16 17

Aflexia rubranura is a specialist on prairie dropseed; Erythroneura carbonata is a specialist on sea- but despite being completely flightless it is found on milkwort, a primrose relative that carpets saline alvars on islands in Lake Huron as well as in eastern depressions. prairies from Manitoba to northern Illinois.

Memnonia consobrina, found in arid grasslands in Memnonia consobrina western Canada, has subterranean nymphs. female

Neohecalus magnificus, found in Canada only in the Neohecalus magnificus Red River valley and at Ojibway Prairie in Windsor, female Ontario, has very different males and females.

Stirellus bicolor – in the United States this has bicolor two forms: in spring it is brown and in high summer (spring form) it is iridescent; but in Canada the two forms may be taken together.

Arthropods of Canadian Grasslands Number 10, 2004 16 17 and their relatives (Homoptera, ) from the Canadian Great Plains

K.G. Andrew Hamilton Biodiversity, Agriculture & Agri-Food Canada, CEF Ottawa Ont. K1A 0C6, [email protected]

The Canadian Great Plains are often referred to as “prairies”. This term is French for “meadows” and properly describes only the eastern plains grass- lands, particularly those around the southern end of Lake Manitoba or “Lac des Prairies.” There, prairie grasses are lush even in the driest part of the sum- mer, yet seldom if ever attain the height of “tall- grass” prairies farther south. The Auchenorrhyncha fauna or “short-horned bugs” of this region of Aphelonema simplex (Caliscelidae) is one of the most Manitoba is unique (Hamilton 1999). Just how common and distinctive bugs of the Canadian plains, unique, and how extensive this fauna is, has nev- the male being bright orange, and the head unusually er been elucidated. Recent sampling suggests that short and broad. Such bugs are called “piglet bugs” it extends all the way into eastern Saskatchewan! because their tiny wings expose their very robust bodies, and they often have a short “snout” on the head. A preliminary analysis of this fauna generates oth- er surprising results. For example, the biodiversity (east vs. west, north vs. south, regional endemism “hot spot” on the Canadian Great Plains appears vs. migration from southern localities) can be deter- to be Elbow, Saskatchewan which is at the head mined easily. of Lake Diefenbaker behind the Gardiner and Qu- This synthesis is based on nine surveys across Appelle dams. the Canadian Great Plains that were made since I was a student and began collecting leafhoppers at Winnipeg in 1965. The initial work was greatly as- sisted by the catches from mosquito traps main- tained at the University of Manitoba and at Glenlea Research Station (Hamilton 1972). This work has been supplemented by studies on other northern grasslands since I first found a Aprairie@ leafhop- per on a heathland near Manitoulin Island, Ontario Laccocera () are western with in 1975. brightly coloured males, but their females are ivory without obvious markings. What, exactly, is an endemic Aprairie@ bug? It is one that feeds only on a grassland-associated plant Short-horned bugs include a large number of species endemic to the Great Plains. The follow- ing account of the Canadian fauna is presented as a species list. An ongoing analysis of detailed bio- geographic patterns and their implications for un- derstanding the ecology of the Great Plains requires a great deal of analysis and will be published as sep- arate papers. The listing presented here gives an Bruchomorpha jocosa (Caliscelidae) is a distinctive overview by grouping the sites by latitude and lon- inhabitant of eastern prairies, one of many species gitude so that the most obvious distribution patterns that specialize on little bluestem.

Arthropods of Canadian Grasslands Number 10, 2004 18 19 or is rarely, if ever, taken far from a native prairie. The definition must be expanded somewhat to in- clude the insects feeding on plants that are large- ly associated with the prairies, but by virtue of their preference for moist habitats, find their way along water courses a considerable distance from the Great Plains (sandbar or Awolf@ willow and certain diminutive spike-rushes are good examples). Oliarus (Cixiidae) are externally rather undistinguished bugs, but they have perhaps the most elaborate male genitalia of any in the world. (MB), where the prairie is much more localized and fragmented, there are 6.7 sites per quadrat. The maximum number is 17 sites in quadrat 2 of south- western Manitoba (between 49oN 100oW and 50oN 101oW) which has been a favourite collecting area Peltonotellus bivittatus (Caliscelidae) is a specialist on for many entomologists. prairie muhly, a grass once common on undisturbed mixed-grass sites but now largely restricted to the This information is taken largely from material sides of coulees. in the Canadian National Collection of insects, supplemented by data from DeLong (1931), Doering Others that are not confined to the Great Plains (1940), Strickland (1953), Medler (1958), Nielson are found in areas that were probably once connect- (1965), Kramer (1967), and Hamilton (1972). The ed to the prairies, but as world temperatures declined results of this compilation indicate that there are over the last 7000 years have become isolated in for- at least 226 prairie-endemic Auchenorrhyncha, ested regions. They now inhabit sheltered valleys in of which 168 are leafhoppers (Cicadellidae), 47 the mountains as far north as the Yukon, or patches are (Fulgoroidea), 7 are of sandy soils in boreal forest, or limestone plains (Membracidae), 3 are (Cicadidae) and 1 is a (alvars) in the vicinity of the Great Lakes, and even spittlebug (Cercopidae). Almost all the planthoppers on saline or sandy areas of the Atlantic coast. are Delphacidae, but these are rare compared to the Caliscelidae or Apiglet bugs,@ with 6 species of Bruchomorpha plus one of Aphelonema and two of Peltonotellus, most of which are common and widespread on the plains. The remaining families of Fulgoroidea represented in the endemic fauna are Fulgoridae (or Dictyopharidae) with one species of Apartridge-bug@ Scolops, and Cixiidae with one species each in the genera Myndus and Oliarus. Peltonotellus rugosus (Caliscelidae) is a rare bug restricted to the driest situations. Scolops (Fulgoridae) This summary of what we now know about the are known as Auchenorrhynch fauna of the Canadian prairies is “partridge bugs” based on more than 340 sites (Table 1). Samples tak- because they sit en within a 5 km radius of each other are Alumped@ up with their beak- like head process to make data presentation simpler. These sites are in the air. then grouped according to one-degree quadrats (Fig. 1). In Alberta (AB) and Saskatchewan (SK) there are an average of 4-5 sites per quadrat; in Manitoba

