905: 476 Type locality: Ayaguz (ca.47°55’N, 80°23’E), eastern Kazakhstan Lectotype: ZISP 6839.1, designated by Peters, 1984 Phrynocephalus bergi Elpatjevsky, 1906 "1907": 51 Type locality: vicinity of Malaykuduk village, Almaty region, Kazakhstan Lectotype: ZISP 10384.1, designated by Barabanov and Ananjeva, 2007 Phrynocephalus caudivolovulus ocellata Bedriaga, 1909: 368 Type locality: Lepsinskaja Staniza, shore of Alakol Lake, Kazakhstan Syntypes: ZISP, not located Type locality: Kyzylkum sands near the Kurchum village left side of the Buhtarminskoe reservoir, eastern Kazakhstan Neotype: ZMMU Re-5611, designated by Semenov et al., 1987 Distribution: Russia: southern Siberia; eastern Kazakhstan; China: Xinjiang

Phrynocephalus guttatus salsatus Golubev, Gorelov, Dunayev & Kotenko, 1995 Phrynocephalus guttatus salsatus Golubev, Gorelov, Dunayev & Kotenko, 1995: 34 Type locality: north Turkmeniya, salt hollow Kazakhlyshor between sands Kumsebshen and the sink of Ustjurt (40º17' N, 55º46' E), Turkmenistan Holotype: IZANU Re-21/1 Distribution: Turkmenistan

Phrynocephalus haeckeli Bedriaga, 1907 Phrynocephalus haeckeli Bedriaga, 1907: 237 Type locality: in Saur mountains of the Tarbagatay range, Tacheng district, northern Xinjiang Uygur Autonomous Region, China Holotype: ZISP 5163 Distribution: China: northern Xinjiang Uygur Autonomous Region

Phrynocephalus helioscopus (Pallas, 1771) Subspecies: Phrynocephalus helioscopus helioscopus (Pallas, 1771) Lacerta helioscopa Pallas, 1771: 457

R Knight

Synonym Lacerta uralensis Gmelin, 1789: 1073 Type locality: Ural desert Lectotype: Iconotype: Lepechin (1774) pl. 22, Fig. 1, designated by Barabanov-Ananjeva 2007 Type locality: Inderskija steppe, area of the lower Ural River, North Kazakhstan Lectotype: ZMB 781, designated by Denzer et al., 1997. Distribution: Armenia eastward to E Kazakhstan southern Russia; northern Iran; Iraq; Turkey

Phrynocephalus helioscopus saidalievi Sattorov, 1981 Phrynocephalus helioscopus saidalievi Sattorov, 1981: 7 Type locality: Konibodom village, approximately 40º17' N, 70º25' E, Tajikistan Holotype: IZANU Re 12. Distribution: Uzbekistan; Tajikistan; Kyrgyzstan

Phrynocephalus helioscopus varius Eichwald, 1831 Phrynocephalus varius Eichwald, 1831:186 Synonyms: Phrynocephalus nigricans Eichwald, 1831: 186. Type locality: southern Siberia Lectotype; ZISP 8719.1, designated by Barabanov-Ananjeva 2007 Phrynocephalus helioscopus cameranoi Bedriaga, 1907: 243 Type locality: Shi (ca. 43°54’N, 81°18’E), western Xinjiang Uygur Autonomous Region, China Lectotype: ZISP 10036.1 , designated by Barabanov and Ananjeva, 2007 Phrynocephalus helioscopus saposhnikovi Kashtchenko, 1909: 122 Type locality: vicinity of the Karatal village, approximately 47º36' N, 85º10' E, eastern Kazakhstan Type: Tomsk State University Zoological Museum (ZMTU), not located Phrynocephalus helioscopus orientalis Bedriaga,1912:751 Type locality: Ulungur river, Xinjiang Uygur Autonomous region, China Lectotype: ZISP 6553.1, designated by Barabanov and Ananjeva, 2007 Type locality: southern Siberia, near the Atlai Mountains, eastern Kazakhstan Holotype: ZISP 8718 Distribution: from Eastern Kazakhstan through Xinjiang Uygur Autonomous Region, China to western Mongolia

Phrynocephalus hongyuanensis Zhao, Jiang & Huang,1980 Phrynocephalus vlangalii hongyuanensis Zhao, Jiang & Huang,1980: 178 Type locality: Waqên, Hongyuan County (33º03’N; 102º37’E), Sichuan Province, China Holotype: CIB 785795 Distribution: China: northwestern portion of the Tibetan Plateau

Phrynocephalus horváthi Méhely, 1894 Phrynocephalus horváthi Méhely, 1894: 79

D Melinkov Type locality: Aralik village (39º52'N, 44º31'E), Idgir Province, base of Mount Ararat, Turkey Neotype: ZISP 5544.1, designated by Barabanov & Ananjeva 2007 Distribution: northeastern Turkey; Armenia

Phrynocephalus interscapularis Lichtenstein, 1856

D Melnikov Subspecies: Phrynocephalus interscapularis interscapularis Lichtenstein, 1856 Phrynocephalus interscapularis Lichtenstein, 1856:12 Type locality: Bukhara, Uzbekistan Lectotype: ZMB 796, designated by Barabanov and Ananjeva, 2007 Distribution: Turkmenistan; southern Kazakhstan; Uzbekistan; northeastern Iran; northeastern Afghanistan

Phrynocephalus interscapularis sogdianus Chernov, 1948 Phrynocephalus interscapularis sogdianus Chernov, 1948: 135 Type locality: Pyandza (Pjantsch), Tadjikistan Holotype: ZISP 16887 Distribution: southern Uzbekistan; southwestern Tajikistan; northern Afghanistan

Phrynocephalus isseli von Bedriaga, 1907 Phrynocephalus isseli von Bedriaga, 1907: 229 Type locality: in Saur mountains of the Tarbagatay range, Tacheng district, northern Xinjiang Uygur Auto- nomous Region, China Holotype: ZISP 5164 Distribution: China: Xinjiang Uygur Autonomous Region

Phrynocephalus longicaudatus Haas, 1957 Phrynocephalus maculatus longicaudatus Haas, 1957: 67 Type locality : Doha Dhalum, Saudi Arabia Holotype : CAS 84449 Distribution: Saudi Arabia; Oman; United Arab Emirates; Kuwait

Phrynocephalus lutensis Kamali and Anderson, 2015 Phrynocephalus lutensis Kamali and Anderson, 2015: 250 Type locality: Mega Dune (30° 27’ N, 59° 21’ E), Lut Desert, Kerman Province, near the junction of Kerman, South Khorasan and Sistan & Baluchistan Provinces. Holotype: ZISP 28014 Distribution: Lut Desert, Iran

Phrynocephalus luteoguttatus Boulenger, 1887 Phrynocephalus luteoguttatus Boulenger, 1887: 497 Type locality: between Nushki and Helmand, Afghanistan Syntypes: BMNH 1946.8.28.44-49, BMNH 1946.8.28.36-38; ZISP 7363 Distribution: southeastern Iran; southwestern Afghanistan; southwestern Pakistan:Baluchistan

Phrynocephalus macropeltis Haas, 1957 Phrynocephalus nejdensis macropeltis Haas, 1957: 69 Type locality: Dhahran [Az Zahrãn], approximately 26º17' N, 50º12' E, Saudi Arabia Holotype: CAS 84619 Distribution: Saudi Arabia

Phrynocephalus maculatus Anderson, 1872

K Kamai Phrynocephalus maculatus Anderson, 1872: 389 Synonym Phrynocephalus spiniventris Nikolsky, 1896: 370 Type locality: Sistan and Baluchestan province, Iran Holotype: ZISP 8780 Type locality: Abadeh (31º10' N, 52º37' E), Fars Province, Iran Holotype: ZSI 4825 Distribution: Iran; Turkmenistan; Syria; Jordan; Iraq; Pakistan; Afghanistan

Phrynocephalus mystaceus (Pallas, 1776)

D Melnikov Subspecies: Phrynocephalus mystaceus mystaceus (Pallas, 1776) Lacerta mystacea Pallas, 1776: 702 Synonym Phrynocephalus mystaceus dagestanicus Krassowsky, 1932: 228 Type locality: Kumtorkala (ca. 42°59’N, 47°15’E), Daghestan Autonomous Republic, Russia. Neotype: ZISP 15252.1, designated by Barabanov and Ananjeva, 2007 Type locality: Ryn Peski (Ryn Sands), Uralskaya Region, northwestern Kazakhstan Neotype: ZISP 8735.1, designated by Barabanov & Anajeva 2007 Distribution: Russia: coast of Caspian Sea; western Kazakhstan; northeastern Iran; western Turkmenistan

Phrynocephalus mystaceus galli Krassowsky, 1932 Phrynocephalus mystaceus galli Krassowsky, 1932: 227 Synonym: Phrynocephalus mystaceus aurantiacaudatus Semenov & Shenbrot, 1990: 79 Type locality: 70 km NNW track Ushtobe (45º50' N; 77º40' E), Semirechije , eastern Kazakhstan Lectotype ZMMU Re-6413, designated by Semenov & Shenbrot, 1990. Type locality: vicinity of Repetek station, approximately 38º35' N, 63º11' E, Lebapsky Region, Turkmenistan Distribution: eastern Turkmenistan; Uzbekistan; eastern Kazakhstan; Afghanistan; China: Xinjiang

Phrynocephalus mystaceus khorasanus Solovyeva, Dunayev, Nazarov, Radjabizadeh, Poyarkov. 2018. Phrynocephalus mystaceus khorasanus Solovyeva, Dunayev, Nazarov, Radjabizadeh, Poyarkov. 2018:114 Type locality . Iran, Khorasan historical area, sand dunes (34°39'N, 58°43'E), Khorasan Razavi Province ), environs of Gonabad, the right bank of the Kale-Shur River Holotype . ZMMU R-11913 Distribution: Iran: Khorasan Razavi and eastern Semnan Provinces

Phrynocephalus nejdensis Haas, 1957 Phrynocephalus nejdensis Haas, 1957: 68 Type locality: Nafud, near Qana, north Nedjd, wnw of Hail, Saudi Arabia Holotype: HUJ 2711 Distribution: eastern Saudi Arabia; Oman; northeastern Yemen

Phrynocephalus ornatus Boulenger, 1887 Phrynocephalus ornatus Boulenger, 1887: 496 Type locality: between Nushki and Heland River, Baluchistan, Afghanistan Lectotype : BMNH 1946.8.28.20, designated by Anderson & Leviton, 1967 Distribution: southwestern Pakistan: Baluchistan; southwstern Afghanistan; eastern Iran

Phrynocephalus persicus De Filippi, 1863 Phrynocephalus persicus De Filippi, 1863: 387 Type locality: Desert fields in Iran by the road from Sultaniyeh to Teheran Lectotype : ZISP 8844, designated by Melnikov et al. 2013 Distribution: northern Iran; eastern Turkey; Azerbaijan, southern Armenia; southern Turkmenistan

Phrynocephalus przewalskii Strauch, 1876 Phrynocephalus przewalskii Strauch, 1876: 10 Synonyms Phrynocephalus affinis Strauch, 1876: 10 Type locality: Ordos desert, Inner Mongolia, China Lectotype: ZISP 3922, designated by Peters, 1984 Phrynocephalus steindachneri Bedriaga, 1907: 167 Type locality: Ruo river, northern limit of Nanshan mountains near Gaotai (ca. 39°22’N, 99°48’E), Gansu province, China Holotype : ZISP 8157 Phrynocephalus carinilabris Bedriaga, 1909: 412 Type locality: Zhugqu (ca.33°48’N, 104°30’E), southeast Gansu province, China Holotype: ZISP 10025 Phrynocephalus elegans Zarevsky, 1927: 305 pampa_isla Type locality: Tengger desert, from Dingyuanying (Inner Mongolia) to Sharuzan-Sume (Ningxia Hui), China Lectotype: ZISP 12193.1, designated by Peters, 1984 Phrynocephalus birulai Zarevsky, 1927: 304 Type locality: Tengger desert, Inner Mongolia, China Lectotype: ZISP 12183.1, designated by Peters, 1984 Phrynocephalus carinatus Zarevsky, 1927: 305 Type locality: Huang Shui, tributary of upper Huang He, eastern Nanshan, border area of Gansu and Qinghai provinces, China Lectotype: ZISP 12187.1, designated by Peters, 1984. Phrynocephalus parvulus Zarevsky, 1927: 305 Type locality: Tengger desert, from Dingyuanying (Inner Mongolia) to Sharuzan-Sume (Ningxia Hui), China Lectotype: ZISP 12189.1, designated by Peters, 1984 Phrynocephalus suschkinianus Zarevsky, 1927: 304 Type locality: vicinity of the Chalkar Teniz lake, Turgai depression, Kazakhstan Lectotype: ZISP 12192.1, designated by Barabanov and Ananjeva, 2007 Type locality: Tengger desert, Inner Mongolia, China Holotype: ZISP 3928 Distribution: China: Nei Mongol (= Inner Mongolia), Ningxia, Gansu, Qinghai; eastern Kazakhstan

Phrynocephalus raddei Boettger, 1888 Subspecies: Phrynocephalus raddei raddei Boettger, 1888 Phrynocephalus raddei Boettger, 1888: 262

D Melnikov Synonyms Phrynocephalus raddei bilkewitschi Nikolsky, 1915: 194 Type locality: Turkmenistan. Type: not located Phrynocephalus raddei zardunyi Nikolsky, 1915: 194 Type locality: Kelif (ca. 37°20’N, 66°18’E), Lebap province, Turkmenistan Lectotype: ZISP 10722.1, designated by Barabanov and Ananjeva, 2007 Type locality: Perevalnaja railroad station, southwestern Turkmenistan Lectotype : SMF 10247, designated by Mertens, 1967 Distribution: southern Turkmenistan; northern Iran; northern Afghanistan

Phrynocephalus raddei boettgeri Bedriaga, 1906 Phrynocephalus raddei boettgeri Bedriaga, 1906: 468 Lectotype: ZISP 6117.1, designated by Barabanov and Ananjeva, 2007 Type locality: Sherabad, approx. 37°42’ N, 67°04’E, Surkhandarya province, Uzbekistan Distribution: southern Uzbekistan; southwestern Tadjikistan

Phrynocephalus reticulatus Eichwald, 1831 Subspecies: Phrynocephalus reticulatus reticulatus Eichwald, 1831 Phrynocephalus reticulatus Eichwald, 1831:186 Synoynms Agama ocellata Lichtenstein, 1823: 143 Type locality: Bukhoro, Uzbekistan Lectotype: ZMB 787, designated by Golubev 1991 Phrynocephalus nikolskii Bedriaga in Nikolsky, 1905: 463 Type locality: Kyzylkum desert, Uzbekistan Lectotype: ZISP 5231.1, designated by Barabanov and Ananjeva, 2007 Phrynocephalus boulengeri Bedriaga in Nikolsky, 1905: 466 Type locality: Karshi desert, SE from Bukhara, southern Uzbekistan Lectotype : ZISP 7330.1, designated by Barabanov and Ananjeva 2007 Type locality: banks of the Oxus (=Amu Darya) Holotype: ZISP 8720 Distribution: northern Afghanistan; southern Kazakhstan; western Uzbekistan; southern Turkmenista; northern India: Kashmir

