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Msea9catalogocarabidae.Pdf MONOGRAFÍAS S.E.A. — vol. 9 Primera Edición: Zaragoza, 15 Septiembre, 2003. Título: Catálogo de los Carabidae (Coleoptera) de la Península Ibérica / Catalogue of the Carabidae (Coleoptera) of the Iberian Peninsula. Autor: José Serrano. Edita: Sociedad Entomológica Aragonesa (SEA). Avda. Radio Juventud, 37 50012 – Zaragoza (España) Director de publicaciones: A. Melic [email protected] http://entomologia.rediris.es/sea Maquetación y Diseño: A. Melic Portada: Trechus apoduvalipenis Salgado y Ortuño 1998. Autor: Vicente M. Ortuño Imprime: Gorfi, S.A. c/.Menéndez Pelayo, 4 50009 – Zaragoza (España) I.S.B.N.: 84 – 932807– 2 – 0 Depósito Legal: Z – 1789 – 94 © El autor (por la obra). © SEA (por la edición). Queda prohibida la reproducción total o parcial del presente volumen, o de cualquiera de sus partes, por cualquier medio, sin el previo y expreso consentimiento por escrito del autor y editora. Publicación gratuita para socios SEA (ejercicio 2003). Precio de venta al público: 18 euros (IVA incluido). Gastos de envío no incluidos. Solicitudes: SEA. Catálogo de los Carabidae (Coleoptera) de la Península Ibérica Catalogue of the Carabidae (Coleoptera) of the Iberian Peninsula José Serrano MONOGRAFÍAS SEA, vol. 9 ZARAGOZA, 2003 a Dedicatoria A mi mujer, Bárbara y a mis hijos José Enrique y Antonio. Por todo. Índice Prólogo ......................................................................... 5 Agradecimiento................................................................... 6 Notas a las distintas partes de la obra................................................... 7 El catálogo...................................................................... 11 Cambio nomenclatural ............................................................ 82 Las novedades................................................................... 83 Relación sintética de la Sistemática empleada y estadísticas del catálogo....................... 85 Los mapas de las regiones naturales de la Península Ibérica................................. 91 La bibliografía ................................................................... 93 Índice taxonómico............................................................... 107 Index Foreword........................................................................ 5 Acknowledgements................................................................ 6 Notes to the the chapters of this work .................................................. 7 The catalogue ................................................................... 11 Proposal of nomenclatural change.................................................... 82 New taxa included in the catalogue................................................... 83 A synthesis of the Systematics and the number of taxa..................................... 85 The maps....................................................................... 91 The references................................................................... 93 Taxonomical index .............................................................. 107 4 José Serrano Catálogo de los Carabidae (Coleoptera) de la Península Ibérica 5 Prólogo Foreword La idea de escribir este catálogo obedece a dos moti- The idea of writing this book is due to two main vos principales. El primero de ellos es la publicación reasons. The first is the publication of the book Nomi- del Nomina Carabidarum (Lorenz, 1998), que facilita na Carabidarum (Lorenz, 1998), that makes easy the notablemente el disponer de una nomenclatura revisa- use of an updated nomenclature for the family. This is da de la familia. Este es posiblemente el aspecto menos possibly the aspect less elaborated of the excellent elaborado del excelente catálogo de Zaballos y Jeanne catalogue of Zaballos and Jeanne (1994). This task has (1994). Esta tarea de revisión de la nomenclatura se ha been much helped in the last moments by the appea- visto complementada a última hora con la publicación ring of a new Palaearctic catalogue (Löbl and Smetana, del catálogo paleártico (Löbl y Smetana, 2003). El 2003). The second reason is the remarkable increase of segundo motivo es el incremento manifiesto de nuevos new taxa for the Iberian carabid fauna during the last taxones que ha experimentado la fauna peninsular en years. The new works dealing with the fauna of los últimos años, lo que unido al estudio de la fauna de different Iberian regions (Galicia, Álava and the regiones concretas de la Península (Galicia, País Basque Country, the Montes de Toledo in central Vasco, Montes de Toledo, las sierras béticas, etc.) y la Spain, the north-eastern Betic mountains, etc.), and the aparición de revisiones de taxones peninsulares y publication of revisions and monographs concerning extrapeninsulares (tribus Carabini, Dyschiriini, Platy- Iberian and extra Iberian taxa (tribes Carabini and derus, Zabrus, etc.), suponen un volumen importante Dyschiriini, the genera Platyderus, Zabrus, etc.), make de nueva información. up altogether an important volume of recent informa- Para llevar a cabo esta labor he contado con la tion. For accomplishing with this work I have received ayuda generosa de numerosos colegas, que han atendi- the generous help of many colleagues, who have do solícitos las peticiones de sus publicaciones o la attended promptly my requests of recent papers or old fotocopia de obras antiguas. En todo momento he publications. tenido presente que esta es una obra abierta, inconclu- I have tried to reduce the critical comments to a sa por su propia naturaleza, y que será mejorada en los minimum, and thus have usually followed the criteria próximos años. Muchas propuestas sobre el estado of Lorenz, Löbl and Smetana, Turin et al. (the genus taxonómico de algunas especies y subespecies, así Carabus), Fedorenko (the tribe Dyschiriini), etc. como sobre su distribución deben ser entendidas como However, I have introduced some changes arising form hipótesis de trabajo. Por ello figuran algunas modifica- my own data or inferences. In any case, I realise that a ciones con respecto a la nomenclatura propuesta por catalogue is an open and endless work by its own Lorenz (1998), Löbl y Smetana (2003), o por otros nature, that should be improved in forthcoming years. autores, como ocurre con los Carabus peninsulares Some proposals about the taxonomic status and con respecto a la monografía de Turin et al. (2003). En distribution of particular species and subspecies, must la medida en que tales propuestas generen investiga- be understood as working hypotheses that should be ciones fructíferas (en vez de polémicas estériles de corroborated in the future. If these hypotheses generate salón), me sentiré satisfecho del esfuerzo realizado en productive research instead of sterile discussion, I pro de la comunidad de entomólogos que se interesan shall be satisfied of having dedicated many efforts for o trabajan con los Carabidae ibéricos. the benefit of the entomologist’s community interested Este libro no hubiera sido posible sin el esfuerzo in the Iberian Carabidae. de Claude Jeanne y Juan Pérez Zaballos, los autores de This work would have not been possible without los catálogos anteriores a éste (1986, 1994). De hecho, the efforts of Claude Jeanne and Juan Pérez Zaballos, el objetivo fundamental que me ha guiado durante su the authors of the former catalogues (1986, 1994). In elaboración, ha sido dar continuidad, de forma digna fact, the aim that has guided me during the realisation y eficaz, a los logros manifiestos de los catálogos of the book has been to continue, in a properly and elaborados por estos dos grandes amigos y colegas. efficient way, the goals attained by these two great Finalmente, agradecería a todos los lectores que me friends and colleagues with their past two catalogues. fueran indicando los errores y omisiones (siempre Finally, I acknowledge in advance to all the involuntarios) que adviertan. Quién sabe, a lo mejor colleagues the indications of mistakes and omissions dentro de diez años hay ganas y motivos para empezar that they may here find. Who knows, perhaps in next con otro nuevo catálogo... years I have again reasons and -most importantly- wishes for starting with a new catalogue... Murcia, 15 de julio de 2003 Prof. José Serrano Departamento de Zoología, Facultad de Veterinaria Universidad de Murcia Aptdo. 4021. 30071 Murcia (España) [email protected] 6 José Serrano Agradecimiento Acknowledgements Para la realización de esta obra he contado con la I have been helped by many colleagues to whom I ayuda imprescindible de numerosos colegas, que me greatly indebted. They have sent their publications or han facilitado publicaciones o información sobre las bibliographic information that are essential for upda- mismas, de cara a la actualización de los datos de ting the taxonomic and the faunistic records. These taxonomía y distribución. Entre ellos están Miguel colleagues are Miguel Ángel Alonso Zarazaga, Anto- Ángel Alonso Zarazaga, Antonio Andújar, Thorsten nio Andújar, Thorsten Assmann, George E. Ball, Assmann, George E. Ball, Pietro y Tulia Brandmayr, Pietro and Tulia Brandmayr, Emilio Carabajal, Ana Emilio Carabajal, Ana Cárdenas, Fabio Cassola, Juan Cárdenas, Fabio Cassola, Juan de Ferrer, Olegario del de Ferrer, Olegario del Junco, Oleguer Escolà, José Junco, Oleguer Escolà, José Galián, José García Galián, José García Carrillo, Carles Hernando, Fritz Carrillo, Carles Hernando, Fritz Hieke, Bernd Jaeger, Hieke, Bernd Jaeger, Claude Jeanne, Boris M. Kata- Claude Jeanne, Boris M. Kataev, José Luís Lencina,
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