Arthropods of Canadian Grasslands Number 10, 2004 18 19 University of Kansas 41 (22), University of Kansas Science Bulletin 26 (2): 83-167. Kramer, J.P. 1967. A taxonomic study of the brachypterous North American leafhoppers of the Lonatura (Homoptera: Cicadellidae: ). Transactions of the American Entomological Society 93: 433-462. Amblysellus wyomus is a western species that feeds Medler, J.T. 1958. A review of the genus Scleroracus in on June grass; it looks quite unlike the other members North America (Homoptera: Cicadellidae). Annals of its genus, which have longer heads and are grey of the Entomological Society of America 51:230- spotted with black. 241. The species are grouped roughly by over- Hamilton, K.G.A. 1972. The Manitoban fauna of all distribution. Fifty-five species are found wide- leafhoppers (Homoptera: Cicadellidae) Pt. II: ly across the Great Plains; 32 of these also occur in the fauna of macro-leafhoppers. The Canadian Entomologist 104: 1137-1148. other Canadian grasslands, either along the Atlantic coast, or in alvars and relict grasslands of Ontario Hamilton, K.G.A. 1999. Leafhoppers (Insecta: (ON), intermontane grasslands of British Columbia Homoptera: Cicadellidae) as indicators of endangered ecosystems. Pp. 103-113 in J. Loo (BC) and the Yukon Territories (YT), or in the Peace and M. Gorman (Comp.), Protected Areas and the River District of Alberta (Table 2). These wide- Bottom Line. Proceedings of the 1997 Conference spread insects are therefore characteristic of the of the Canadian Council on Ecological Areas, Northern Grasslands Biome. When these are known September 14-16, 1997, Fredericton, New only (or primarily) from the Canadian Great Plains, Brunswick. Canadian Forest Service. they are indicated as Aendemic.@ Nielson, M.W. 1965. A revision of the genus Cuerna (Homoptera: Cicadellidae). USDA Technical Commellus comma is a specialist on Bulletin 1318. Canada wild rye. It Strickland, E.H. 1953. An annotated list of the is common in the (s.l.) of Alberta. The Canadian USA but is found in Entomologist 85: 193-214. Canada only in the canyon of the South Acknowledgments Saskatchewan. This work has been greatly assisted by many pri- Species characteristic of only parts of the Great vate and governmental agencies, notably The Nature Conservancy, Manitoba and Ontario Ministries of Natural Plains can be divided into two biotic provinces, 34 Resources, Michigan Natural Features Inventory, Minnesota species characteristic of eastern grasslands or Atrue and Wisconsin Departments of Natural Resources and Ohio prairie@ and 39 that are widespread on the western State University (for a complete list of participants, see The plains or ACanadian steppe@ (Table 3), with the Canadian Entomologist 132 July/August 2000, page 500). remaining species divided between 40 on moist Particular mention should be made of the following indi- steppe or Aaspen parkland@ and 58 on dry steppe viduals who helped by trapping thousands of prairie in- or Ashort-grass prairie@ (Table 4). The intervening sects on Canadian sites over an extended period: Albert area or Amixed grass prairie@ appears to have no Finnamore at the Alberta Provincial Museum, Edmonton; characteristic bugs. Jeanette Pepper of the Fish and Wildlife Branch, Regina, Saskatchewan; and Rob Roughley at the University of References Manitoba, Winnipeg. DeLong, D.M. 1931. A revision of the American spe- cies of Empoasca known to occur north of Mexico. Hecalus finnamorei USDA Technical Bulletin 231. is known only from Grasslands National Doering, K. 1940. Part III: a contribution to the Park. of the subfamily Issinae in America north of Mexico (Fulgoridae, Homoptera). Bulletin of the

Arthropods of Canadian Grasslands Number 10, 2004 20 21

Figure 1. Distribution of sampled sites on the Canadian Great Plains by one-degree quadrats, with quadrats for each province numbered from west to east, beginning with southern row. Individual sites are listed north to south and west to east within each quadrat, as shown in enlarged view of quadrat 2 of Manitoba (lower left).

Arthropods of Canadian Grasslands Number 10, 2004 20 21

Table 1. Sites sampled on the Great Plains of Canada (conserved sites in boldface), numbered and lettered as in Fig. 1.