Phrynocephalus roborowskii Bedriaga, 1906 Phrynocephalus vlangalii roborowskii Bedriaga, 1906:172 Type locality: Qaidam basin, Qinghai province, China Lectotype: ZISP 7060, designated by Peters, 1984 Distribution: northwestern China: Qinghai, Xinjiang

Phrynocephalus rossikowi Nikolsky, 1898 Subspecies: Phrynocephalus rossikowi rossikowi Nikolsky, 1898 Phrynocephalus rossikowi Nikolsky, 1898: 286 Synonym Phrynocephalus rossikowi relictus Nikolsky, 1915: 169 Type locality: Kushkanatau Mountain in the Amu-Darja river mouth, Uzbekistan Holotype: ZDKU, not found Type locality: mountains near Nukus, Karakalpakstan, Uzbekistan Holotype: ZISP 9112. Distribution: Turkmenistan; Uzbekistan

Phrynocephalus rossikowi shammakowi Szczerbak & Golubev, 1979 Phrynocephalus rossikowi shammakowi Szczerbak & Golubev, 1979: 81 Holotype: IZANU Re 11 Type locality: central Kara-Kum Desert, Turkmenistan Distribution: southeastern Turkmenistan

Phrynocephalus sakoi Melnikov, Nazarov, Al-Johany & Ananjeva , 2015 Phrynocephalus sakoi Melnikov, Nazarov, Al-Johany & Ananjeva, 2015: 304

D Melnikov Type locality: Al Sharqiyah Sands, Filim vicinity (20°41’19.7” N, 58°17’41.6” E), northeastern Oman Holotype: ZISP 28705 Distribution: northeastern Oman: Al Sharqiyah Sands

Phrynocephalus salenskyi Bedriaga, 1907 Phrynocephalus salenskyi Bedriaga, 1907: 141 Type locality : Ulungur river, Xinjiang Uygur Autonomous Region, China Lectotype: ZISP 7035.1, designated by Peters, 1984: 141 Distribution: Junggur Depression of northwestern China, and the Zaysan Depression of northeastern Kazakhstan.

Phrynocephalus scutellatus (Olivier, 1807) Agama scutellata Olivier, 1807:110

V Zahiri Synonyms Phrynocephalus tickelii Gray, 1845: 260 Type locality: Afghanistan Holotype: BMNH XXIV.70a. Phrynocephalus olivieri brevipes Nikolsky, 1907: 274 Type locality: Naim-Abad (36°14’N, 54°39’E) , Mazandaran province, Iran Lectotype: ZISP 10235, designated by Barabanov and Ananjeva, 2007 Phrynocephalus olivieri carinipes Nikolsky, 1907: 273 Type locality: Tudeshg, Isfahan province, Iran Lectotype: ZISP 10236, designated by Barabanov and Ananjeva, 2007 Type locality: Mt. Sophia, near Esfahan, Esfahan Province, Iran Holotype: MNHP 0.6947 Distribution: Iran; Afghanistan; southwestern Pakistan: Baluchistan

Phrynocephalus strauchi Nikolsky, 1899 Phrynocephalus strauchi Nikolsky, 1899: 21 Type locality: Khodzhent (40º17' N, 69º37' E), northern Tajikistan Lectotype : ZMMU Re-2116, designated by Dunayev, 1995 Distribution: Tajikistan; Uzbekistan; Kyrgyzstan

Phrynocephalus theobaldi Blyth, 1863 Phrynocephalus theobaldi Blyth, 1863: 90

Y Qi Synonyms Phrynocephalus stolickai Steindachner, 1867: 23 Type locality: Rong village, Indus valley, Jammu and Kashmir, India. Syntypes: ZISP 5022.1 – 5022.3, NMHW 23482 Phrynocephalus alticola Peters, 1984: 58 Type locality: Rupshu in Ladakh, Jammu and Kashmir State, India Holotype: BMNH 60.3.19.38A Phrynocephalus zetangensis Wang, Zeng, Fang, Liu., Wu, Papenfuss & Macey, 1996: 27 Type locality: Tsedang, Tibet Holotype: CIB 0156 Type locality: Lake Chomoriri (=Tsho-marari), Jammu and Kashmir State, northern India Syntypes: ZSI 5391–93 Distribution: India: Jammu, Kashmir; Nepal; China: southern Xizang, Xinjiang

Phrynocephalus turcomanus Dunayev, Solovyeva & Poyarkov, 2012 Phrynocephalus helioscopus turcomanus Dunayev, Solovyeva & Poyarkov, 2012: 1391 Type locality: Turkmenbashi (40° 01 ′ 00 ′′ N, 52°58 ′ 00 ′′ E), Turkmenistan Holotype: ZMMU R-13435 Distribution: southwestern Turkmenistan

Phrynocephalus vindumi Golubev, 1998 Phrynocephalus ornatus vindumi Golubev, 1998: 163 Type locality : 35 km. N of Gonabad on road to Torbat-E. Heydariyeh (ca. 34º49' N, 58º47' E), Khorasan Prov., Iran Holotype: CAS 141200 Distribution : Afghanistan:(eastern part of Kavir-i-Namak Desert; eastern Iran: northeast slopes of Kelat (Qalat) Range and Kayen Mountains to Iranian-Afghanistan border

Phrynocephalus versicolor Strauch, 1876

P Bogomolov Subspecies: Phrynocephalus versicolor versicolor Strauch, 1876 Phrynocephalus versicolor Strauch, 1876: 18 Synonyms Phrynocephalus guentheri Bedriaga, 1906: 168 Type locality: Ruo Shui, Inner Mongolia, China Holotype: ZISP 7458 Phrynocephalus blanfordi Bedriaga, 1907: 381 Type locality: Nanshan range along Qinghai – Gansu border, China Lectotype: ZISP 7483.1, designated by Peters, 1984 Phrynocephalus ciliaris Bedriaga, 1907:297 Type locality: Ruo Shui river, from Gaotaï and Jiuquan (Gansu province) to Gaxun Nur (Inner Mongolia), China Holotype: ZISP 7462 Phrynocephalus pewzowi Bedriaga, 1907: 139 Type locality: Ziko’u or Hsi-ku, now Zhugqu, approximately 33º48' N, 104º30' E, southeast Gansu Province, China Holotype: ZISP 7440. Type locality: Alashan Desert, northern China Lectotype : ZISP 3929.1 (designated by Peters, 1984) Distribution : western China: Xinjiang, Gansu, Ningxia, Nei Mongol (= Inner Mongolia); Mongolia; eastern Kazakhstan

Phrynocephalus versicolor siebenrocki Bedriaga, 1909: 333 Phrynocephalus versicolor siebenrocki Bedriaga, 1909: 333 Synonyms : Phrynocephalus bogdanowi Bedriaga, 1909:334 Type locality: Galbyn-Gobi, Mongolia Lectotype: ZISP 7027.1, designated by Peters, 1984 Phrynocephalus helioscopus levis Kashtchenko, 1909: 122 Type locality: vicinity of the Karatal village, approximately 47º36' N, 85º10' E, eastern Kazakhstan Type: ZMTU, not located Phrynocephalus potanini Bedriaga, 1909: 389 Type locality: Ulan Muren, Ordos desert, Inner Mongolia, China Lectotype : ZISP 7438.1, designated by Peters, 1984 Phrynocephalus rostralis Zarevsky, 1930: 216 Type locality: 100 – 200 km from Ulaanbaatar to Kalgan, Mongolia Lectotype: ZISP 12194.1, designated by Peters, 1984 Phrynocephalus immaculatus Zhao, 1995: 47 Type locality: Siziwang Qi (41.5 N, 111.7 E), Inner Mongolia [Autonomous Region], China Holotype: SRNC R8017 Type locality : between Metschin-Ola [= Mieh-Ch’eng-O-La Shan, about 30 km northeast of Barkol] and Barkul town [= Barkol, approximately 43º55' N, 93º00 E, Barkol County]”, Xinjiang Uygur Autonomous Region, China Holotype : ZISP 5156 Distribution: western China: Xinjiang, Gansu, Ningxia, Nei Mongol (= Inner Mongolia); Mongolia; eastern Kazakhstan

Phrynocephalus versicolor doriai Bedriaga, 1909 Phrynocephalus versicolor doriai Bedriaga, 1909: 331 Type locality: Ebinur Lake, Xinjiang Uygur Autonomous Region, China Lectotype: ZISP 5117.1, designated by Peters, 1984 Distribution: China: northwestern Xinjiang

Phrynocephalus versicolor hispidus Bedriaga, 1909 Phrynocephalus versicolor hispidus Bedriaga, 1909: 332 Type locality: Junggar Basin, Xinjiang Uygur Autonomous Region, China Lectotype: ZISP 6637.1, designated by Peters, 1984 Distribution: China: northern Xinjiang

Phrynocephalus versicolor kulagini Bedriaga, 1909 Phrynocephalus versicolor kulagini Bedriaga, 1909: 329 Type locality: Chirgis-nur lake, Uvs province, Mongolia Lectotype : ZISP 5840, designated by Peters, 1984 Distribution: northwestern Mongolia; Russia: Tyva Autonomous Region

Phrynocephalus vlangalii Strauch, 1876 Phrynocephalus vlangalii Strauch, 1876: 20 Phrynocephalus vlangalii Strauch, 1876: 20

D Melnikov Synonyms Phrynocephalus vlangalii nanschanica Bedriaga,1907: 172 Type locality: Nanshan mountains, Xinjiang Uygur Autonomous Region, China Lectotype: ZISP 7081.1, designated by Peters, 1984 Phrynocephalus putjatai Bedriaga 1909: 302 Type locality: Guide County (36º00' N, 101º 40' E), southeast of Qinghai (Kuku-Nor) Lake, Qinghai Province, China Lectotype: (designated by Peters, 1984: 44), ZISP 7447.1 Phrynocephalus vlangalii geckoides Bedriaga, 1909: 466 Type locality: Estuary of the Tschumar River, Qinghai Province, China. Lectotype: ZISP 6601.1, designated by Peters, 1984 Phrynocephalus vlangalii parva Bedriaga, 1909: 468 Type locality: Moron Us and Tuotuo rivers, Qinghai Province, China Lectotype: ZISP 7278.1, designated by Peters, 1984 Phrynocephalus vlangalii pylzowi Bedriaga, 1909: 471 Type locality: Hoang-ho-Quellgebiet, China. Lectotype: ZISP 7084.1, designated by Peters, 1984 Phrynocephalus reldoe Bedriaga,, 1909: 495 Type locality: near Gong He County (36º20' N, 100º60' E), south of Qinghai (Kuku-Nor) Lake, Qinghai Province, China. Holotype : ZISP 6630 Phrynocephalus vlangalii grombtschewskii Bedriaga, 1909: 463 Type locality: Qaidam basin, south of Qinghai Hu mountain, Qinghai province, China Lectotype: ZISP 7269.1, designated by Peters, 1984 Phrynocephalus vlangalii lidskii Bedriaga 1909: 479 Type locality: Yutian mountains, southern Xinjiang Uygur Autonomous Region – northern Xizang (Tibet) Autonomous Region, China Lectotype : ZISP 7258.1, designated by Peters, 1984 Type locality: Lake Kuku-nor, Qinghai Hu, Qinghai province, China Holotype: ZISP 3946 Distribution: China: Xinjiang, Gansu, Qinghai, northwestern Sichuan

Physignathus Cuvier, 1829 Cuvier, G. 1829. Le Règne Animal, distribué d'après son organisation, pour servir de base à l'histoire naturelle des animaux et d'introduction à l'anatomie comparée . Paris : Déterville Vol. 2, Edn 2, i-xv + pp. 1-406. Type species: Phyhignathus cocincinus Cuvier, 1829, by monotypy

Physignathus cocincinus Cuvier, 1829 Physignathus cocincinus Cuvier, 1829: 41

M Pickersgill Synonyms Lophura cuvieri Gray, 1831: 60 Type locality: south Vietnam Holotype: MNHP, not found Istiurus cochinsinensis Cuvier, 1837: plate 18 Type locality: Vietnam Syntypes: MNHP 0.1856, 2536 Dilophyrus mentager Günther, 1861: 188 Type locality: Chantaburi province, southeastern Thailand Holotype: BMNH 1860.8.28.15 Physignathus cocincinus caudicinctus Barbour, 1912 : 192 Type locality: Laokay, north Vietnam Holotype: MCZ 8352 Type locality: Cochinchine, south Vietnam Holotype: MNHP 0. 2537 Distribution: eastern Thailand; Laos; Vietnam; Cambodia

Pogona Storr, 1982 Storr, G.M. 1982. Revision of the bearded dragons (Lacertilia: Agamidae) of Western Australia with notes on the dismemberment of the genus Amphibolurus . Records of the Western Australian Museum 10(2): 199-214 Type species: Agama barbata Cuvier, 1829 by original designation

Pogona barbata (Cuvier, 1829) Agama barbata Cuvier, 1829: 35

S Zozaya Type locality: Australia Syntypes: MNHP 0.7375, MNHP 0.7489 Distribution: Australia: south of Cairns Queensland, New South Wales, Victoria, far as Eyre Peninsula South Australia,

Pogona henrylawsoni Wells & Wellington, 1985 Pogona henrylawsoni Wells & Wellington, 1985: 19

S Zozaya Synonym Pogona brevis Witten, 1994: 331 Type locality: Croydon, Queensland Holotype: QM J32292 Type locality: 118 km W Richmond, Queensland, Australia Holotype: AMS R116984 Distribution: Australia: central Queensland

Pogona microlepidota (Glauert, 1952) Amphibolurus barbatus microlepidotus Glauert, 1952: 168

S Zozaya Type locality: Drysdale River Mission, north Kimberley, Western Australia Syntypes: WAM R591–592 Distribution: Australia: Western Australia: northwestern Kimberley

Pogona minor (Sternfeld, 1919) Amphibolurus barbatus minor Sternfeld, 1919: 78

S Zozaya Subspecies: Pogona minor minor (Sternfeld, 1919 ) Amphibolurus barbatus minor Sternfeld, 1919: 78 Type locality: Hermannsburg Mission, upper Finke River, Northern Territory, Australia Lectotype: SMF 10339, designated by Mertens 1967 Distribution: Australia: western half of South Australia; Western Australia to western Kimberley; southwestern Northern Territory

Pogona minima (Loveridge, 1933) Amphibolurus barbatus minimus Loveridge, 1933: 69 Type locality : West Wallaby Is., Houtman's Abrolhos, Western Australia Holotype : MCZ 32972 Distribution : Australia: Western Australia: Houtman's Abrolhos Islands