Alberta (AB) 7a Weyburn, b Stoughton 5 km W, c 42a Cochin 1a Porcupine Hills (44 km N Lundbreck), Torquay Manitoba (MB) 8a Kenosee, b Kisbey 8 km W, c Kisbey 5 b Frank, c Bellevue 2a Brandon 5-7 mi N, b Oak Lake 11 km km E, d Arcola, e Glen Ewen, f Estevan, 2a Granum, b Smashed-in-Head Buffalo N, c Virden, d Brandon, e Oak Lake, g Roch Percée, h North Portal Jump, c Fort McLeod, d Cowley, e f Griswold 4 km W, g Brandon 15 mi 9a Maryfield, b Redvers 15 km N, c Pincher Ck, f Brocket, g Glenwood, h S (Carroll), h Hartney, i Napinka, j Wauchope 4 km E Waterton Melita, k Deloraine, m Whitewater Lk, 11a Gascoign, b Fox Valley, c Cliftonville, 3a Iron Springs, b Taber 15 km N, c Taber, n Boissevain, p Waskada, q Horton, r d Maple Creek 10 mi NW, e Piapot and d Chin, e Diamond City, f Lethbridge, Goodlands, s Turtle Mountain Prov. 2 mi NW g Welling, h Stirling, i Magrath, j Park 12a Lacedena, b Great Sand Hills Raymond, k New Dayton, m Del Bonito 3a Onah, b Carberry, c Melbourne, d Shilo (Hazlet), c Hazlet 6 mi E, d Swift 10 km NW (Ross Lake), n Milk River, p 5 mi SW, e Carberry 7 mi S, f Glenboro Current 10 mi W, e Webb, f Gull Lk 8 mi Kimball, q Del Bonito 9 km E 13 mi N (Bald Head Hills), g Glenboro N, g Tompkins, h Gull Lake 4a Purple Spgs 5 km W, b Burdett, c 14-18 km N, h Rounthwaite, i Treesbank 13a Matador Prairie, b Saskatchewan Whitla, d Foremost, e Clarinda, f Milk (Aweme), j Stockton MB 2 mi W, k Ldg (Stewart Valley 9 mi W), c River 10 km E, g Coutts, h Writing-on- Glenboro MB and 5 km N, m Ninette, n Waldeck, d Swift Current, e Simmie 5 Stone, i Aden Baldur, p Ninga, q Rock Lake mi NE 5a Irvine, b Walsh, c Elkwater, d 4a MacGregor 3 mi E, b Austin MB 16 14a Riverhurst, b Tugaske, c Eyebrow 3 Orion, e Manyberries, f Orion 8-10 km S, c La Riviere, d Pilot Mound, e mi SE, d Chaplin, e Bateman km S, g Onefour, h Orion 40 km S, i Crystal City, f Morden, g Morden 5 km 15a Aylesbury, b Brownlee, c Moose Jaw 5 Manyberries 45 km S W, h Morden 6 km S, i Snowflake mi S, d Crestwynd 4 mi N 11a Hartell, b Pekisko 5 km W, c 5a St. Fr-Xavier, b Winnipeg (St. Charles 16a Craven 2 km NW, b Lumsden, c Longview 22 km S, d Longview 30 km S Rifle Range), c Winnipeg (airport, Regina, d Davin 12a Gleichen, b Nanton Living Prairie Museum), d Transcona, 17a Duff 2 km E, b Fort Qu=appelle 13a Bassano, b Scandia, c Vauxhall e Winnipeg (S), f Ft Whyte, g Glenlea, h (Lebret), c Katepwa Bch, d Qu=Appelle, 14a Steveville, b Duchess 13 mi N, c Niverville, i Myrtle, j Morris, k Arnaud, e Indian Head, f Qu=Appelle 7 mi S Brooks, d Tilley, e Tilley 5 mi E m Altona (Neuenburg 2 km W ), n 18a Melville, b Duff 8 km E, c Stockholm 15a Empress, b Suffield CFB (Amiens), c Dominion City, p Emerson and 5 km W, d Esterhazy, e Round Lake Medicine Hat 6a Steinbach 9 km S, b Marchand 2 mi 6 km E Birds Point, f B roadview, g 21a Olds, b Didsbury, c Ghost Dam, d W, c Marchand 6 mi E, d Stuartburn Pipestone Creek, h Pipestone Creek 6 Calgary 9 mi W (Chestermere Lake), e 8 km ENE, e Tolstoi 2-6 km E, f km SE St. Hubert, i Peebles, j Kennedy Calgary Gardenton and 3-5 km W 21a Marengo, b Kindersley 13 km W 22a Beiseker 5 mi E 11a Russell, b Binscarth, c Birtle 22a Plato 23a Morrin, b Delia, c Michichi 4 mi N, 12a Rossburn, b Solsgirth, c Kelloe, d 23a Harris, b Rosetown, c Dinsmore d Drumheller, e Rosedale, f Rosebud, g Shoal Lake, e Strathclair, f Rapid City 6 24a Saskatoon 14 km SE, b Beaver Creek, Hussar mi E g Oakner c Pike Lake, d Dundurn, e Dundurn 24a Sheerness 10 mi NW, b Sunnynook 12 13a St. Amelie, b McCreary 3 km E, c 5 mi S (Rudy), f Hanley 3 mi SE, g mi SW Minnedosa and 5 mi N, d Forrest Station Bladworth, h Davidson 11 mi W, i 31a Spruce View 9 mi N Elbow, j Elbow 8 mi SE (Douglas Prov. 32a Wetaskiwin, b Battle River 1st- 3rd 14a Mulvihill 4 km N, b Oak Point 3 mi Park) crossing, c Lacombe, d Red Deer S, c Gladstone, d Woodside 5 km E, e 25a Viscount 11 km E, b Imperial, c 33a Nevis, b Stettler Westbourne, f High Bluff 3 mi S Imperial 11-18 km E (Last Mountain 34a Castor 15a Fraserwood, b Komarno, c St. Laurent WMA), d Davidson 35a Wainwright, b Czar, c Provost 13 km E, d St. Laurent 8 km S, e Lake 26a Dafoe, b Kandahar 42a Edmonton, b Devon Francis 3 mi NW, f St. Ambroise 7- 9 27a Foam Lake, b Ituna 3 km N 44a Ranfurly, b Mannville km E, g Woodlands and 3 mi SE, h 28a Canora, b Willowbrook 12 km E, c 45a Vermilion, b Islay 4 mi S, c Blackfoot Stony Mountain 12 km N, i Grosse Isle Yorkton, d Rokeby, e Crescent Lake Saskatchewan (SK) 21a Roblin 31a Neilburg, b Cut Knife, c Rutland 1a Cypress Hills 22a Pulp River 13 km E, b Ethelbert, 32a Battleford, b Denholm 2a Gull Lk 14 mi S, b Illerbrun 2 mi S, c c Ashville 16 km N (Crocus Hill), 33a Redberry, b Radisson, c Borden, d Simmie 6-7 mi SW, d Scotsguard 3 mi N, d Ashville 11 km N, e Ashville, f Ceepee, e Asquith, f Biggar 16 mi N e Dollard, f Eastend Dauphin, g Keld 34a Blaine Lake, b Duck Lake, c Rosthern, 3a Cadillac 9-11 mi S, b Val Marie 23a Winnipegosis 1 km NW, b Toutes d Fish Creek, e Hepburn, f Vonda, g (Grasslands N Pk) Aides, c Rorketon 6 km SE, d Ochre Martensville, h Saskatoon 3 mi NW, i 4a Assiniboia, b Limerick, c Wood River 4 km N Saskatoon 4 mi NE, j Saskatoon Mountain, d Lisieux, e Killdeer 24a Moosehorn, b Wapah 7-10 km E, c 35a Dana 5a Mossbank, b Ardill, c Willows, d Camper 2.5 km SSE 41a Tangleflags, b Lloydminster 4 mi SE, c Readlyn 5 mi W, e Verwood, f St. Victor, 31a Mafeking Marshall 3 mi NW, d Lashburn 3.5 mi E, g Willow Bunch, h Rockglen, i Fife Lake 32a Cowan e Maidstone 6a Pangman, b Minton

Arthropods of Canadian Grasslands Number 10, 2004 22 23

Table 2. AShort-horned@ bug fauna characteristic of the Northern Grasslands biome including much of the Great Plains of Canada, with distribution (codes as in Table 1). Leafhoppers Flexamia serrata Beamer & Tuthill: BC; AB Mesamia nigridorsum (Ball): AB 21e, 45c; Amblysellus acuerus DeLong & Hamilton: 35a, 44a; SK 8abf, 18a, 24fi, 25c, 27b, SK 4cd, 24d, 34i; MB 3ip, 5hp, 6e, 15di; SK 3a, 5f; MB 15d, 24b. 28de, 31a, 34gi; MB 4g, 6ef, 12c, 14b-d, ON. Athysanella attenuata Baker: BC; AB 3f, 5ce- 15cdghi, 22a, 23c. Mocuellus americanus Emeljanov: AB Peace h, 13c, 14b, 15a-c, 22a, 23ae, 32a, 42b; Gypona contana DeLong: AB 2b, 3fn, 5cg, R District, 45a; SK 3b, 5b, 8b, 17e, 18ch, SK 2a, 3ab, 5d, 8h, 11bd, 12be, 13ad, 14a, 14b, 15c, 21e, 22a, 23dg, 33b; SK 2f, 3b, 25c, 28b, 34bij, 42a; MB 2df, 3k, 4b, 12c, 15b-d, 16b, 17d, 18b, 21b, 24hi, 25d, 28d, 6b, 8f, 9a, 11e, 12bg, 13d, 17c, 24bcdi, 28b, 14a, 15cg-i, 22f, 23cd, 24bc; ON. 33bf, 34i-km; MB 3km, 5c, 14d. 32b, 33e, 34g-j; MB 2a, 3in, 4gi, 5bdehp, Paraphlepsius continuus (DeLong): AB 33b, Athysanella kadokana Knull: AB 5g, 15a; 15dfi. 42a; SK 31b; MB 15e; ON. SK 5bd, 24fi, 25c, 26ab; MB 3j, 12c, 14d, Hebecephalus occidentalis Beamer & Tuthill: Paraphlepsius lascivius (Ball): YT; BC; AB 15gh. AB 3fq, 14ce, 15a-c, 21e, 22a, 23f, 24ab, Peace R District, 3c, 14c, 15b, 23d, 33b; Athysanella secunda Blocker & Wesley: SK 42b, 44b; SK 5de, 8bch, 11e, 13d, 15cd, SK 3b, 4b-d, 5gh, 15a, 21a, 23b, 24cd, 26b, 24f, 25c, 34gi; MB 14b, 15d. Endemic sp. 17bde, 21b, 24h-j, 25ad, 28bde, 32b, 33c, 33b, 34cj; MB 4af, 5ef, 15i. Athysanella. terebrans Gillette & Baker: AB 34gj, 42a; MB 2bf, 3c, 4a, 5e, 12c, 14d, Paraphlepsius solidaginis (Walker): AB 3f, 5fhi, 15ab, 35a; SK 3b, 8f, 11e, 18e, 24c, 15i, 23d, 24c. 5e, 15c, 33b, 35a; SK 4d, 5gh, 6b, 13d, 33c; MB 3cij, 5e. Hebecephalus truncatus Beamer & Tuthill: 28d, 31ac, 34i; MB 3im, 5n, 6e, 12dg, 14d, Attenuipyga minor (Osborn): AB 3b, 4abf, YT; BC; AB 3m, 15b, 23a; SK 24c, 34j; 15di; Atlantic coast. 5afh, 13b, 15bc, 21ce, 23a; SK 2bc, 3ab, MB 2dr, 5e. Paraphlepsius turpiculus (Ball): AB 35a; SK 12bf, 13e, 18bce, 22a, 24aci, 25c, 33a, 34j; Hecalus viridis (Uhler): AB 2b, 3bcfmq, 4adf, 4c, 24i; MB 2deh, 3i, 4f, 11a, 22a; ON. MB 3bin, 4af, 5dimp, 12f, 13b, 14bcd. 5abe-h, 13b, 14a, 15a-c, 23df, 35a; SK 1a, Pendarus magnus (Osborn & Ball): AB 15b; Auridius sandaraca Hamilton: YT; AB 42b; 3ab, 4a, 5g, 11e, 12h, 15c, 21b, 24i, 25c, SK 9c, 12b, 18bc, 24i, 28d, 34j; MB 4g, 6e, MB 3c, 6a; ON. 34j; MB 3bhim, 5cip, 6e, 13d, 14d; ON. 14b, 15di. Ceratagallia viator Hamilton: AB 1a, 2h, 3f, Idiocerus freytagi Hamilton: BC; AB Peace Prairiana cinerea (Uhler): YT; AB 1b, 2h, 5c, 11a-c, 13b, 14a, 15bc, 22a, 23d, 24a, R District, 3f, 5ce, 15c; SK 13b, 16b, 17bd, 3fq, 4i, 5ace-h, 15bc, 21e, 23acde, 33b; SK 33b, 34a, 35a, 42a; SK 2a-d, 3ab, 5bd, 7a, 24j, 32a, 33d, 34ij; MB 2a, 4eg, 5e, 12bd; 2b, 3ab, 4c, 5acdg, 8a, 13e, 15c, 18f, 24ai, 8bf, 9bc, 13e, 15c, 17a, 18cehi, 24ci, 27b, ON. 25c, 33a, 34gj; MB 2c, 3beim, 4f, 5abe, 6e, 28e, 31b, 34j; MB 2d, 3bcimn, 4eg, 5bcdjp, Idiocerus raphus Freytag: BC; AB 3f, 5ce, 14b, 21a. 6ef, 12c, 13b, 14c, 22cf, 23a-d, 31a, 32a. 15c, 23d; SK 5a, 13b, 17b, 24j, 32a, 33d, Prairiana kansana (Ball): AB 15b; SK 2c, 3b; Cicadula longiseta (Van Duzee): YT; BC; AB 34ij; MB 2ar, 3di, 4e-g, 5eg, 12b, 14f, 31a; MB 3i, 4g, 5b, 6e, 15b, 24c; ON. Peace R District, 12b, 21b, 22a, 33b, 44a; ON. Psammotettix knullae Greene: YT; BC; AB SK 8b, 21c, 31a, 33b, 34ag, 41b; MB 