Pogona mitchelli (Badham, 1976) Amphibolurus mitchelli Badham, 1976:435 S Zozaya Type locality: Derby, Western Australia Holotype: WAM R26828 Distribution: Australia: northern Western Australia, northwestern Northern Territory

Pogona nullarbor (Badham, 1976) Amphibolurus nullarbor Badham, 1976: 440 Type locality: 16 km NW of Naretha Railway Station, Western Australia Holotype: WAM R29667 Distribution: Australia: Nullarbor Plain of South Australia and Western Australia

Pogona vitticeps (Ahl, 1926) Amphibolurus vitticeps Ahl, 1926:189

S Zozaya Type locality: Australia Holotype: ZMB 30101 Distribution: Australia: interior New South Wales, eastern Northern Territory, interior Queensland, eastern South Australia, northwestern Victoria

Psammophilus Fitzinger, 1843 Fitzinger, L. 1843. Systema Reptilium , fasciculus primus, Amblyglossae. Braumüller et Seidel, Wien: 106 pp. Type species: Agama dorsalis Gray in Griffith & Pidgeon, 1831, by monotypy

Psammophilus blanfordanus (Stoliczka, 1871) Charasia blanfordana Stoliczka, 1871: 194

I Das Type locality: central India Syntypes: ZSI 6592–93, 6595–97, ZSI 6604, ZSI 6609, ZSI 6611–13; ZMB 8098 Distribution: India: Gujarat, Bihar, Orissa, Madhya Pradesh, Chhattisgarh, Maharashtra, Andhra Pradesh, Tamil Nadu, Kerala

Psammophilus dorsalis (Gray, 1831) Agama dorsalis Gray in Griffith & Pidgeon, 1831:56

I Das Type locality: southern India Holotype: BMNH 1846.11.12.6 Distribution: India: Bihar, Jharkhand, Orissa, Madhya Pradesh, Karnataka, Andhra Pradesh, Kerala, Tamil Nadu

Pseudocalotes Fitzinger, 1843 Fitzinger, L. 1843. Systema Reptilium , fasciculus primus, Amblyglossae. Braumüller et Seidel, Wien: 106 pp Type species: Calotes tympanistra Gray, 1831, by monotypy

Pseudocalotes andamanensis (Boulenger, 1891) Calotes andamanensis Boulenger, 1891: 288

Cory Type locality: Andaman Islands, India Holotype: ZMUC 36944146 Distribution: India: Nicobar Islands

Pseudocalotes austeniana (Annandale, 1908) Salea austeniana Annandale, 1908: 97 Type locality: Dafla Hills, Assam , India Holotype: ZSI 3976 Distribution: India: Assam, Arunachal Pradesh; Bhutan

Pseudocalotes baliomus Harvey, Shaney, Hamidy, Kurniawan & Smith, 2017 Pseudocalotes baliomus Harvey, Shaney, Hamidy, Kurniawan & Smith, 2017: 218

EN Smith Type locality: forest west of the mountain crest next to the road from Tapan to Sungai Penuh (2.04294°S, 101.31129°E), Sumatera Barat Department, Sumatra, Indonesia Holotype: MZB 9813 Distribution: Indonesia: West Sumatra

Pseudocalotes bapoensis (Yang & Su, 1979) Calotes kingdonwardi bapoensis Yang & Su, 1979:186 Type locality: Bapo , Gongshan Co., Yunnan Prov., China Holotype: KIZ 730010 Distribution: China: western Yunnan)

Pseudocalotes brevipes (Werner, 1904) Calotes brevipes Werner, 1904: 462 Synonym Acanthosaura quinquicarinata Fan, 1931: 28 Type locality: Luoxiang, Guangxi Zhuang AR, China Holotype: SYSU Type locality: Man-Son Mts, north Vietnam Lectotype : MNHP 1907.164, designated by Hallermann 2000 Distribution: southern China: Guangxi; north Vietnam

Pseudocalotes cybelidermus Harvey, Hamidy, Kurniawan, Shaney & Smith, 2014 Pseudocalotes cybelidermus Harvey, Hamidy, Kurniawan, Shaney & Smith, 2014: 214 Type locality: montane forest at Maura Dua (4.91°S, 104.13°E), Remanan Jaya, Gunung Pesagi (locally known as Masagi), Sumatera Selatan, Indonesia Holotype: MZB 9767 Distribution: Bukit Barisan Range, Sumatra

Pseudocalotes dringi Hallermann & Böhme, 2000 Pseudocalotes dringi Hallermann & Böhme, 2000: 195 Type locality: Gunung Tahan, Pahang, West Malaysia Holotype: BMNH 1906.2.28.10 Distribution: West Malaysia: Gunung Tahan and Gunung Lawit

Pseudocalotes drogon Grismer, Quah, Wood, Davis, Murdoch and Anthony J. Cobos, 2016 Pseudocalotes drogon Grismer, Quah, Wood, Davis, Murdoch and Anthony J. Cobos, 2016: 472 Type locality: 1 km south of Air Terjun Jeriau (3°43.283 N, 101°43.035 E), Fraser’s Hill, Pahang, Peninsular Malaysia Holotype: LSUHC 12223 Distribution: West Malaysia: Pahang State: Fraser’s Hill, Genting Highlands

Pseudocalotes flavigula (Smith, 1924) Calotes flavigula Smith, 1924: 182 Type locality: Guning Berinchang, Pahang, West Malaysia Holotype: BMNH 1946.8.11.14 Distribution: West Malaysia: Cameron Highlands

Pseudocalotes floweri (Boulenger, 1912) Calotes floweri Boulenger 1912: 70 Type locality: Chantaburi, southeastern Thailand Lectotype: BMNH 1946.8.11.25, designated by Hallermann & Böhme 2000 Distribution: southern Thailand: Chantaburi; Cambodia: Elephant Mountains, Cardamom Mountains

Pseudocalotes guttalineatus Harvey, Hamidy, Kurniawan, Shaney & Smith, 2014 Pseudocalotes guttalineatus Harvey, Hamidy, Kurniawan, Shaney & Smith, 2014: 220 A Hamidy Type locality: montane forest above Ngarip (5.28°S, 104.56° E), Lampung, Sumatra, Indonesia Holotype: MZB 9796 Distribution: Bukit Barisan Range of Lampung and Sumatera Selatan, Sumatra

Pseudocalotes kakhienensis (Anderson, 1879) Oriocalotes kakhienensis Anderson, 1879: 806 Synonyms Calotes feae Boulenger, 1888: 477 Type locality: Mogok (17°34’0”N, 95°5’0”E), Pyin Oo Lwin District, Mandalay Division,Burma. Syntypes: BMNH 1893.10.93-94 Salea brachydactyla Rendahl, 1937: 4 Type locality: Kambaiti Pass (25°23’60” N, 98°09’0” E), Kachin State, Burma Holotype: NRM 3958 Type locality: Ponsee (=Pengxi), western Yunnan, China Type: lost Distribution: southern China: western Yunnan; Burma: east of Irrawadi River

Pseudocalotes khaonanensis Chan-ard, Cota, Makchai & Laoteow, 2008 Pseudocalotes khaonanensis Chan-ard, Cota, Makchai & Laoteow, 2008: 26

M Cota Type locality: upper montane scrub of Khao Nan National Park (08°45’49’’N, 99°32’19’’E), Thailand Holotype: THNHM 11791 Distribution: Thailand: Nakhon Si Thammarat Mountains

Pseudocalotes kingdonwardi (Smith, 1935) Subspecies: Pseudocalotes kingdonwardi kingdonwardi (Smith, 1935) Calotes kingdonwardi Smith, 1935: 204 Synonyms Japalura kaulbacki Smith, 1937: 755 Type locality: Nam Tamai Valley, Burma-Tibet border Holotype: BMNH 1946.9.4.72 Type locality: Adung Valley, on the Burma-Tibetan border Holotype: BMNH 1932.6.8.8 Distribution: northern Burma

Pseudocalotes larutensis Hallermann & McGuire, 2001 Pseudocalotes larutensis Hallermann & McGuire, 2001: 256

M Muin Type locality: approx. 200 road meters below the summit of Bukit Larut along the paved road leading to the Bukit Larut Telekom Station, Perak, Malaysia Holotype: TNHC 58040 Distribution: West Malaysia: Perak

Pseudocalotes microlepis (Boulenger, 1888) Calotes microlepis Boulenger, 1888: 476

R Nazarov Type locality: Pla-pu, west of Mt. Muleyit, northern Tenasserim, Burma Holotype: BMNH 1946.8.11.21 Distribution: China: Hainan, Guizhou; southern Thailand: Phong Saly; northern Laos: Langbian Plateau;

Pseudocalotes poilani (Bourret, 1939) Paracalotes poilani Bourret, 1939: 20 Type locality: between Pak Song and Pak Sé, Bas Laos Holotype: MNHP 1948.00.50 Distribution: southern Laos

Pseudocalotes rhaegal Grismer, Quah, Anuar, Muin, Wood, Davis, Murdoch, Burch & Cobos, 2016 Pseudocalotes rhaegal Grismer, Quah, Anuar, Muin, Wood, Davis, Murdoch, Burch & Cobos, 2016: 477 Type locality: Robinson Falls (04°43.283 N, 101°23.129 E), Cameron Highlands, Pahang, Peninsular Malaysia Holotype: LSUHC 12178 Distribution: West Malaysia: Pahang State: Cameron Highlands, Banjaran Titiwangsa range

Pseudocalotes rhammanotus Harvey, Hamidy, Kurniawan, Shaney & Smith, 2014 Pseudocalotes rhammanotus Harvey, Hamidy, Kurniawan, Shaney & Smith, 2014: 226

A Hamidy Type locality: montane forest along the ridge of a mountain south of Danau Ranau (= Lake Ranau) (4.9394° S, 103.85292° E), Lampung, Sumatra, Indonesia Holotype: MZB 10804 Distribution: Barisan Range, southwestern Sumatra, Indonesia

Pseudocalotes saravacensis Inger & Stuebing, 1994 Pseudocalotes saravacensis Inger & Stuebing, 1994: 961 Type locality: Nanga Segerag (125ºN, 112ºE), headwaters of the Engkari River, Lanjak-Entimau Wildlife Sanctuary, Second Division, Sarawak, Borneo, East Malaysia Holotype: FMNH 251522 Distribution: East Malaysia: Sarawak (Borneo) Last seen 1994

Pseudocalotes tympanistriga (Gray, 1831) Calotes tympanistra Gray, 1831:56 Synonym Dendragama fruhstorferi Boettger, 1893: 335 Type locality: Tjisurupan volcano , Java, Indonesia Holotype: SMF 9742 Type locality: Java, Indonesia Holotype: RMNH 3872 Distribution: Indonesia: Java, Sumatra

Pseudocalotes viserion Grismer, Quah, Wood, Anuar & Muin, 2016 Pseudocalotes viserion Grismer, Quah, Wood, Anuar & Muin, 2016: 480 Type locality: crossing the radar tower road at Ulu Kali at Genting Highlands (03°26.166 N, 101°47.021 E), Pahang, Peninsular Malaysia Holotype: LSUHC 12227 Distribution: West Malaysia: Pahang State: Genting Highlands

Pseudocalotes ziegleri Hallermann, Truong, Orlov & Ananjeva, 2010 Pseudocalotes ziegleri Hallermann, Truong, Orlov & Ananjeva, 2010: 32 Type locality: Nuoc Ka forest, near Mang Canh (14°41’57’’ N, 108°12’ 5.7’’ E), Kon Plong District, Kon Tum Province, central Vietnam Holotype: IEBR 330 Distribution: Central Vietnam: Kon Tum Province

Roman Nazarov

Pseudocophotis Manthey, 1997 Manthey, U., and W. Grossmann. 1997. Amphibien & Reptilien Südostasiens. Münster: Natur und Tier . Type species: Cophotis sumatrana Hubrecht, 1879

Roman Nazarov Pseudocophotis kontumensis Ananjeva, Orlov, Truong & Nazarov, 2007 Pseudocophotis kontumensis Ananjeva, Orlov, Truong & Nazarov, 2007:154 Type locality: Kon Du (14°41’25’’N, 108°19’31’’), Mang Canh village, Konplong district, Kon Tum Province, Vietnam Holotype: ZISP 24251 Distribution: central Vietnam: Kon Tum Province

Pseudocophotis sumatrana (Hubrecht, 1879) Cophotis sumatrana Hubrecht, 1879: 243 Type locality: near Padang, Sumatra, Indonesia Holotype : RMNH 3872 Distribution: Indonesia: Sumatra, Java

Pseudotrapelus Fitzinger, 1843 Fitzinger, L. 1843. Systema Reptilium , fasciculus primus, Amblyglossae. Braumüller et Seidel, Wien: 106 pp Type species: Agama sinaita Heyden, 1827, by monotypy

Pseudotrapelus aqabensis Melnikov, Nazarov, Ananjeva & Disi, 2012 Pseudotrapelus aqabensis Melnikov, Nazarov, Ananjeva & Disi, 2012: 146

D Melnikov Type locality: Al Aqabah, southern Jordan Holotype: ZISP 26382 Distribution: southern Jordan; southern Israel; Egypt: Sinai

Pseudotrapelus chlodnickii Melnikov, Śmieowski, Melnikova, Nazarov, & Ananjeva, 2015 Pseudotrapelus chlodnickii Melnikov, Śmieowski, Melnikova, Nazarov, & Ananjeva, 2015: 55

D Melnikov Type locality: Gamamiya (19°27’18.3”N, 32°48’40.5”E), Bayuda Desert, Sudan. Holotype: ZISP 28540 Distribution: Bayuda Desert, Sudan

Pseudotrapelus dhofarensis Melnikov & Pierson, 2012 Pseudotrapelus dhofarensis Melnikov & Pierson, 2012: 145 T Pierson Type locality: Jabal Samhan, Dhofar, Oman Holotype: ZISP 26531 Distribution: Oman: Dhofar

Pseudotrapelus jensvindumi Melnikov, Ananjeva & Papenfuss, 2013 Pseudotrapelus jensvindumi Melnikov, Ananjeva & Papenfuss, 2013: 80

D Fischer Type locality: Jebel al Akhdah, Qatana, Nizwa Wilayat, Oman Holotype: CAS 225340 Distribution: eastern Oman

Pseudotrapelus neumanni Tornier, 1905 Agama neumanni Tornier, 1905:384 Type locality: Haitthalhin, Ladadj, Saudi Arabia Syntypes: NMHW 23341, ZMB 27418 Distribution: Yemen; western Saudi Arabia

Pseudotrapelus sinaitus (Heyden, 1827) Subspecies: Pseudotrapelus sinaitus sinaitus (Heyden, 1827) Agama sinaita Heyden, 1827: 10 D Melnikov Synonym Agama arenaria Heyden, 1827: 12 Type locality: upper Egypt Lectotype: SMF 9972, designated by Mertens 1967 Type locality: Sinai, Egypt Holotype: SMF 9971 Distribution: souteastern Libya; eastern Egypt and Sinai; eastern Sudan

Pseudotrapelus sinaitus werneri Moravec, 2002 Pseudotrapelus sinaitus werneri Moravec, 2002: 135 Holotype: NMP6V 34860/1 Type locality: Ar’ Raqiyeh (=Er Raqaïyé)(32.18ºN, 37.08ºE), As- Suwayda governate, Syria Distribution: southern Syria; northern Jordan

Ptyctolaemus Peters, 1865 Peters, 1867 : Über Flederthiere (Pteropus gouldii, Rhinolophus deckenii, Vespertilio lobipes, Vesperugo temminckii) und Amphibien (Hypsilurus godeffroyi, Lygosoma scutatum, Stenostoma narirostre, Onychocephalus unguirostris, Ahaetulla poylepis, Pseudechis scutellatus, Hoplobatrachus reinhardtii, Hyla coriacea) . Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, vol. 1867, p. 703-712 Type species: Otocryptis gularis Peters, 1864, by monotypy

Ptyctolaemus collicristatus Schulte & Vindum, 2004 Ptyctolaemus collicristatus Schulte & Vindum, 2004: 230

H Tun Type locality: Min Dat Township (21°22 ′20.1 ″N, 93°58 ′34.6 ″E), Min Dat District, Chin State, Burma Holotype: CAS 227489 Distribution: Burma: Chin state

Ptyctolaemus gularis (Peters, 1864) Otocryptis gularis Peters, 1864: 386 S Harikishnan Type locality: Margherita, in Patkai Mountains, Upper Assam, India Holotype: ZMB 5004 Distribution: northeastern India: Assam, Arunachal Pradesh, Mizoram; northern Bangladesh; China: Xizang = Tibet; Burma: Kachin, Chin

Rankinia Wells & Wellington, 1984 Wells, R.W. & Wellington, C.R. 1984. A synopsis of the class Reptilia in Australia. Australian Journal of Herpetology 1(3-4): 73-129 Type species: Grammatophora muricata diemensis Gray, 1841, by original designation

Rankinia diemensis (Gray, 1841) Grammatophora muricata diemensis Gray, 1841: 88

S Zozaya Synonyms Grammatophora angulifera Gray, 1845: 252 Type locality: Tasmania Syntypes: BMNH 1946.9.4.29–30, BMNH 1946.9.4.58–66, BMNH xxiv.49.f Agama caelaticeps Smith, 1849: pl 74 Type locality: South Africa, in error Syntypes: BMNH 1946.9.4.33–34 Rankinia boylani Wells & Wellington, 1984: 82 Type locality: Mt Victoria (33°35'S 150°15'E), New South Wales Holotype: AMS R111951 Type locality: Tasmania Syntypes: BMNH XXIV.49.g-i Distribution: Australia: coast ranges of New South Wales and Victoria, northern and eastern Tasmania

Salea Gray, 1845 Gray, J.E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum . Trustees of the British Museum/Edward Newman, London: xxvii + 289 pp. Type species: Salea horsfieldii Gray, 1845, by monotypy

Salea anamallayana (Beddome, 1878) Lophosalea anamallayana Beddome, 1878: 153

Praveenmac Synonyms Lophosalea indica Beddome, 1878: 153 Type locality: Pulney Hills, India Syntype: ZMB 10223 Salea anamallayana triangularis Roux, 1928: 452 Type locality: Mariyanshola, Palnis Hills, India Holotype: NHMB 9687 Type locality: Anamallay Mountains, Travancore, India Holotype: BMNH 1964.8.14.84 Distribution: India: Kerala, Tamil Nadu

Salea gularis Blyth, 1854 Salea gularis Blyth in Jerdon, 1854: 473 Type locality: Mirzapur, Uttar Pradesh, India Type: not found Distribution: India: Uttar Pradesh Last seen 1854

Salea horsfieldii Gray, 1845 Salea horsfieldii Gray, 1845:242

L. Shyamal Synonyms Calotes hirsutus Duméril, 1851: 88 Type locality: India Holotype: MNHP 0.6892 Calotes trispinosus Duméril, 1851:88 Type locality: Nilgiri Hills, India Syntypes: MNHP 0.2558-59 Mecolepis sulcatus Duméril, 1851: 89 Type locality: Nilgiri Hills, India Holotype: MNHP 0.6893 Type locality: southern India Syntypes: BMNH 1946.8.14.11–12 Distribution: India: Western Ghats, Tamil Nadu

Sarada Deepak, Karanth and Giri, 2016 Deepak, V, V.B. Giri, M. Asif, S.K. Dutta, R. Vyas, A.M. Zambre, H. Bhosale, K.P. Karanth. 2016. Systematics and phylogeny of Sitana (Reptilia: Agamidae) of Peninsular India, with the description of one new genus and five new species. Contributions to Zoology , 85 (1) 67-111 Type species: Sitana deccanensis Jerdon, 1870

Sarada darwini Deepak, Karanth, Dutta and Giri, 2016 Type locality : Bidnal(5.32724°N, 75.15823°E), Dharwad District, Karnataka State Holotype: NCBS-AQ051 Distribution: Belgaum & Dharwad Districts, Karnataka State; Kolhapur & Sangli District, southern Maharashtra State

Sarada deccanensis (Jerdon, 1870) Sitana deccanensis Jerdon, 1870: 76

N Sant Type locality : Deccan, India Syntypes: BMNH 1946.8.27.39-40 Distribution: Aurangabad, Jalnat, Nanded Districts, Maharashtra State, southwestern India

Sarada superba Deepak, Zambre, Bhosale and Giri, 2016 Sarada superba Deepak, Zambre, Bhosale and Giri, 2016: 109

V Giri Type locality : Chalkewadi (17.57635°N, 73.82430°E), Satara District, Maharashtra State. Holotype: NCBS-AQ052 Distribution: Satara District, Maharashtra state, India

Sitana Cuvier, 1829 Cuvier, G.J.L.N.F.D. 1829. Le Regne Animal Distribué, d'apres son Organisation, pur servir de base à l'Histoire naturelle des Animaux et d'introduction à l'Anatomie Comparé . Nouvelle Edition [second edition]. Vol. 2. Les Reptiles. Déterville, Paris, i-xvi, 1-406 Type species: Sitana ponticeriana Cuvier, 1829, by monotypy

Sitana attenboroughii Sadasivan, Ramesh, Palot, Mirza 2018 Sitana attenboroughii Sadasivan, Ramesh, Palot, Mirza 2018: 547 Type locality: Trivandrum (8.313387°, 77.070933°), Kerala, India Holotype: BNHS 2481 Distribution: India: Kerala State: coastal sand dunes of Poovar in Trivandrum district

Sitana bahiri Amarasinghe, Ineich & Karunarathna, 2015 Sitana bahiri Amarasinghe, Ineich & Karunarathna, 2015: 70 Type locality: Yala National Park (6°22 ′N, 81°31 ′E), Sri Lanka Holotype: WHT 1434A Distribution: Drier coastal areas in southeastern Sri Lanka

Sitana devakai Amarasinghe, Ineich & Karunarathna, 2015 Sitana devakai Amarasinghe, Ineich & Karunarathna, 2015: 77

V Weeratunge Type locality: Nagagamuwa-Puttalam (8°10 ′N, 79°50 ′E), Sri Lanka Holotype: WHT 0111E Distribution: Drier coastal areas in the north and north-western parts of Sri Lanka

Sitana fusca Schleich & Kästle, 1998 Sitana fusca Schleich & Kästle, 1998: 94 Type locality: Ghauri Dhara Road Camp (26º59'25.0" N, 85º53'45.6" E), 0.8 km north of the road Janatir- Bardibas, eastern Nepal Holotype: ZMS 853/2012 Distribution: eastern Nepal Last seen 1998

Sitana gokakensis Deepak, Khandekar, Chaitanya, Karanth, 2018 Sitana gokakensis Deepak, Khandekar, Chaitanya, Karanth, 2018: 336 Type locality: Gokak plateau (16.18618°N, 74.75952°E), Belagavi district, Karnataka, India Holotype: BNHS 2490 Distribution: India: Karnatka State: Gokak Plateau

Sitana kalesari Bahuguna, 2015 Sitana kalesari Bahuguna, 2015: 527 Type locality: Paddal line (30°23'12.2”N, 077°01'44.5”E), Kalesar National Park, Wildlife Sanctuary (District: Yammuna Nagar), Haryana, India Holotype: NRC,ZSI 151 Distribution: northern India: Haryana State

Sitana laticeps Deepak and Giri, 2016 Sitana laticeps Deepak and Giri, 2016: 97 Type locality : Bopdev Ghat (18.372818°N, 73.9128°E), Pune District, Maharashtra State , Holotype : NCBS-AQ056 Distribution: Balaghat Hill Range, Maharashtra State, India

Sitana marudhamneydhal Deepak, Khandekar, Varma, Chaitanya, 2016 Sitana marudhamneydhal Deepak, Khandekar, Varma, Chaitanya, 2016: 171 Type locality : Kallidaikurichi (8°39’54” N; 77°27’15” E), Tirunelveli District, Tamil Nadu, India Holotype: NCBS AQ460 Distribution: southern Tamil Nadu State, India

Sitana ponticeriana Cuvier, 1829 Sitana ponticeriana Cuvier, 1829: 43

Jahnavi Pai Synonyms Sitana minor Günther, 1864: 135 Type locality: Madras, India Syntypes: BMNH 1946.8.27.41-44, 1946.8.30.53-56 Sitana ponticeriana mucronata Deraniyagala, 1958: 4 Type locality: Hiniduma (06°20’N, 80°18’E), Sri Lanka Holotype: NMSL, lost Type locality: Pondichéry , Coromandel, India Lectotype: MNHP 0. 6901 Distribution: northern plains of Tamil Nadu State, India

Sitana schleichi Anders & Kästle, 2002 Sitana schleichi Anders & Kästle, 2002: 653 Type locality: Suklaphantah National Park, western Nepal Holotype: ZMS 848/2012 Distribution: western Nepal

Sitana sivalensis Schleich, Kästle & Shah, 1998 Sitana sivalensis Schleich, Kästle & Shah, 1998: 87 Type locality: between Shivpur (Shivapur) and Bankas near the Surai Khola (27º43'17"N, 82º50'40"E), Lumbini Zone, Kapilabastu District, in southern West Nepal Holotype: SHHS 95/1 Distribution: southern Nepal

Sitana spinaecephalus Deepak, Vyas and Giri, 2016 Sitana spinaecephalus Deepak, Vyas and Giri, 2016: 93 K Upadhay Type locality: Halol (22.43163°N, 73.62966°E), Panchmahal District, Gujarat State Holotype: NCBS-AQ055 Distribution: Gujarat, Maharashtra, Rajasthan states, India

Sitana thondalu Deepak, Khandekar, Chaitanya, Karanth, 2018 Sitana thondalu Deepak, Khandekar, Chaitanya, Karanth, 2018: 343 Type locality : east bank of Nagarjuna Sagar reservoir (16.55557°N, 79.30134°E), Guntur district, Andhra Pradesh, India Holotype: BNHS 2492 Distribution: India: Andhra Pradesh State: Macherala and Nagarjuna Sagar

Sitana visiri Deepak, 2016 Sitana visiri Deepak, 2016: 89

V Deepak Type locality: Tuticorin (8.784993°N, E78.156341°E), Thoothukudi, Tamil Nadu state, India Holotype: NCBS-AQ054 Distribution: southern plains of Tamil Nadu south of the Cauvery River, India

Thaumatorhynchus Parker, 1924 Parker, H.W. 1924. Description of a new agamid lizard from Sumatra. Annals and magazine of natural history (9) 14: 624-625 Type species: Thaumatorhynchus brooksi Parker, 1924, by original designation

Thaumatorhynchus brooksi Parker, 1924 Thaumatorhynchus brooksi Parker, 1924:624

N Cegalerba

Type locality: Lebong Tandai, Res. Benkoelen, Sumatra , Indonesia Holotype: BMNH 1946.8.30.90 Distribution: Indonesia: Sumatra

Trapelus Cuvier, 1817 Cuvier, G. 1817. Le règne animal distribué d’après son organisation, pour servir de base a l’historire naturelle des animaux et d’introduction a l’anatomie comparée . Vol. 2. Les reptiles, les poissons, les mollusques et les annélides. Déterville, Paris Type species: Agama mutabilis Merrem, 1820, by monotypy

Trapelus agilis (Olivier, 1807)

I Das Subspecies: Trapelus agilis agilis (Olivier, 1807) Agama agilis Olivier, 1807: 394 Type locality: vicinity of Bagdad, Iraq Syntypes: MHNP 0.5108, BMNH 1874.11.23.113 Synonyms Agama kirmanensis Nikolsky, 1899: 389 Type locality: Kurin, Sargad region, Sistan and Baluchestan province, Iran Holotype: ZISP 9321 Agama kirmanensis brevicauda Nikolsky, 1907: 272 Type locality: Kohrud (33°38’N, 51°27’E), Esfahan province, Iran Holotype: ZISP 10335 Distribution: central Iran; Iraq; southern Afghanistan; southwestern Pakistan

Trapelus agilis isolepis (Boulenger, 1885) Agama isolepis Boulenger, 1885: 342 Type locality: between Magas and Bampur, Iran Lectotype: BMNH 1874.11.23.113, designated by Anderson 1966 Distribution: Afghanistan; Iran

Trapelus agilis khuzistanensis Rastegar-Pouyani, 1999 Trapelus agilis khuzistanensis Rastegar-Pouyani, 1999: 91 Type locality: 5 km northwest of Haft-Gel on the road to Shushtar, Khuzistan Province, Iran Holotype: GNHM Re. ex. 5224 Distribution: southwestern Iran

Trapelus agilis pakistanensis Rastegar-Pouyani, 1999 Trapelus agilis pakistanensis Rastegar-Pouyani, 1999: 94 Type locality: Gaj-River, Kirthar Range, southeastern Baluchistan, Pakistan Holotype: SMF 63258 . Distribution: southeastern Pakistan; adjacent northwestern India

Trapelus agnetae (Werner, 1929) Agama agnetae Werner, 1929: 239

D Melnikov Synonym Agama pallidus haasi Werner, 1971:222 Type locality: Azraq, Jordan Holotype: BMNH 1965.800 Type locality: Azraq, Jordan Holotype : NMHW 23349 Distribution: Jordan; Iraq; northern Saudi Arabia; Kuwait

Trapelus boehmei Wagner, Melville, Wilms & Schmitz, 2011 Trapelus boehmei Wagner, Melville, Wilms & Schmitz, 2011: 900 A Roux Type locality: between Akka and Icht, Morocco Holotype: ZFMK 49751 Distribution: Morocco; Mauretania; Algeria

Trapelus flavimaculatus Rüppell, 1835 Trapelus flavimaculatus Rüppell, 1835: 12

D Fischer Type locality: Jeddah, Saudi Arabia Holotype: BMNH 1920.1.20.1241 Distribution: eastern Saudi Arabia; Oman; United Arab Emirates