2a, Laevicephalus poudris Tuthill: AB 2h, 34a, 5e, 14c, 15bc, 22a, 44a; SK 2e, 4a, 8b, 22a. 35a, 44a; SK 2b, 18a, 24fi, 31ac, 34gi, 13d, 24di, 32b, 33b, 34acj, 41e; MB 2d, 4b, Commellus sexvittatus (Van Duzee): YT; BC; 35a; MB 6ef, 13d, 14b-d, 15cdf-i, 22f. 13b, 14c, 22a, 23acd, 24b, 31a. AB Peace R District, 5begh, 12a, 14a, Laevicephalus saskatchewanensis Hamilton Texananus marmor (Sanders & DeLong): 15a-c, 21e, 23de, 42b; SK 2b-d, 3b, 5c, 6a, & Ross: SK 5b, 12e, 18b, 24h, 28d, 33b, AB 23d, 33b, 34a, 44b, 45a; SK 18b, 23a, 7a, 8h, 12beh, 13ae, 15c, 24i, 28d, 34j; MB 35a, 41b; MB 5c, 12c, 14bd. Endemic sp. 24bi, 34bj; MB 3abeij, 11a; On. 2hj, 4d, 5c, 11a, 15b, 32a. Limotettix symphoricarpae (Ball): MB 2h, 5e. Xerophloea zionis Lawson: BC; AB 15c; SK Deltocephalus serpentinus Hamilton & Ross: Limotettix uneolus (Ball): YT; BC; AB 3f, 5c, 2b-d, 3a, 13e; MB 2p, 3i, 5e. AB 21d, 23d, 33b; SK 3b, 5d, 8b, 11d, 14c, 15bc, 22a, 33b; SK 2e, 3b, 5dg, 17b, Other bugs 17bd, 18b, 24fhi, 28de, 34j, 41c; MB 12d, 24gi, 33be, 34ej; MB 2an, 3i, 4df, 5e. Aphelonema simplex Uhler: AB 3c, 5g, 35a; 14d, 24b, 31a; coast of James Bay, ON. Limotettix utahnus (Lawson): BC; AB 42a; SK 5g, 8c, 14e, 18ac, 24i, 26b, 28de, 31c, Destria crocea (Beirne): SK 8g 14e, 24fi, 31a, SK 15d, 17f, 24i, 31b, 34j; MB 4f, 5efg. 34ij; MB 2nq, 4d, 5bdi, 6e, 12c, 13b, 14b- 34i; MB 3i, 4f-h, 5bce, 15dhi; Atlantic Lonatura salsura (Ball): BC; AB 3c, 14a, e, 15dghi, 22af. coast. 15ab, 33b, 45bc; SK 5bdg, 8b, 18bc, 24fhi, Bruchomorpha beameri Doering: BC; AB Driotura robusta Osborn & Ball: AB 5h, 25c, 33b, 34a, 41b; MB 2m, 6e, 14d, 15di, Peace R District, 2bh, 5ceg, 14b, 22a, 15b; SK 3ab, 8f, 18e, 26b, 28d; MB 2jkr, 23ab, 31a. 23cdf, 24ab, 33b, 35ac, 44b; SK 1a, 2b, 3beg. Lonatura teretis Beamer: AB 32c; SK 18eh, 3ab, 5bdeg, 8cf, 11ae, 12b, 13bde, 15b-d, Erythroneura carbonata McAtee: AB 45b; SK 24fi, 27b, 34i; MB 15di. 17bf, 18a-c, 24bci, 25c, 28bd, 31ab, 32ab, 8c, 34j; MB 13b, 23a. Macropsis feminis Hamilton: AB 2c, 23d; 33bcf, 34bgj; MB 2gn, 3bdei, 4b, 5b. Euscelis maculipennis DeLong & Davidson: SK 18d, 33d, 34j; MB 2e, 3bi, 4g, 5e, 6b, Caenodelphax nigriscutellata Beamer: AB BC; AB 3f, 5cg, 15b, 35a; SK 3b, 5g, 16b; 12bf, 13c. 5fi,13b, 15b; MB 3g, 5b; ON. MB 4f, 5e. Macropsis rufescens Hamilton: AB 5e, 15c, Cixius cultus (Ball): AB Peace R District, Extrusanus oryssus Hamilton: AB Peace R 23d; SK 4c, 16b, 33d, 34j; MB 2d, 3b, 5e. 3cf-jn, 5ceg, 14cd, 15a-c, 23g; SK 1a, 3b, District, 3f, 5cgh, 11b, 15a-c, 23d, 32d, Macropsis rufocephala Osborn: AB 21b, 23d, 5i, 8a, 13d, 24i, 34j. 35a, 42b; SK 1a, 2bc, 3ab, 4c, 5cg, 6a, 8a, 17b, 24i, 27a, 32a, 33d, 34ij; MB 2s, 3i, Delphacodes sp.nov. A: AB Peace R District, 13e, 16b, 18g, 21b, 24ai, 25c, 28ac, 31ac, 4ag, 5e, 12b, 13c; ON. 4f; SK 24i, 26b, 34j; MB 14b. 33a, 34j; MB 2iqs, 3bimnq, 4g, 5b-ep 6aef, Memnonia anthalopus Hamilton: AB 35a; Pissonotus delicatus Van Duzee: BC; AB 3cf, 14bd, 15cdhi, 21a, 22e, 24b, 31a. SK 3b, 24i, 25c, 31a, 34j; MB 6ef, 14bd, 14acd, 15ac, 23d; SK 5g, 8b, 11a, 13d, Flexamia decora Beamer & Tuthill: AB Peace 15cdhi, 24b. 16bc, 24i, 31a, 32b, 34gj; MB 2jm, 5i, R District, 2h, 12a, 15b, 21e, 35a; SK 2bc, Mesamia ludovicia Ball: AB Peace R District, 11ab, 14d, 23ab. 3b, 5h, 8a, 11d, 13a, 18a, 24fi, 28d, 31ac, 2b, 3cf, 15b, 23d, 34a; SK 2c, 3b, 8f, 13d, Publilia brunnea (Ball): AB 3f, 4a, 5acde, 34gij, 41a; MB 6e, 14bd, 15cdfhi, 24bc. 18c, 24bi, 25c, 34gj; MB 15i. 13b, 15abc; SK 11a, 12bh, 13bd, 14d, 16a, 18ce, 24bci, 25b, 28ab, 33e; MB 2b, 4g, 5bi, 15a, 22cd, 23c, 24a; ON.

Arthropods of Canadian Grasslands Number 10, 2004 22 23

Table 3. “Short-horned” bug fauna characteristic of two major subdivisions (biotic provinces) of the Great Plains of Canada, with distribution (codes as in Table 1). BIOTIC PROVINCE: PRAIRIES 14bd, 15dghi, 22cd, 24c, 32a; ON. 3ab, 11a, 13e, 24ai, 25c, 34j. Leafhoppers Delphacodes sp.nov. B: MB 5c. Hebecephalus rostratus Beamer & Tuthill: AB Aflexia rubranura (DeLong): MB5b, 14b, Megamelus metzaria Crawford: MB 2g, 5b, 2b, 3cf, 5ceg, 14be, 15c, 22a, 23a-d, 34a; 15dfhi, 24b; ON. 6e, 15h, 23a; ON; Atlantic coast. SK 2d, 3b, 5bd, 8ae, 11d, 12f, 13ade, 14bc, Athysanella incongrua Baker: MB 3g, 15g. Megamelanus sp.nov.: MB 14d. 15bd, 17be, 18b, 24e-j, 25d, 33c, 34gij; Attenuipyga joyceae Hamilton: MB 14b, crocea (Van Duzee): SK 16a, 17d; MB 2a, 4g, 14d. 15di. Endemic sp. MB 4gh, 5bcdfim, 6e, 13a, 14a, 15f, 23a; Idiocerus ramentosus Uhler: BC; AB Peace R Commellus colon (Osborn & Ball): SK 18eh; Atlantic coast. District, 3fnp, 5ade, 15c, 23d, 44b; SK 5a, MB 3di, 5bc, 11a, 15di. gibbosa Ball: 7 SK 8g; MB 3abdfi, 7c, 12b, 17b, 24cj, 32a, 33e, 34j; MB 2ak, Commellus comma (Van Duzee): AB 15b; 4b; Atlantic coast [MA]. 4ceg, 12a, 13c; ON. SK 24i. Tortistilus minutus Cald.: 3 MB 3b, 4g, 14b. Memnonia brunnea (Ball): AB 3g, 5a; SK 8b, Cuerna fenestella Hamilton: MB 3im, 5b, 15i, 24i, 34j; MB 14d. BIOTIC PROVINCE: STEPPE 24c; ON. Mocuellus caprillus Ross & Hamilton: AB 1c, Leafhoppers Flexamia atlantica (DeLong): SK 18e; MB 2bh, 3bf, 4ai, 5afg, 12a, 13b, 14bc, 15a-c, Acinopterus viridis Van Duzee: BC; AB14be, 15g. 21e, 22a, 23d-f, 24b, 33b, 34a, 35a, 44b; 23d, 24b, 33b, 34a; SK 3b, 5e, 13a, 23bc, Flexamia delongi Ross & Cooley: SK 18e; SK 2b-d, 3ab, 4a, 5g, 8h, 11d, 12beh, 13ae, 24ej, 34g. MB 3i, 14bd, 15bcdghi, 24c; ON. 15cd, 16b, 17bd, 23b, 24ci, 25cd, 33f, 34ij; Amblysellus valens (Bm. & Tut.): AB 5c, 23f, Flexamia graminea (DeLong): SK 9c, 18ch; MB 2j. 24a, Peace R District; SK 2b, 3ab, 34j. MB 2f, 4g, 13b, 23c. Oncopsis marilynae Hamilton: BC; SK 8f, Athysanella bifida Ball & Beamer: AB 3bfq, Flexamia inflata (Osborn & Ball): BC; MB 34j; MB 2ns, 3b. 5cegh, 13b, 14a-ce, 15c, 22a, 23de, 35a, 5d, 6ef, 13b, 14d, 15h, 22f, 23a, 32a; ON. Orocastus labeculus (DeLong): BC; AB 5c, 45c; SK 2a-d, 3ab, 4c, 8abfh, 11b, 12h, Flexamia prairiana DeLong: MB 4g, 5b-d, 12a, 13b, 14ace, 15a-c, 21e, 24ab, 34a, 13ae, 15bcd, 18g, 24ci, 25c, 34gj; MB 2q, 6ef, 14c, 15dgi, 22bc, 24b; ON. 35a; SK 2bc, 3ab, 11d, 12h, 13e, 14a, 3bj. Flexamia reflexa (Osborn & Ball): MB14d, 15cd, 24i, 34j. Athysanella occidentalis Baker: AB 3fq, 4a, 22f; ON. Orocastus perpusillus (Ball & DeL.): YT; BC; 5ce, 12a, 15ab; SK 2a, 3b, 11b 12h, 15c, Flexamia sp.nov.: MB 14c. Endemic sp. AB Peace R District, 3bf, 4a, 12a, 13b, 24ci, 25d, 34j. Graminella ampla (Beamer): SK 8g; MB 5c. 14a-c, 15a-c, 21e, 23df, 34a, 35a, 44b; SK Athysanella robusta Baker: YT; BC; AB Laevicephalus unicoloratus (Gillette & 2b-d, 3ab, 8a, 11d, 12h, 13ae, 17b, 18a, Peace R District, 3mq, 4a, 5aceghi, 14a, Baker): SK 18ceh, 28b; MB 2f, 3k, 4g, 5dp, 22a, 24i, 33cf, 34gj; MB 2q, 3cdik, 22c. 15a-c, 21e, 23e, 32a; SK 1a, 2a-d, 3ab, 6a, 6ad-f, 13b, 14b-d, 15cdf-i, 23cd, 24c; ON. Rosenus cruciatus (Osborn & Ball): YT; BC; 7a, 8h, 11a, 12h, 13ae, 24acdi, 25c, 33a, Laevicephalus vannus Knull: MB 6b, 23a? AB 3bfq, 4adg, 5ce-i, 15b, 23ac-e, 32b, 34fj; MB 3i. Latalus