Trapelus jayakari (Anderson, 1896) Agama jayakari Anderson, 1896: 65

J. Pfleiderer Type locality: Masqa ṭ, Oman Syntypes: BMNH 1946.8.14.86-91 Distribution: southeastern Arabia; Oman

Trapelus megalonyx Günther, 1864 Trapelus megalonyx Günther, 1864: 159 Type locality: eastern Afghanistan Holotype: BMNH 1946.8.11.34 Distribution: western Pakistan: Baluchistan; eastern Afghanistan; southeastern Iran

Trapelus microtympanum (Werner, 1895) Agama microtympanum Werner, 1895:15 Type locality: Iran Holotype: NMHW 19457 Distribution: Iran

Trapelus mutabilis (Merrem, 1820)

L Fidel Subspecies: Trapelus mutabilis mutabilis (Merrem, 1820) Agama mutabilis Merrem, 1820: 50 Synonyms Trapelus aegyptius Cuvier, 1829: 54 Type locality: Egypt Holotype: MNHP 0.2608 Agama inermis Reuss, 1834: 33 Type locality: southern Egypt Lectotype: SMF 9991, designated by Mertens 1967 Agama gularis Reuss, 1834: 36 Type locality: southern Egypt Lectotype: SMF 9993, designated by Mertens 1967 Agama aspera Werner, 1893: 359 Type locality: Algerian Sahara, between Kef-el-Dhor and Chegga Type: not located Agama latastii Boulenger, 1885: 344 Type locality: Egypt Holotype: BMNH XXIV.64.c Type locality: 10 km north-west of Cairo, Egypt Neotype: ZFMK 64395, designated by Wagner et al. 2011 Distribution western Egypt

Trapelus mutabilis pallidus (Reuss, 1834) Agama pallida Reuss, 1834: 38 Synonyms: Agama deserti Lichtenstein, 1823:111 Type locality: Egypt Syntypes: ZMB 767-769, ZMB 55736 Agama loricata Reuss, 1834: 40 Type locality:southern Egypt Lectotype: SMF 10008, designated by Mertens 1967 Agama nigrofasciata Reuss, 1834: 40 Type: Nubia, Sudan Lectotype: SMF 1001, designated by Mertens 1967 Agama leucostygma Reuss, 1834: 44 Type locality: southern Egypt Lectotype: SMF 10027, designated by Mertens 1967 Agama straminea Lichtenstein, 1845: 259 [in Gray 1845] Type locality: Arabia Syntypes: ZMB 759-760 Type locality: upper Egypt Holoytype: SMF 10007 Distribution: northeastern Egypt, Sinai ; Sudan; Saudi Arabia;, Syria; Israel; Jordan; Iraq

Trapelus mutabilis poppeki Wagner, Melville, Wilms, Schmitz, 2011 Trapelus mutabilis poppeki Wagner, Melville, Wilms, Schmitz, 2011: 896 Type locality: east of Tarbu (24.48.27ºN, 16.19.11ºE), Libya Holotype: ZFMK 63678 Distribution: Libya

Trapelus rubrigularis Blanford, 1875 Trapelus rubrigularis Blanford, 1875: 233 Type locality: foot of the Khirthar range, Upper Sind, Pakistan Syntypes: BMNH 1946.8.11.18-19; ZSI 5043; ZMB 8684 Distribution: Iran; Pakistan: Baluchistan, Waziristan, Sind

Trapelus ruderatus (Olivier, 1804)

D Hofmann Subspecies: Trapelus ruderatus ruderatus (Olivier, 1804) Agama ruderata Olivier, 1804: 395 Synonyms Agama lessonae De Filippi, 1865: 353 Type locality: Iran Holotype: MZUT R307 Agama persica Blanford,1881: 674 Type locality: Iran Syntypes: BMNH 1946.8.11.39-42 , 1946.8.11.30 Type locality: Isfahan, in central Iran Neotype : MZUT R307, designated by Ananjeva et al. 2013 Distribution: southern Azerbaijan; Syria; Lebanon; northern Saudi Arabia; Iraq; Iran; northern Afghanistan; northern Pakistan; eastern Turkey

Trapelus ruderatus fieldii (Haas & Werner, 1969) Agama persicus fieldii Haas & Werner, 1969: 337 Type locality: between Al-Qaysumeh and Turaif, northeastern Saudi Arabia Holotype: MCZ 56866 Distribution: Jordan; northern Saudi Arabia; Israel; Kuwait

Trapelus sanguinolentus (Pallas, 1814) D Melnikov Subspecies: Trapelus sanguinolentus sanguinolentus (Pallas, 1814) Lacerta sanguinolenta Pallas, 1814: 23 Type locality: Kum-Ankatar, Terek River Valley , Kazakhstan Type: unknown Distribution: Russia: Dagestanp; Kazakhstan; Kyrgyzstan; southwestern Tajikistan; northern Iran; northern Afghanistan; northwestern China: western Xinjiang

Trapelus sanguinolentus aralensis (Lichtenstein, 1823) Agama aralensis Lichtenstein, 1823: 144 Type locality: east side of the Aral Sea, Kazakhstan Syntypes: ZMB 753-758, Distribution: Kazakhstan, Uzbekistan

Trapelus savignii (Duméril & Bibron, 1837) Agama savignii Duméril & Bibron, 1837: 508 Type locality: El Arish, northern Sinai, Egypt Neotype : ZFMK 77470, designated by Wagner & Crochet 2009 Distribution: northern Egypt; southern Israel

Trapelus schmitzi Wagner & Böhme, 2006 Trapelus schmitzi Wagner & Böhme, 2006: 900 Type locality: Guelta Archei, Ennedi Mountains, Chad Holotype: ZFMK 2590 Distribution: mountains of eastern Chad to southern Algeria

Trapelus tournevillei (Lataste, 1880) Agama tournevillei Lataste, 1880: 325 A Fellous Type locality: Ouargla (Wargla?), Algeria Holotype: BMNH 1920.1.20.1241 Distribution: Algeria: Sahara; southern Tunisia

Tropicagama Melville, Ritchie, Chapple, Glor & Schulte, 2018 . Melville, J., E.J. Ritchie, S.N.J. Chapple, R.E. Glor, J.A. Schulte. 2018. Diversity in Australia’s tropical savannas: An integrative taxonomic revision of agamid lizards from the genera Amphibolurus and Lophognathus (Lacertilia: Agamidae). Memoirs of Museum Victoria 77: 41–61 . Type species: Grammatophora temporalis (part.) Günther, 1867.

Stephen Zozaya Tropicagama temporalis (Günther, 1867) Grammatophora temporalis Günther, 1867: 52 Synonyms: Lophognathus lateralis Macleay, 1877:103 Type locality: Mawatta, Binaturi River (as Katow), Papua New Guinea Holotype: AMS R31882 Lophognathus labialis Boulenger, 1883:225 Type locality: Port Essington, Northern Territory Syntypes: BMNH 1946.8.12.72, BMNH 1946.8.12.63 Lophognathus maculilabris Boulenger, 1883: 226 Type locality: Tanimbar Islands, Indonesia Syntypes: BMNH 1946.8.28.70-71 Type locality : Port Essington, Northern Territory. Lectotype : BMNH 1946.8.28.72, designated by Cogger 1983 Distribution: Indonesia: Papua, Kei Islands-Tam and Kur, Tanimbar Islands; southern Papua New Guinea; Australia: northern North Territory, northern Queensland, extreme northeastern West Australia

Tympanocryptis Peters, 1863 Peters, W. 1863. Übersicht der von Hrn. Richard Schomburgk an das zoologische Museum eingesandten Amphibien, aus Buchsfelde bei Adelaide in Südaustralien. Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1863: 228-236 Type species: Tympanocryptis lineata Peters, 1863, by monotypy

Tympanocryptis centralis Sternfeld, 1925 Tympanocryptis centralis Sternfeld, 1925: 234

S Macdonald Type locality: Hermannsberg Mission, upper Finke River, Northern Territory, Australia Lectotype: SMF 10349, designated by Mertens 1967 Distribution: Australia: southern Northern Territory and adjacent eastern Western Australia and northern South Australia

Tympanocryptis cephalus Günther, 1867 Tympanocryptis cephalus Günther, 1867: 52

J Vos Synonym Tympanocryptis cephalus gigas Mitchell, 1948: 65 Type locality: between Ashburton and Gascoyne Rivers, WA. Holotype: SAMA R2434 Type locality: Nickol Bay, Western Australia Lectotype : BMNH 1946.8.12.78, designated by Cogger 1983 Distribution: Australia: central Western Australia

Tympanocryptis condaminensis Melville, Smith, Hobson, Hunjan, Shoo, 2014 Tympanocryptis condaminensis Melville, Smith, Hobson, Hunjan, Shoo, 2014: 8 R Ashdown Type locality: 30 km south of Oakey (27° 39’ S, 151° 36’ E), Queensland, Australia. Holotype: QM J34744 Distribution: eastern Darling Downs: north to the Pirrinuan/Jimbour area, west as far as Dalby and south to the township of Clifton, Queensland, Australia

Tympanocryptis diabolicus Doughty, Kealley, Shoo & Melville, 2015 Tympanocryptis diabolicus Doughty, Kealley, Shoo & Melville, 2015: 100

J Vos Type localit y: Mount Brockman (22°17'31"S, 117°16'23"E), Western Australia Holotype: WAM R135413 Distribution: Australia: Western Australia: Pilbara region, Hamersley Range

Tympanocryptis fortescuensis Doughty, Kealley, Shoo & Melville, 2015 Tympanocryptis fortescuensis Doughty, Kealley, Shoo & Melville, 2015: 104

J Vos Type locality : 4 km south-south-east of Wanna Wanna Pool (21°23'41"S, 117°10'14"E), Western Australia Holotype : WAM R158076 Distribution: Australia: Western Australia: Pilbara region

Tympanocryptis gigas Mitchell, 1948 Tympanocryptis gigas Mitchell, 1948: 65 Type locality: Ashburton and Gascoyne Rivers, Western Australia Neotype: SAMA R68044 Distribution: Australia: Western Australia: Gascoyne region

Tympanocryptis houstoni Storr, 1982 Tympanocryptis houstoni Storr, 1982: 62 Type locality: 10 km SSE Cocklebiddy (32°07'S 126°06'E), Western Australia Holotype: WAM R53427 Distribution: Australia: Nullarbor Plain of western South Australia and southeastern Western Australia

Tympanocryptis intima Mitchell, 1948 Tympanocryptis intima Mitchell, 1948: 60

S Zozaya Type locality: Oodnadatta, South Australia Holotype: SAMA R2331 Distribution: Australia: southeastern Northern Territory, southwestern Queensland, northern South Australia, northwestern New South Wales

Tympanocryptis lineata Peters, 1863 Subspecies: Tympanocryptis lineata lineata Peters, 1863 Tympanocryptis lineata Peters, 1863: 230 Synonym Tympanocryptis karumba Wells & Wellington,1985:20 Type locality: Karumba (17°29'S 140°50'E), Queensland Holotype: AMS R63438 Tympanocryptis telecom Wells & Wellington, 1985 Type locality: Black Mountain, Australian Capital Territory Holotype: not located Type locality: Loos, 4.5 km W Gawler, South Australia Lectotype: ZMB 740 Distribution: Australia: Northern Territory, southern Queensland, northern New South Wales, South Australia

Tympanocryptis lineata macra Storr, 1982 Tympanocryptis lineata macra Storr, 1982: 61 Type locality: 16 km south of main dam at Lake Argyle (16°15'S, 128°40'E), Western Australia Holotype: WAM R44553 Distribution: eastern Western Australia

Tympanocryptis mccartneyi Melville, Chaplin, Hutchinson, Sumner, Gruber, MacDonald, Sarre, 2019 Tympanocryptis mccartneyi Melville, Chaplin, Hutchinson, Sumner, Gruber, MacDonald, Sarre, 2019: 19 Type locality : Bathurst, NSW Holotype : AMS R26077 Distribution : Australia: New South Wales: Central Tablelands

Tympanocryptis osbornei Melville, Chaplin, Hutchinson, Sumner, Gruber, MacDonald, Sarre, 2019 Tympanocryptis osbornei Melville, Chaplin, Hutchinson, Sumner, Gruber, MacDonald, Sarre, 2019: 18 Type locality : Cooma, NSW Holotype: SAMA R43098 Distribution : Australia: New South Wales: Monaro tablelands

Tympanocryptis pentalineata Melville, Smith, Hobson, Hunjan, Shoo, 2014 Tympanocryptis pentalineata Melville, Smith, Hobson, Hunjan, Shoo, 2014: 9 Type locality: Type locality: 47.9 km south-west of the intersection between the Burke Development Rd (Route 83) and the Gulf Development Rd (Route 1)(18° 06’ 29’’ S, 140° 53’ 00’’ E), Queensland, Australia . Holotype: NMV D74074 Distribution: south of Normanton in the gulf region of far northern Queensland, Australia

Tympanocryptis pinguicolla Mitchell, 1948 Tympanocryptis lineata pinguicolla Mitchell, 1948: 70

S Macdonald Type locality: s outhern Victoria, Australia Holotype: SAMA R2468a Distribution: Australia: central Victoria, southeastern New South Wales

Tympanocryptis pseudopsephos Doughty, Kealley, Shoo & Melville, 2015 Tympanocryptis pseudopsephos Doughty, Kealley, Shoo & Melville, 2015: 107

J Vos Type locality : 31 km south-west of Doolgunna Homestead (25°52'33"S, 119°01'41"E), Western Australia Holotype: WAM R154790 Distribution: Australia: Western Australia: Goldfields region

Tympanocryptis tetraporophora Lucas & Frost, 1895 Tympanocryptis tetraporophora Lucas & Frost, 1895: 65

S Zozaya Type locality: Adminga and Dalhousie, South Australia Lectotype : NMV D7701, designated by Coventry, 1970 Distribution: Australia: northwestern New South Wales, eastern Northern Territory, western Queensland, South Australia

Tympanocryptis uniformis Mitchell, 1948 Tympanocryptis uniformis Mitchell, 1948:76 Type locality: near Darwin, Northern Territory, Australia Holotype : SAMA R705 Distribution: Australia: northern Northern Territory

Tympanocryptis wilsoni Melville, Smith, Hobson, Hunjan, Shoo, 2014 Tympanocryptis wilsoni Melville, Smith, Hobson, Hunjan, Shoo, 2014: 11

S Zozaya Type locality: Mount Abundance Rd, 40 km southwest of Roma (26° 42’ 10’’ S, 148° 29’ 10’’ E), Queensland, Australia Holotype: QM J89119 Distribution: Australia : near the town of Roma: from Hodgson approximately 20 km west to Mt Abundance approximately 50 km southwest of Roma, Queensland.

Uromastyx Merrem, 1820 Merrem, B. 1820. Versuch eines Systems der Amphibien I (Tentamen Systematis Amphibiorum). J. C. Krieger, Marburg, 191 pp Type species: Stellio spinipes Daudin, 1802 (= Uromastyx aegyptia ), by subsequent designation

Uromastyx acanthinura Bell, 1825 Uromastyx acanthinura Bell, 1825:45

M O’Shea Synonym Uromastix mutabilis Fischer, 1885: 272 Type locality: unknown Type: unknown Type locality: near Biskra, north to el Kantara, Algeria Holotype: OUM 7845

Uromastyx aegyptia (Forskal, 1775)

M O’Shea Subspecies: Uromastyx aegyptia aegyptia (Forskål, 1775) Lacerta aegyptia Forskål, 1775: 13 Synonyms Lacerta harbai Forskål, 1775: 9 Type locality: Cairo, Egypt Type: unknown Stellio spinipes Daudin, 1802: 31 Type locality: Egypt Holotype: MNHP 0.8297 Type locality: Suez, at the road to Cairo, Egypt Neotype: ZFMK 44216 Distribution: northeastern Egypt, the Sinai Peninsula; Palestine; extreme northwestern Saudi Arabia.

Uromastyx aegyptia microlepis Blanford, 1874 Uromastix microlepis Blanford, 1874: 658 Type locality: Basrah, Iraq Lectotype: BMNH 1946.8.14.55, designated by Wilms & Bohme, 2000 Distribution: Saudi Arabia; Yemen; Oman; United Arab Emirates; Qatar; Kuwait; Jordan; Syria; Iraq; coastal Iran.

Uromastyx aegyptia leptieni Wilms & Böhme, 2000 Uromastyx leptieni Wilms & Böhme, 2000: 142 Type locality: Wadi Siji, United Arab Emirates Holotype: ZFMK 52398 Distribution: northern Oman; northeastern United Arab Emirates

Uromastyx alfredschmidti Wilms & Böhme, 2001 Uromastyx alfredschmidti Wilms & Böhme, 2001: 95 Type locality: Tassili N’Ajjer, Tamrit Plateau, approx. 30 km northeast Djanet, Algeria Holotype: ZFMK 24643 Distribution: southern Algeria; southwestern Libya

Uromastyx asmussi (Strauch, 1863) Centrotrachelus asmussi Strauch, 1863: 479

O Mozaffari Type locality: Sar-i-tschah, Iran Holotype : ZISP 3029 Distribution: Iran: Esfehan, Kerman, Khorasan, Baluchistan-Sistan; Pakistan: Baluchistan; southern Afghanistan

Uromastyx benti (Anderson, 1894) Aporoscelis benti Anderson, 1894: 376

R Vandevender Synonym Uromastix simonyi Steindachner, 1899: 143 Type locality: Azzan, Yemen Syntypes: NMHW 21214:1–2 Type locality: Wadi Hadramaut, Yemen Lectotype: BMNH 1946.8.11.72, designated by Wilms & Bohme 2000 Distribution: southern and southeastern Yemen; southwestern Oman

Uromastyx dispar Heyden, 1827 Subspecies: Uromastyx dispar dispar Heyden, 1827 Uromastyx dispar Heyden, 1827: 5 Type locality: desert near Ambukol and Dongola, Sudan Holotype: SMF 10417 Distribution: western Sudan; Chad: Tibesti and Ennedi mountains

Uromastyx dispar flavifasciata Mertens, 1962 Uromastyx dispar flavifasciata Mertens, 1962: 427

M O’Shea Type locality: approx. 50 km north of Dakar, Senegal Holotype: SMF 58032 Synonym: Uromastyx flavifasciata obscura Mateo, Geniez, Lopez-Jurado & Bons, 1999: 104 Type locality: Gor el Carrashit (21°23’N, 14°27’W), Western Sahara. Holotype: DB.ULPGC 7 Distribution: southern Western Sahara, Mauritania; southwestern Algeria

Uromastyx dispar hodhensis Trapé & Trapé, 2012 Uromastyx dispar hodhensis Trapé & Trapé, 2012: 33 Type locality: on the pass of Enji (18º06N, 08º02E), Mauretania Holotype: MNHP 2011.0272 Distribution: southeastern Mauretania

Uromastyx dispar maliensis Joger & Lambert, 1996 Uromastyx maliensis Joger & Lambert, 1996: 24 Type locality: 40 km southeast of Gao, Mali Holotype: HLMD RA 1545 Distribution: northwestern Mali; southwestern Algeria

Uromastyx geyri Müller, 1922 Uromastyx geyri Müller, 1922: 193

M O’Shea Type locality: Gara Djenoum, Ahaggar Mountains, southern Algeria Neotype: ZFMK 9230, designated by Muller 1951 Distribution: northeastern Mali; southern Algeria

Uromastyx hardwickii Gray, 1827 Uromastyx hardwicki Gray in Hardwicke & Gray, 1827: 219

M O’Shea Synonyms Uromastyx griseus Cuvier, 1829: 34 Type locality: Bengal, India Holotype: MNHP 0.1448 Uromastyx reticulatus Cuvier, 1829:34 Type locality: unknown Syntypes: MNHP 0.2635, 6978 Centrocercus similis Fitzinger, 1843: 86 Type locality: unknown (Australia in error) Type: unknown Type locality: plains of Kanauj district, Uttar Pradesh, India Holotype: BMNH 1946.8.14.44 Distribution: northwest India, eastern Pakistan, eastern Afghanistan

Uromastyx loricata (Blanford, 1874) Centrotrachelus loricatus Blanford, 1874: 660 Synonym Uromastix costatus Müller, 1885: 294 Type locality: unknown Holotype: NHMB 3491 Type locality: Bushire, Iran Holotype : BMNH 1946.8.11.59 Distribution: eastern Iraq; southwest Iran:Kurdestan-Kermanshah, Kuzestan-Lorestan, Fars

Uromastyx macfadyeni Parker, 1932 Uromastyx macfadyeni Parker, 1932: 353 Type locality: near Berbara, British Somaliland, Somalia Holotype: BMNH 1946.8.14.54 Distribution: Somalia: between Berbera and Heis on the Gulf of Aden; southern Eritrea; Djibouti

Uromastyx nigriventris Rothschild & Hartert, 1912 Uromastix acanthinurus nigriventris Rothschild & Hartert, 1912: 468 G Mtnez Synonyms Uromastyx acanthinura nigerrima Hartert, 1913:79 Type locality: southern Oued Mya, Algeria Holotype: BMNH (Tring), not located Uromastix acanthinurus werneri Müller, 1922: 201 Type locality: Ain Sefra, Oràn, Algeria Holotype: ZSM 436/0 Uromastix acanthinurus pluriscutata Fejérvary, 1927: 514 Type locality: Morocco Holotype: HMN 2813/1 Type locality: Tilrhempt, between Laghouat and Ghardaia, Algeria Holotype: BMNH 1969.2074 Distribution: Morocco: east and south of the Atlas Mountains; western Algeria: Sahara Atlas and in the regions northwest, northeast and southwest of the Great Western Erg

Uromastyx occidentalis Mateo, Geniez, Lopez-Jurado & Bons, 1999 Uromastyx occidentalis Mateo, Geniez, Lopez-Jurado & Bons, 1999: 106 Type locality: Aagtel Agmumuit (21° 52'N, 15° 31'W), between Yeloua and Mades, Adrar Souttouf, Western Sahara Holotype: DB.ULPGC-5 Distribution: Western Sahara

Uromastyx ocellata Lichtenstein, 1823 Uromastyx ocellata Lichtenstein, 1823: 107

D Melnikov Type locality: Nubia (=Sudan) Syntypes: ZMB 809-813 Distribution: Somalia: Borama district; Djibouti; Eritrea; eastern Ethiopia; Sudan; southeastern Egypt

Uromastyx ornata Heyden, 1827 Subspecies: Uromastyx ornata ornata Heyden, 1827 Uromastyx ornatus Heyden, 1827: 1

M Samuni Type locality: Al Muwaylih, Saudi Arabia Holotype: SMF 10403 Distribution: eastern Egypt: Sinai; Israel: Negev; west coast of Saudi Arabia

Uromastyx ornata philbyi Parker, 1938 Uromastyx philbyi Parker, 1938: 484 Type locality: between Makkah and Shabwa, southern Hejaz, between the mountains and Rub al Khali, Saudi Arabia Holotype: BMNH 1946.8.14.65 Distribution: western Saudi Arabia; northwestern Yemen

Uromastyx princeps O’Shaughnessy, 1880 Uromastix princeps O’Shaughnessy, 1880:445 Type locality: Candala, northeastern Somalia Holotype: BMNH 1946.814.56 Distribution: Somalia: Sanaag, Bari, Nogal and Mudug provinces; Ethiopia: Ogaden

Uromastyx shobraki Wilms & Schmitz, 2007 Uromastyx yemenensis shobraki Wilms & Schmitz, 2007:16 Type locality: Mafraq Mocca (Mafraq al-Mukha), km 13.5, Yemen Holotype: ZFMK 48681 Distribution: southwestern Yemen

Uromastyx thomasi Parker, 1930 Uromastix thomasi Parker, 1930: 595 Type locality: Bu Ju’ay, Rub al Khali, Dhofar, Oman Holotype: BMNH 1946.8.14.43 Distribution: southern Oman

Uromastyx yemenensis Wilms & Schmitz, 2007 Uromastyx yemenensis yemenensis Wilms & Schmitz, 2007: 12 Type locality: Abyan Governorate, vicinity of Lodar (= Lawdar),Yemen Holotype: ZFMK 47861 Distribution: eastern Yemen

Xenagama Boulenger, 1895 Boulenger, G.A. 1895. An account of the reptiles and batrachians collected by Dr. A. Donaldson Smith in western Somaliland and the Galla Country. Proceedings of the Zoological Society of London 1895: 530-540 Type species: Uromastix batilliferus Vaillant, 1882, by original designation

Xenagama batillifera (Vaillant, 1882) Uromastix batilliferus Vaillant, 1882: 10

Moscow Zoo Type locality: Bender-Meraya, Somalia Lectotype: MNHP 0.5830, designated by Wagner et al 2013 Distribution: northern Somalia: eastern Ethiopia

Xenagama taylori (Parker, 1935) Agama taylori Parker, 1935: 525

H Zell Type locality: Haud (44°49 ′ E., 8°43 ′N), Somalia Holotype: BMNH 1946.8.28.11 Distribution: northwestern Somalia; northeastern Ethiopia

Xenagama wilmsi Wagner, Mazuch & Bauer, 2013 Xenagama wilmsi Wagner, Mazuch & Bauer, 2013: 158 T Mazuch Type locality: ca. 42 km SE of (Ogaden) (09°09.353 ′N, 43°07.96 ′2E), Ethiopia Holotype: ZFMK 93255 Distribution: Ethiopia: central, and Ogaden; northern Somalia

Xenagama zonura (Boulenger, 1895) Agama zonura Boulenger, 1895: 533 Type locality: Shek Mohamed, near Delo ( 07°’ 15’ N, 40° 28’ E), near Ginir, Oromia Region, Ethiopia Holotype: BMNH 1946.8.28.10 Distribution: Ethiopia: central highlands

ACKNOWLEGEMENTS

This work could not have been completed without the help of numerous individuals. We thus acknowledge the help and advice of Patrick D Campbell (BMNH), Frank Tillack (ZMB ), Esther Dondorp (RMNH), Lemmy Mashinini (TMP), Nikolai Poyarkov (LUM),

The authors express their gratitude to the photographers who have contributed to this book, in particular, Stephen Zozaya, Mark O’Shea, Indraneil Das, Mohd Abdul Muin, Martin Pickersgill, S Harikishnan, Stewart Macdonald, Tom Charlton, Niranjan Sant, and many others.

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Taxon Index

A abbotti (see Gonocephalus ) abbreviatus (see Draco ) acanthinura (see Uromastyx ) acanthocephalus (see Monilesaurus ) achlios (see Lophocalotes ) aculeata (see Agama) acutirostris (see Aphaniotis ) acutirostris (see Phrynocephalus guttatus ) adelaidensis (see Ctenophorus ) adramitanus (see Acanthocercus ) adductus (see Diporiphora ) aegyptia (see Uromastyx ) aegyptius (see Trapelus mutabilis ) affinis (see Phrynocephalus przewalskii ) africana (see Agama) agama (see Agama) agilis (see Trapelus ) agnetae (see Trapelus ) agrorensis (see Laudakia ) ahvazicus (see Phrynocephalus ) albertisii (see Hypsilurus papuensis ) albilabris (see Diporiphora ) albolineatus (see Phrynocephalus ) alfredschmidti (see Uromastyx ) alpherakii (see Phrynocephalus guttatus ) altaica (see Laudakia stoliczkana ) alticola (see Phrynocephalus theobaldi ) alticristatus (see Calotes emma ) amarambalamensis (see Monilesaurus ellioti ) amboinensis (see Draco lineatus ) amboinensis (see Hydrosaurus ) ameliae (see Diporiphora ) amphiboluroides (see Diporiphora ) anamallayana (see Salea ) ananjevae (see Phrynocephalus ) anchietae (see Agama) andamanensis (see Pseudocalotes ) andersoniana (see Japalura ) angulifera (see Rankinia diemensis ) annamensis (see Leiolepis guttata ) annectens (see Acanthocercus ) aqabensis (see Pseudotrapelus ) arabicus (see Phrynocephalus ) aralensis (see Trapelus sanguinolentus ) arcellazzii (see Phrynocephalus) archiducissae (see ) arenaria (see Pseudotrapelus sinaitus ) armata (see Acanthosaura ) armata (see Agama) arnhemica (see Diporiphora ) asmussi (see Uromastyx ) aspera (see Trapelus mutabilis ) atra (see Agama) atricollis (see Acanthocercus ) attenboroughii (see Sitana ) auffenbergi (see Laudakia pakistanica ) aurantiacaudatus (Phrynocephalus mystaceus ) aurantolabium (see Microauris ) aurita (see Cryptagama ) auritus (see Hypsilurus ) austeniana (see Pseudocalotes ) australis (see Diporiphora ) australis (see Dendragama ) axillaris (see Phrynocephalus )