latidens (Sanders & DeLong): MB 2d. 44b, 45c; SK 2bc, 3ab, 6a, 11b, 13e, 15d, Auridius auratus (Gillette & Baker): BC; Limotettix urnura Hamilton: SK 5g; MB 5eg, 18eh, 24aci, 25c, 32a, 34j; MB 3deijk, 22c. AB Peace R District, 1c, 4ai, 5e, 12a, 14c, 6e, 15i; ON. Stenometopiellus cookei (Gillette): AB 3q, 15ab, 21de, 23c, 31a, 33b, 35a; SK 3ab, Macrosteles flavalis Hamilton: 5 MB2d, 3d, 5ei, 13c, 15b; SK 2a, 3ab, 13e, 24ai, 34j. 11d, 12e, 13a, 24fi, 32a, 33a, 34ij. 4f, 5eg. Endemic sp. Auridius ordinatus (Ball): YT; BC; AB Peace Other bugs Macrosteles sp.nov.: MB 24b. Endemic sp. R District, 2b, 3j, 12a, 15b, 21e, 23f, 32b, Delphacodes paransera Beamer: AB Peace R Memnonia panzeri Hamilton: MB 5bp, 6ef, 42a; SK 2bd, 3b, 5h, 8ah, 11d, 12de, 13ae, District, 35a; SK 5f, 8a, 24j; MB 2c, 3i. 14b, 15dhi; ON. 15c, 17e, 18g, 24i, 25c, 31c, 33f, 34j; MB Elachodelphax indistincta (Crawford): AB 3f, Neohecalus magnificus Hamilton: MB5b-eik, 2q, 6c. 5c, 15c; SK 3b, 24di, 34j; MB 3i. 6e, 14b, 15di; ON. Ceratagallia semiarida Hamilton: AB 3f, 5c, Eurybregma magnifrons (Crawford): YT; BC; Paraphlepsius hemicolor (Sanders & 13b, 14c, 15c, 23bd, 35b; SK 3b, 11a, 12a, AB Peace R District, 13b, 15ab, 23d; SK DeLong): MB 4f, 5ej. 15c, 24bfi, 34j; MB 3i. 3b, 23a, 34j. Paraphlepsius lobatus (Osborn): SK 9b, 17a, Deltocephalus lineatifrons Oman: BC; AB Laccocera obesa Van Duzee: BC; AB 2b, 3f, 18ceh; MB 4g, 12c, 15i, 22cd; ON. 13c, 15b; MB 2h. 5adegh, 14b, 15a-c; SK 3b, 5g, 11ab, 13ae, Paraphlepsius umbrosus (Sanders & Empoasca medora DeLong: AB 3e, 5ab, 14c, 21b, 24i, 34j. DeLong): SK 18ce; MB 3b, 4g, 5e; ON. 15a-c, 34a, 35a; SK 3b, 24i; MB 11a. Nothodelphax gillettei (Van Duzee): BC; AB Polyamia caperata Ball: SK 18ceh; MB 4gh, Flexamia abbreviata (Osborn & Ball): AB 2b, 3fnq, 5cfg, 11c, 21e, 23d; SK 3b, 24i, 28a, 5bc, 13b, 14c, 15di, 22d; ON. 3f, 12a, 14ab, 15bc, 21e, 23df, 24b, 35a; 33a. (Van D): SK 18c, 28b; MB SK 2b, 3b, 5cf, 8f, 11d, 13ae, 15cd, 18eh, Peltonotellus bivittatus Ball: BC; AB Peace R 2f, 4g, 5bp, 6e, 14bd, 15cdghi, 23c, 24c. 24i, 25c, 34j; MB 2j, 3cdegi, 13c. District, 2b, 5g, 14b, 15c, 23d; SK 3b, 5g, Other bugs Flexamia flexulosa (Ball): AB 2b, 3f, 5ceg, 8af, 18eh, 24i, 34j. Bruchomorpha dorsata Fitch: SK 18eh; MB 14a-c, 15bc, 22a, 24b; SK 2b, 3ab, 8fh, Pentagramma vittatifrons (Uhler): AB 3ef, 5g, 4g, 5bc, 6ef, 15cdf; ON. 11d, 13ae, 24i, 34j; MB 2q, 22c. 15ac, 23g, 35a; SK 4c, 8f, 11e, 17b, 24di, Bruchomorpha jocosa Stål: SK 8cf, 18ceh, Flexamia grammica (Ball): AB 2b, 5ei, 15a, 26b, 31b, 33a, 34j; MB 2g, 4d, 12e. 28ab; MB 3c, 5p, 6e, 13b, 15dfi, 22c, 23c. 23d, 35a; SK 11e, 12b, 13b, 24di, 34j. Prokelisia salina (Ball): BC; AB 2bg, 15b; Bruchomorpha keidensia Doering: AB Peace Flexamia stylata (Ball): AB 2b, 5a, 15b, 21e; SK 11e, 12b, 17d, 21b, 25a, 26b, 34i; MB R District; MB 6e, 15bdi, 22dfg, 31a. SK 3b, 5g, 13a, 18ehj, 24i; MB 2jq. 3cdfj; ON. Endemic sp. Frigartus frigidus (Ball): AB 15b, 22a, 34a, Campylenchia rugosa (Fowler): BC; AB 1c, Bruchomorpha pallidipes Stål: SK 18c; MB 35a; SK 2bc, 3b, 8c, 13e, 23b, 24i, 34ij; 2dh, 3ef, 4i, 5ce, 13b, 15abc, 21e, 33b, 35a; 3a, 23c. MB 3i. SK 3b, 4e, 5gh, 12bc, 13bd, 17c, 18c, 24ci, Delphacodes angulata Beamer: MB 4g. Hardya dentata (Osborn & Ball): BC; AB 1a, 31bc, 34j; MB 2eq, 3bin, 11b, 22d. Delphacodes parvula (Ball): MB 5b-dp, 6ae, 3mq, 5acehi, 11ad, 15b, 23d, 34a; SK 2a-c,