B bachae (see Calotes ) badakhshana (see Laudakia ) badius (see Ctenophorus maculatus ) bahiri (see Sitana ) baliomus (see Pseudocalotes ) bannikovi (see Phrynocephalus ) bapoensis (see Pseudocalotes kingdonwardi ) barbata (see Pogona ) batangensis (see Diploderma ) batillifera (see Xenagama ) beccarii (see Draco ) beccarii (see Harpesaurus ) beddomei (see Otocryptis ) bedrjagai (see Phrynocephalus ) bellii (see Gonocephalus ) bellii (see Leiolepis belliana ) belliana (see Leiolepis ) bengalensis (see Japalura variegata ) bennettii (see Diporiphora ) benti (see Uromastyx ) benueensis (see Agama doriae ) bergi (see Phrynocephalus guttatus ) beyschlagi (see Gonocephalus ) bhutanensis (see Calotes ) biaro (see Draco ) bilineata (see Diporiphora ) bilkewitschi (see Phrynocephalus raddei ) bimaculatus (see Draco ) binotatus (see Hypsilurus ) bintangensis (see Acanthosaura ) birulai (see Phrynocephalus przewalskii ) bivittata (see Otocryptis wiegmanni ) blanfordana (see Psammophilus ) blanfordi (see Phrynocephalus versicolor ) blanfordii (see Draco ) bochariensis (see Laudakia ) bocourti (see Agama) boehmei (see Leiolepis ) boehmei (see Trapelus ) boensis (see Agama) boettgeri (see Phrynocephalus raddei ) bogdanowi (see Phrynocephalus versicolor ) borneensis (see Harpesaurus ) borneensis (see Phoxophrys ) bornensis (see Gonocephalus ) borstschewskyi (see Laudakia lehmanni ) boschmai (see Draco ) bottegi (see Agama) boueti (see Agama) boulengeri (see Agama) boulengeri (see Dendragama ) boulengeri (see Phrynocephalus reticulatus ) bourouniensis (see Draco ) boydii (see Hypsilurus ) boylani (see Rankinia diemensis ) brachypoda (see Acanthosaura ) branchi (see Acanthocercus ) branchialis (see Intellagama lesueurii ) brachydactyla (see Laudakia stellio ) brachydactyla (see Pseudocalotes kakhienensis ) braueri (see Acanthosaura lepidogaster ) brevicauda (see Diporiphora bilineata ) brevicauda (see Diploderma ) brevicauda (see Trapelus agilis ) brevicaudus (see Coryphophylax ) brevipes (see Diploderma ) brevipes (see Phrynocephalus scutellatus ) brevipes (see Pseudocalotes ) brevis (see Pogona henrylawsoni ) brooksi (see Thaumatorhynchus ) bruijnii (see Hypsilurus ) burmana (see Bronchocela cristatella ) brunnea (see Chelosania ) burnsi (see Amphibolurus ) butlerorum (see Ctenophorus )

C caelaticeps (see Rankinia diemensis ) caerulhians (see Draco ) calotes (see Calotes ) cameranoi (see Phrynocephalus helioscopus ) capensis (see Acanthocercus atricollis ) capra (see Acanthosaura ) capreolatus (see Hypsilurus ) cardamomensis (see Acanthosaura ) carinatus (see Hypsilurus modestus ) carinatus (see Laudakia nupta ) carinatus (see Phrynocephalus przewalskii ) carinilabris (see Phrynocephalus przewalskii ) carinipes (see Phrynocephalus scutellatus ) cariniventris (see Agama mossambica ) carpentariensis (see Diporiphora ) castroviejoi (see Agama boueti ) caucasica (see Laudakia ) caudicinctus (see Ctenophorus ) caudicinctus (see Physignathus cocincinus ) caudivolvula (see Phrynocephalus guttatus ) caudospinosa (see Agama ) celebensis (see Bronchocela ) celebensis (see Hydrosaurus amboinensis ) centralis (see Amphibolurus ) centralis (see Tympanocryptis ) cephalum (see Phoxophrys ) cephalus (see Tympanocryptis ) ceylanica (see Cophotis ) ceylonensis (see Calotes ) chamaeleontinus (see Gonocephalus ) chapaensis (see Diploderma ) chapmani (see Ctenophorus ) chernovi (see Laudakia bochariensis ) chincollium (see Calotes ) chlodnickii (see Pseudotrapelus ) ciliaris (see Phrynocephalus versicolor ) citrinus (see Ctenophorus isolepis ) clamosa (see Lyriocephalus scutatus ) clarkorum (see Phrynocephalus ) clayi (see Ctenophorus ) cochinsinensis (see Physignathus cocincinus ) cocincinus (see Physignathus ) coggeri (see Diporiphora nobbi ) collicristatus (see Ptyctolaemus ) colonorum (see Agama agama ) condaminensis (see Tympanocryptis ) congica (see Agama agama ) convergens (see Diporiphora ) cordofanensis (see Agama doriae ) cordylina (see Laudakia stellio ) cornii (see Agama hartmanni ) cornutus (see Draco ) costatus (see Uromastyx loricata ) cristata (see Agama) cristatella (see Bronchocela ) cristatellus (see Draco ) cristatus (see Calotes versicolor ) cristatus (see Ctenophorus ) crucigera (see Acanthosaura ) cryptotis (see Draco maximus ) cuvieri (see Physignathus cocincinus ) cyanocephalus (see Acanthocercus ) cyanogaster (see Acanthocercus ) cyanolaemus (see Draco blanfordii ) cyanopterus (see Draco ) cybelidermus (see Pseudocalotes ) cypriaca (see Laudakia stellio )

D daani (see Laudakia stellio ) dagestanicus (see Phrynocephalus mystaceus ) danieli (see Bronchocela ) darlingtoni (see ) dasi (see Japalura ) daudinii (see Draco volans ) dayana (see Laudakia ) deccanensis (see Sitana ponticeriana ) decresii (see Ctenophorus ) denzeri (see Gonocephalus bornensis ) deserti (see Trapelus mutabilis ) desilvai (see Calotes ) devakai (see Sitana ) dhofarensis (see Pseudotrapelus ) diabolicus (see Tympanocryptis ) diemensis (see Rankinia ) dilophus (see Hypsilurus ) dioidema (see Dendragama ) dispar (see Uromastyx ) distanti (see Agama aculeata ) divergens (see Draco maculatus ) dodomae (see Agama lionotus ) donan (see Diploderma polygonata ) doriae (see Agama ) doriae (see Gonocephalus ) doriai (see Phrynocephalus versicolor ) dorsalis (see Psammophilus ) dringi (see Pseudocalotes ) drogon (see Pseudocalotes ) drukdaypo (see Diploderma ) dualis (see Ctenophorus maculatus ) dumbara (see Cophotis ) dussumieri (see Draco ) dymondi (see Diploderma )

E elegans (see Phrynocephalus przewalskii ) elgonis (see Agama lionotus ) ellioti (see Monilesaurus ) elliotti (see Japalura tricarinata) emma (see Calotes ) ensicauda (see Harpesaurus ) eraduensis (see Gowidon longirostris ) erdeleni (see Ceratophora ) ericae (see Phrynocephalus axillaris ) erythrogaster (see Laudakia ) erythrurus (see Phrynocephalus ) etoshae (see Agama ) euptilopus (see Phrynocephalus ) everetti (see Draco cyanopterus )

F farooqi (see Calotes versicolor ) fasciata (see Diploderma ) fasciata (see Lophognathus gilberti ) feae (see Pseudocalotes kakhienensis ) femoralis (see Ctenophorus ) fieldii (see Trapelus ruderatus ) fimbriatus (see Draco ) finchi (see Agama ) fionni (see Ctenophorus ) fitzsimonsi (see Agama kirkii ) flavicauda (see Laudakia nupta ) flaviceps (see Diploderma ) flavifasciata (see Uromastyx dispar ) flavigula (see Pseudocalotes ) flavimaculatus (see Trapelus ) floweri (see ) floweri (see Pseudocalotes ) fordi (see Ctenophorus ) formosensis (see Diploderma swinhonis ) formosus (see Draco ) forsythia (see Phrynocephalus ) fortescuensis (see Tympanocryptis ) frontalis (see Phrynocephalus ) fruhstorferi (see Acanthosaura lepidogaster ) fusca (see Aphaniotis ) fusca (see Laudakia nupta ) fusca (see Sitana ) fuscus (see Draco volans )

G galli (see Phrynocephalus mystaceus ) geckoides (see Phrynocephalus vlangalii ) geelvinkianus (see Hypsilurus ) geyri (see Uromastyx ) gibba (see Ctenophorus ) gibbosus (see Moloch horridus ) gigantea (see Gonocephalus chamaeleontinus ) gigas (see Calotes versicolor ) gigas (see Tympanocryptis ) gilberti (see Lophognathus ) godeffroyi (see Hypsilurus ) gokakensis (see Sitana ) golubewii ( see Phrynocephalus ) graafi (see Ctenophorus caudicinctus ) gracilimembris (see Agama ) gracilipes (see ) gracilis (see Draco cornutus ) gracilis (see Diporiphora ) grahami (see Diploderma ) grandis (see Draco fimbriatus ) grandis (see Gonocephalus ) grandisquamis (see Calotes ) granulifera (see Diporiphora ) gregorii (see Acanthocercus ) griseus (see Ctenophorus maculatus) griseus (see Uromastyx hardwicki ) grombtschewskii (see Phrynocephalus vlangalii ) grumgrzimailoi (see Phrynocephalus ) guentheri (see Draco ) guentheri (see Phrynocephalus versicolor ) guentherpetersi (see Acanthocercus ) guentherpetersi (see Leiolepis ) guinanensis (see Phrynocephalus ) gularis (see Ctenophorus isolepis ) gularis (see Ptyctolaemus ) gularis (see Salea ) gularis (see Trapelus mutabilis ) guttalineatus (see Pseudocalotes ) guttata (see Leiolepis ) guttatus (see Phrynocephalus ) gutturosa (see Bronchocela cristatella )

H haasei (see Draco maculatus ) haasi (see Trapelus agnetae ) haeckeli (see Phrynocephalus ) haematopogon (see Draco ) hainanensis (see Acanthosaura lepidogaster ) hamptoni (see Diploderma ) harbai (see Uromastyx aegyptia ) hardwicki (see Uromastyx ) hartmanni (see Agama ) hayeki (see Bronchocela ) helioscopus (see Phrynocephalus ) hennigi (see Draco fimbriatus ) henrylawsoni (see Pogona ) herveyi (see Gonocephalus liogaster ) heterurus (see Intellagama lesueurii ) hikidanus (see Hypsilurus ) himalayana (see Laudakia ) hirsutus (see Salea horsfieldii ) hispida (see Agama ) hispidus (see Phrynocephalus versicolor ) hodhensis (see Uromastyx dispar ) holsti (see ) hongyuanensis (see Phrynocephalus ) horneri (see Lophognathus ) horrescens (see Acanthosaura crucigera ) horridus (see Moloch ) horsfieldii (see Salea ) horváthi (see Phrynocephalus ) houstoni (see Tympanocryptis ) howittii (see Intellagama lesueurii ) htunwini (see Calotes ) hulbertorum (see Agama ) humei (see Coryphophylax subcristatus )

I

idae (see Phrynocephalus axillaris ) iadina (see Diploderma ) imbricatus (see Ctenophorus caudicinctus ) immaculatus (see Phrynocephalus versicolor ) impalearis (see Agama ) incerta (see Phrynocephalus guttatus ) incognitus (see Lophognathus gilberti ) indica (see Monilesaurus ellioti ) indica (see Calotes versicolor ) indica (see Salea anamallayana ) indicus (see ) indochinensis (see Draco ) inermis (see Ctenophorus nuchalis ) inermis (see Trapelus mutabilis ) infans (see Ctenophorus caudicinctus ) infralineata (see Agama aculeata ) inornatus (see Hypsilurus modestus ) insularis (see Agama ) intermedia (see Bronchocela cristatella ) intermedius (see Bronchocela jubata ) intermedius (see Draco maximus ) interruptus (see Gonocephalus ) interscapularis (see Phrynocephalus ) intima (see Tympanocryptis ) irawadi (see Calotes ) ishigakiensis (see Diploderma polygonata ) iskandari (see Draco ) isolepis (see Ctenophorus ) isolepis (see Trapelus agilis ) isozona (see Laudakia bochariensis ) isseli (see Phrynocephalus)

J jacksoniensis (see Amphibolurus muricata ) jareckii (see Draco ) jayakari (see Trapelus ) javanica (see Draco volans ) jensvindumi (see Pseudotrapelus ) jerdoni (see Calotes ) jubata (see Bronchocela ) jugularis (see Diporiphora )

K kaimosae (see Agama ) kakhienensis (see Pseudocalotes ) kalesari (see Sitana ) kalmykus (see Phrynocephalus guttatus ) karu (see Ceratophora ) karumba (see Tympanocryptis lineata ) kaulbacki (see Pseudocalotes kingdonwardi ) kelaarti (see Calotes ceylonensis ) khani (see Laudakia pakistanica ) khaonanensis (see Pseudocalotes ) khorasanus (see Phrynocephalus mystaceus ) khuzistanensis (see Trapelus agilis ) kinabaluensis (see Hypsicalotes ) kingdonwardi (see Pseudocalotes ) kingii (see Chlamydosaurus ) kirkii (see Agama ) kirmanensis (see Trapelus agilis ) kiwuensis (see Acanthocercus ) klossi (see Gonocephalus ) klemenzi (see Phrynocephalus axillaris ) knobeli (see Agama ) kontumensis (see Pseudocophotis ) kozlovi (see Phrynocephalus axillaris ) kuhlii (see Gonocephalus ) kulagini (see Phrynocephalus versicolor ) kumaonensis (see Japalura ) kuschakewitschi (see Phrynocephalus guttatus )

L labialis (see Tropicagama temporalis ) lacunosus (see Gonocephalus ) laetepictus (see Draco obscurus ) laevis (see Ctenophorus reticulatus ) laeviventris (see Diploderma ) lalliae (see Diporiphora ) lamnidentata (see Acanthosaura lepidogaster ) lanzai (see Agama ) larutensis (see Pseudocalotes ) latastii (see Trapelus mutabilis ) lateralis (see Tropicagama temporalis ) laungwalaensis (see Bufoniceps ) lebretoni (see Agama ) lehmanni (see Laudakia ) lepidogaster (see Acanthosaura ) leptieni (see Uromastyx aegyptia ) lessonae (see Trapelus ruderatus ) lesueurii (see Intellagama ) leucerythrolaema (see Agama finchi ) leucostygma (see Trapelus mutabilis ) levis (see Phrynocephalus versicolor ) lidskii (see Phrynocephalus vlangalii ) lineata (see Calotes calotes) lineata (see Tympanocryptis ) lineatus (see Draco ) linga (see Diporiphora ) liocephalus (see Calotes ) liogaster (see Gonocephalus ) liolepis (see Calotes ) lionotus (see Agama ) liratus (see Laudakia melanura ) lobata (see Hypsilurus modestus ) longibarba (see Draco quinquefasciatus ) longicauda (see Hypsilurus papuensis ) longicaudatus (see Phrynocephalus ) longii (see Hypsilurus ) longirostris (see Gowidon ) lophura (see Hydrosaurus amboinensis ) lophyrus (see Gonocephalus chamaeleontinus ) loricata (see Trapelus mutabilis ) loricata (see Uromastyx ) loveridgei (see Acanthocercus gregorii ) lucyae (see Agama ) luei (see Diploderma ) ludekingi (see Lophocalotes ) ludovici (see Phrynocephalus axillaris ) lutensis (see Phrynocephalus ) luteoguttatus (see Phrynocephalus )

M macfadyeni (see Uromastyx ) macgregorii (see Lyriocephalus scutatus ) macra (see Tympanocryptis lineata ) macropus (see Ctenophorus caudicinctus ) macrolepis (see Diporiphora bilineata ) macrolepis (see Hypsilurus ) macropeltis (see Phrynocephalus ) maculata (see Phrynocephalus axillaris ) maculatus (see Ctenophorus ) maculatus (see Draco ) maculatus (see Phrynocephalus ) maculiferus (see Intellagama lesueurii ) maculilabris (see Lophognathus ) maculosus (see Ctenophorus ) magna (see Diporiphora ) magnus (see Hypsilurus ) major (see Ctenophorus nuchalis ) major (see Draco volans ) major (see Japalura ) makarikarika (see Agama hispida ) makii (see Diploderma ) maliensis (see Uromastyx dispar ) manamendrai (see Calotes ) margaretae (see Diporiphora ) margaritaceus (see Lyriocephalus scutatus ) maria (see Calotes ) marmorata (see Bronchocela ) maximiliani (see Coryphophylax subcristatus ) maximus (see Draco ) mccartneyi (see Tympanocryptis ) mckenziei (see Ctenophorus ) medogensis (see Calotes ) megalepis (see Gonocephalus ) megalonyx (see Trapelus ) megapogon (see Hypsilurus dilophus ) mehelyi (see Agama anchietae ) melanopogon (see Draco ) melanura (see Laudakia ) melanurus (see Phrynocephalus guttatus ) mensarum (see Ctenophorus caudicinctus ) mentager (see Physignathus cocincinus ) methueni (see Agama anchietae ) micangshanensis (see Diploderma ) microlepidotus (see Pogona ) microlepis (see Draco haematopogon ) microlepis (see Japalura variegata ) microlepis (see Laudakia ) microlepis (see Pseudocalotes ) microlepis (see Uromastyx aegyptia ) microlophus (see Hydrosaurus amboinensis ) micropholis (see Agama atra ) microterolepis (see Agama atra ) microtympanum (see Trapelus ) mindanensis (see Draco ) minima (see Pogona ) minor (see Brachysaura ) minor (see Draco volans ) minor (see Pogona ) minor (see Sitana ponticeriana ) minutus (see Acanthocercus ) miotympanum (see Gonocephalus bornensis ) mirrityana (see Ctenophorus ) mitchelli (see Pogona ) mitsukurii (see Diploderma swinhonis ) miyakensis (see Diploderma polygonata ) mjobergi (see Gonocephalus ) modestus (see Ctenophorus decresii ) modestus (see Hypsilurus ) modiglianii (see Draco ) modiglianii (see Harpesaurus ) moltschanowi (see Phrynocephalus guttatus ) moluccana (see Bronchocela cristatella ) montana (see Agama ) montanus (see Monilesaurus ) mossambica (see Agama ) mucosoensis (see Agama ) mucronata (see Laudakia erythrogaster ) mucronata (see Sitana ponticeriana ) muricata (see Amphibolurus ) murielis (Phrynocephalus axillaris ) murphyi (see Acanthosaura ) mutabilis (see Trapelus ) mutabilis (see Uromastyx acanthinura ) mwanzae (see Agama ) mykonensis (see Laudakia stellio ) mystaceus (see Calotes ) mystaceus (Phrynocephalus )

N namaquensis (see Agama anchietae ) nanschanica (see Phrynocephalus ) nasatus (see Phrynocephalus axillaris ) nasiri (see Laudakia melanura ) nasuta (see Aphaniotis ornata ) nataliae (see Acanthosaura ) nejdensis (see Phrynocephalus ) nemoricola (see Calotes ) neumanni (see Pseudotrapelus ) ngoclinensis (see Japalura ) ngovantrii (see Leiolepis ) nguyarna (see Ctenophorus ) niger (see Japalura variegata ) nigerrima (see Uromastyx nigriventris ) nigriappendiculatus (see Draco melanopogon ) nigricans (see Phrynocephalus helioscopus ) nigricollis (see Acanthocercus atricollis ) nigricollis (see ) nigrigularis (see Complicitus ) nigrigularis (see Hypsilurus ) nigrilabris (see Calotes ) nigrilabris (see Phoxophrys) nigristigma (see Otocryptis ) nigriventris (see Uromastyx ) nigrofasciata (see Trapelus mutabilis ) nikolskii (see Phrynocephalus reticulatus ) nobbi (see Diporiphora ) norrisi (see Amphibolurus ) norvillii (see Draco ) nuchalis (see Ctenophorus ) nuchalis (see Diporiphora australis ) nullarbor (see Pogona ) nupta (see Laudakia ) nurgeldievi (see Laudakia erythrogaster ) nuristanica (see Laudakia )

O osbornei (see Tympanocryptis ) obscura (see Uromastyx dispar ) obscurus (see Draco ) ocellata (see Leiolepis belliana ) ocellata (see Phrynocephalus guttatus ) ocellata (see Phrynocephalus reticulatus ) occidentalis (see Agama ruppelli ) occidentalis (see Uromastyx ) ochropterus (see Draco lineatus ) octospinosus (see Bronchocela cristatella ) ophiomachus (see Calotes calotes ) orientalis (see Phrynocephalus helioscopus ) orlovi (see Bronchocela ) ornata (see Aphaniotis ) ornata (see Brachysaura minor ) ornata (see Diporiphora australis ) ornata (see Uromastyx ) ornatus (see Ctenophorus ) ornatus (see Draco ) ornatus (see Hypsilurus ) ornatus (see Phrynocephalus ) otai (see Cristidorsa )

P pakistanensis (see Trapelus agilis ) pakistanica (see Laudakia ) palawanensis (see Draco ) pallida (see Laudakia erythrogaster ) pallida (see Diporiphora ) pallidus (see Trapelus mutabilis ) papenfussi (see Laudakia ) papuensis (see Hypsilurus ) paraconvergens (see Diporiphora ) parafricana (see Agama ) paragama (see Agama ) paraskiwi (see Phrynocephalus guttatus ) parva (see Phrynocephalus vlangalii ) parviceps (see Ctenophorus ) parvulus (see Phrynocephalus przewalskii ) paulus (see Calotes ) peguensis (see Leiolepis ) pentalineata (see Diporiphora bilineata ) pentalineata (see Tympanocryptis ) perplexa (see Diporiphora ) persica (see Trapelus ruderatus ) persicus (see Laudakia caucasica ) persicus (see Phrynocephalus ) persimilis (see Agama ) pethiyagodai (see Calotes ) pewzowi (see Phrynocephalus versicolor ) phaeospinosa (see Diporiphora ) philbyi (see Uromastyx ornata ) philippinus (see Bronchocela marmorata ) phillipsii (see Acanthocercus ) phongdienensis (see Acanthosaura ) phuwuanensis (see Mantheyus ) picticauda (see Agama) pictus (see Ctenophorus ) picea (see Laudakia stellio ) pindan (see Diporiphora ) pinguicolla (see Tympanocryptis ) planiceps (see Agama ) planidorsata (see Cristidorsa ) platyceps (see Calotes jerdoni ) pluriscutata (see Uromastyx nigriventris ) poilani (see Pseudocalotes ) polygonata (see Diploderma ) ponticeriana (see Sitana ) popei (see Diploderma swinhonis ) poppeki (see Trapelus mutabilis ) potanini (see Phrynocephalus versicolor ) praepos (see Draco volans ) princeps (see Uromastyx ) przewalskii (see Phrynocephalus ) pseudopsephos (see Tympanocryptis ) pulchella (see Agama aculeata ) punctatus (see Draco fimbriatus ) pustulatus (see Hydrosaurus ) pylzowi (see Phrynocephalus vlangalii )

Q quadrasi (see Draco ) quattuorfasciatus (see Gowidon longirostris ) quinquefasciatus (see Draco ) quinquicarinata (see Pseudocalotes brevipes )

R raddei (see Phrynocephalus ) rayaensis (see Bronchocela ) reginae (see Diporiphora ) reinwardti (see Draco lineatus ) reldoe (see Phrynocephalus vlangalii ) relictus (see Phrynocephalus rossikowi ) reticulata (see Laudakia caucasica ) reticulatus (see Ctenophorus ) reticulatus (see Draco ) reticulatus (see Phrynocephalus ) reticulatus (see Uromastyx hardwicki ) reveesii (see Leiolepis ) rhaegal (see Pseudocalotes ) rhammanotus (see Pseudocalotes ) rhytisma (see Draco ) rizali (see Draco guentheri ) robecchii (see Acanthocercus ) robecchii (see Agama ) robinsoni (see Phoxophrys tuberculata ) robinsonii (see Malayodracon ) rosaceum (see Dendragama boulengeri ) rossikowi (see Phrynocephalus ) rostralis (see Phrynocephalus versicolor ) rostratus (see Draco spilopterus ) rouxii (see Monilesaurus ) rubens (see Ctenophorus ) rubrigularis (see Bronchocela ) rubrigularis (see Trapelus ) rubritaeniata (see Leiolepis ) ruderatus (see Trapelus ) rudis (see Agama atra ) rufescens (see Ctenophorus ) ruppelli (see Agama )

S sacra (see Laudakia ) sagittifera (see Japalura ) saidalievi (see Phrynocephalus helioscopus ) sakoi (see Phrynocephalus) salehi (see Laudakia stellio ) salenskyi (see Phrynocephalus ) saleoides (see Calotes ceylonensis ) salinarum (see Ctenophorus ) salsatus (see Phrynocephalus guttatus ) samaoensis (see Draco volans ) sanchezi (see Bronchocela marmorata ) sanguinolentus (see Trapelus ) sankaranica (see Agama ) saposhnikovi (see Phrynocephalus helioscopus ) saravacensis (see Pseudocalotes ) savattieri (see Agama africana ) savignii (see Trapelus ) schleichi (see Sitana ) schmitzi (see Trapelus ) schneideri (see Dendragama boulengeri ) schoedei (see Hypsilurus ) schultzewestrumi (see Hypsilurus ) scutatus (see Lyriocephalus ) scutellatus (see Phrynocephalus ) scutulatus (see Ctenophorus ) semperi (see Gonocephalus ) sennariensis (see Agama doriae ) septentrionalis (see Agama ruppelli ) shammakowi (see Phrynocephalus rossikowi ) shenlong (see Bronchocela ) shobraki (see Uromastyx ) siebenrocki (see Phrynocephalus versicolor ) similis (see Uromastyx hardwicki ) simonyi (see Uromastyx benti ) sinaitus (see Pseudotrapelus ) sivalensis (see Sitana ) slateri (see Ctenophorus caudicinctus ) slowinskii (see Diploderm a) smaragdina (see Bronchocela ) smithii (see Agama ) sobria (see Diporiphora ) sogdianus (see Phrynocephalus interscapularis ) somalica (see Agama ) sophiae (see Gonocephalus ) spawlsi (see Agama caudospinosa ) spilonotus (see Draco ) spilopterus (see Draco ) spiniceps (see Phoxophrys ) spinipes (see Hypsilurus ) spinipes (see Uromastyx aegyptia ) spiniventris (see Phrynocephalus maculatus ) spinodomus (see Ctenophorus ) spinosa (see Agama ) spinosus (see Bronchocela marmorata ) splendida (see Diploderma ) steindachneri (see Phrynocephalus przewalskii ) stellio (see Laudakia ) stoddartii (see Ceratophora ) stolickai (see Phrynocephalus theobaldi ) stoliczkanus (see Laudakia ) straminea (see Trapelus mutabilis ) strauchi (see Phrynocephalus ) subcristatus (see Coryphophylax ) sulcatus (see Salea horsfieldii ) sumatrana (see Draco ) sumatrana (see Pseudocophotis ) sumatranus (see Gonocephalus chamaeleontinus ) superba (see Diporiphora ) supriatnai (see Draco ) suschkini (see Phrynocephalus guttatus ) suschkinianus (see Phrynocephalus przewalskii ) swinhonis (see Diploderma ) swild (see Diploderma ) sylvana (see Agama ) szechwanensis (see Diploderma fasciata )

T taeniopterus (see Draco ) tarimensis (see Laudakia stoliczkana ) tassiliensis (see Agama ) taylori (see Xenagama ) temporalis (see Tropicagama ) tennentii (see Ceratophora ) tenuicephalus (see Hypsilurus ) tetraporophora (see Tympanocryptis ) theobaldi (see Phrynocephalus ) thescelorhinos (see Harpesaurus borneensis ) thomasi (see Uromastyx ) thondalu (see Sitana ) tibialis (see ) tickelii (see Phrynocephalus scutellatus ) tiedemanni (see Calotes versicolor ) tigrina (see Gonocephalus chamaeleontinus ) timorensis (see Draco ) titiwangsaensis (see Acanthosaura ) tjantjalka (see Ctenophorus ) tongbiguanensis (see Acanthosaura ) tournevillei (see Trapelus ) toxopei (see Draco bourouniensis ) trachypleura (see Acanthocercus phillipsii ) triangularis (see Salea anamallayana ) triannulatus (see Laudakia microlepis ) tricarinata (see Japalura ) tricinctus (see Harpesaurus ) triploida (see Leiolepis ) trispinosus (see Salea horsfieldii ) tuberculata (see Laudakia ) tuberculata (see Phoxophrys ) tuberculatus (see Gonocephalus megalepis ) turcomanus (see Phrynocephalus ) turuensis (see Agama ) tympanistriga (see Pseudocalotes )

U ufipae (see Agama lionotus ) ugandaensis (see Acanthocercus ) uniformis (see Tympanocryptis ) uralensis (see Phrynocephalus helioscopus ) usambarae (see Agama lionotus )

V vadnappa (see Ctenophorus ) valens (see Diporiphora ) varcoae (see Diploderma ) variegata (see Japalura ) varius (see Phrynocephalus helioscopus ) vela (see Diploderma ) versicolor (see Calotes ) versicolor (see Phrynocephalus ) vescus (see Diporiphora ) vietnamensis (see Bronchocela ) vindumi (see Phrynocephalus ) viridis (see Calotes versicolor ) viridis (see Draco volans ) viserion (see Pseudocalotes ) vitticeps (see Pogona ) vlangalii (see Phrynocephalus ) volans (see Draco ) vulgaris (see Laudakia stellio ) vultuosa (see Calotes versicolor )

W wagneri (see Agama ) walker (see Draco ) weberi (see Hydrosaurus ) websteri (see Ctenophorus scutulatus ) weidholzi (see Agama ) werneri (see Pseudotrapelus sinaitus ) werneri (see Uromastyx nigriventris ) whiteheadi (see Draco maculatus ) wiegmanni (see Otocryptis ) wilmsi (see Xenagama ) wilsoni (see Tympanocryptis ) winneckei (see Diporiphora ) wui (see Laudakia )

X xanthostoma (see Diploderma polygonata )

Y yemenensis (see Uromastyx ) yemensis (see Acanthocercus ) yinnietharra (see Ctenophorus ) yulongensis (see Diploderma ) yunnanensis (see Calotes jerdoni ) yunnanensis (see Diploderma )

Z zardunyi (see Phrynocephalus raddei ) zetangensis (see Phrynocephalus theobaldi ) zhaoermii (see Diploderma ) ziegleri (see Pseudocalotes ) zolaiking (see Calotes ) zonura (see Xenagama )