Arthropods of Canadian Grasslands Number 10, 2004 24 25

Table 4. AShort-horned@ bug fauna characteristic of two biotic regions of the steppe of Canada, with distribution (codes as in Table 1). BIOTIC REGION: ASPEN PARKLAND Elachodelphax pediforma (Beamer): BC; AB Memnonia maia Hamilton: AB 3mq, 5ce, Leafhoppers 2h, 5c; SK 8a; MB 2q. 15b; SK 2acd, 3ab. Endemic sp. Amblysellus dorsti Oman: AB 33b, 35a, SK Elachodelphax sp.nov. A: 5 BC; AB 4f, 5c, Mesamia straminea (Osborn): AB 3f; SK 13e. 24i, 34j. 23d, 32d; SK 34f. Neokolla surcula (DeLong & Currie): AB 3f. Ceratagallia cerea Hamilton: AB 5c. Elachodelphax sp.nov. B: MB 2c. Norvellina clarivida (Van Duzee): AB Peace Colladonus olus Nielson: SK 34j. Laccocera canadensis Beirne: AB 3m, 5c, R District, 5a, 14c; SK 24i. Cuerna nielsoni Hamilton: SK 17e; MB 3cm. 23d; SK 1a, 2b, 3a, 24ai, 33a, 34j. Norvellina columbiana (Ball): BC; AB 23d. Endemic sp. Myndus collinus Ball: SK 34j. Stragania atra (Baker): BC; AB 3e, 5e, 15c; Deltocephalus castoreus Ball: BC; AB 33b, Scolopygos pallidus Bartlett: 4 AB 23d, 33b, SK 3b. 35a; SK 16b, 24i, 34a. 35a; SK 31a; Atlantic coast. Telusus sp.nov.: AB 15ab; SK 3b, 24i. Deltocephalus sp.nov. A: SK 31a. Vanduzea triguttata (Burmeister): SK 18c; Endemic sp. Elymana circius Hamilton: BC; AB Peace MB 2d, 3i. Texananus cumulatus (Ball): AB 15c; SK 3a; R District, 2d, 21ae, 22a, 23cd, 33b, 34a, MB 2p, 3a. BIOTIC REGION: SHORT-GRASS 35a; SK 5d, 24fi, 31a, 33bf, 34gij. Texananus dolus DeLong: MB 4de. STEPPE Empoasca digita DeLong: BC; SK 24i; MB Texananus extremus (Ball): AB 15c. Leafhoppers 11c. Unoka gillettei Metcalf: AB 4a. Amblysellus punctatus Osborn & Ball: BC; Empoasca exiguae Ross: BC; AB 3f, 13c; SK Xerophloea peltata (Uhler): AB 2bc, 3f, 4h; AB 14b, 15b, 24b, SK 2bc, 3ab, 13e. 34j; MB 3i. SK 3b, 4a, 5b, 8f, 15d; MB 2s. Amblysellus wyomus Kramer: BC; AB 1c, Empoasca mexicana Gillette: SK 34j. 2b, 3m, 5f, 15b; SK 2b, 3b, 6a, 13e, 25c. Other bugs Empoasca nigroscuta Gilette. & Baker: BC; Aplanus albidus (Ball): AB 23d. Okanagana fratercula Davis: AB 2eh, 5a, 21e. AB Peace R District, 3f, 5g; SK 24bi. Athysanella obesa Ball & Beamer: AB 3mq, Okanagana luteobasalis Davis: AB 3f, 5c, Erythroneura calva Beamer: 3 AB 13b; SK 5acg, 15b; SK 2b-d, 3ab, 4c, 5f, 13e, 24i. 15c. 25c; MB 3i. Athysan ella sinuata Osborn: AB 3f, 4b, 5ceg, Okanagana synodica (Say): AB 3f. Flexamia dakota Young & Beirne: SK 24i. 14a, 15a; SK 11b, 24i. Bruchomorpha sp.nov.: SK 8f. Flexamia sp. SK 3b. Attenuipyga balli Oman: SK: 3b. Delphacodes emeljanovi Wilson: YT; BC; Gyponana vincula DeLong: AB 2b, 3f, 4c, Auridius helvus (DeLong): BC; AB Peace R AB 11bc. 15b, 23f, 24a; SK 18d, 32ab, 34gij; MB District, 2b, 12a, 14be, 15bc; SK 2b-d, 3b, Delphacodes munda (Beamer): AB 3n, 5g. 3m, 15i. 13e; MB 3k. Eurybregma montana (Beamer): AB 5g, 13b, Idiocerus moniliferae Osborn & Ball: SK Ballana ortha DeLong: BC; 2 AB 34a; SK 5g. 14c; SK 12e. 16b, 33d; MB 2a. Ceratagallia califa Oman: AB 15b. Laccocera flava Crawford: AB 3n, 4d, 5eg, Limotettix bisoni Knull: AB 35a; SK 17b, Ceratagallia cinerea (Osborn & Ball): AB 14a, 15a-c; SK 23e, 24i. 31a; MB 2m; On. 5ag, 14b, 15c, 23de, 34a; SK 3b, 5c, 24i. Laccocera lineata Scudder: BC; AB 5ch, 15b; Limotettix brooksi Hamilton: AB 45c. Ceratagallia nanella (Oman): SK 3b. SK 2c, 3a, 13e. Endemic sp. Cuerna alpina Oman & Beamer: AB 2b, Oliarus dondonius Ball: 8 AB Peace R Limotettix conservatus Hamilton: AB 33b; 3dfghjkn, 5dgi, 14ab, 15a-c; SK 2f, 3b, 11e, District, 5aeg; SK 3b, 4c, 5g, 6b, 24i. SK 31b. 13b, 17b, 18e, 24i. Parkana alata Beamer: BC; AB 21d. Limotettix myralis (Medler): AB 23d; SK 34j. Cuerna sayi Nielson: AB 3n, 5de, 15c. Peltonotellus rugosus (Ball): AB Peace R Limotettix shastus (Ball): BC; SK 35a. Deltocephalus artemisiae (Gillette & Baker): District, 2b, 3acfn, 14be, 15ab, 24b; SK 3b, Limotettix taramus (Medler): SK 34j. AB 3f, 14c, 15b; SK 2b, 8b, 13e, 16b, 24i. 6b, 8f, 24ci. Macrosteles snowi (Dorst): AB 32d, 35a; SK Deltocephalus gnarus Ball: AB 3f, 15b; SK Scolops hesperius Uhler: AB 3ch, 14b, 15ac; 24i, 34j; MB 3bf. 3b; MB 3di. SK 13b; MB 3ef. Oncopsis incidens Hamilton: BC; SK 8d, 34j. Deltocephalus sp.nov. B: SK 3b. Yukonodelphax kendallae Wilson: YT; BC; Oncopsis juno Hamilton: BC; SK 24i. Dicyphonia ornata (Baker): AB 15b. AB 13c, 14d, 15c; SK 2b, 3a, 24i. Pinumius sexmaculatus (Gillette & Baker): Empoasca angustifoliae Ross: AB 2f. Ceresa wickhami Van Duzee: BC; AB 1a, 2h, BC; AB Peace R District, 1c, 21e, 32b, Empoasca typhlocyboides Gillette & Baker: 5e, 15b. 42b; SK 12e, 34dj. BC; AB Peace R District, 15b; SK 2bc, Leioscyta ferruginipennis (Goding): AB 5eg; Stragania rufoscutellata (Baker): BC; AB 3ab, 24i. SK 5g, 6b. Peace R District, 2h, 3ef, 4eg, 5a-eh, 15bc, Erythroneura sp.nov.: AB 15b; SK 3b. Telamona viridia Ball: AB 3f, 5e, 15c; SK 21e, 23d, 32d, 33ab, 35a; SK 2f, 8a, 11c, Hecalus finnamorei Hamilton: SK3b. 16c, 18d; MB 3i. 13e, 16d, 17b, 18af, 24ei, 31c, 33a, 34dgj, Endemic sp. 41d; MB 2cdkn, 3bdijmn, 13c. Idiocerus canae Hamilton: AB 15c. Endemic Other bugs sp. Caenodelphax sp.nov.: SK 25c. Endemic sp. Idiocerus interruptus Gillette & Baker: AB 3f. Criomorphus inconspicuus (Uhler): MB 2c. Laevicephalus exiguus Knull: AB 3e; SK 3b, Delphacodes anufrievi Wilson: YT; BC; SK 13b, 24i; MB 3f; ON. 24i, 34j. Limotettix cacheolus (Ball): AB 2a. Delphacodes lutulentoides (Beamer): BC; SK Lonatura megalopa (Osborn & Ball): SK 3b, 8a, 34j; MB 2s. 24i. Okanagana synodica (Cicadidae) Delphacodes stricklandi Metcalf: AB Peace R Lonatura melina (DeLong): AB 15a; SK 5g, District, 13a, 21d; SK 6a, 7b, 8a, 28e, 34j; 24fhi. MB 2m, 3m, 4d, 31a. Memnonia consobrina Ball: AB 15b; SK 2c, Elachodelphax hochae (Wilson): YT; BC; AB 3ab. All illustrations by K.G.A. Hamilton; Peace R District, 5c; SK 8a. Memnonia grandis (Shaw): AB 15b. Cicadellidae unless otherwise noted.

Arthropods of Canadian Grasslands Number 10, 2004