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NuvtsiaVol. 4, No. 1 (1982)

The species are arranged alphabetically except for A. adsurgens, A. kempeana and A. rhoclophloiawhich are discussedunder A. stowardii. The distribution of each species is indicated with respect to its occurrencewithin Beard's (1980) Bolanical Districts. The 1:250000 map referencesare also given so that distributions can be correlated with those listed in Hnatiuk and Maslin (1980) and Maslin and Pedley (in press).

The environment A detailed description of the Pilbara region is given in Beard (1975) and summarized in Beard (1980). The brief notes on climate, geology/geomorphologyand vegelation which follow, are largely taken from these two accounts.

The Hamersley Plateau has an arid-tropical climate with an annual precipitation 'fhe of 250-300mm. climate is largely dorninated by tropical cyclones which occur predominantly from January to March. Average temperatures for Wittenoom, situated on the north of the Plateau, range from about 18'C in June and Julv to about 33'C in the November-March period.

The Hamersley Plateau is a mountainous region lying between the Fortescue and Ashburton Rivers, It conprises massive deposits of Lower Proterozoic sediments (mainly jasperlite and dolomite) and volcanics overlying Archean granite and volcanics. On its northern flank the Plateau is bounded by an abrupt escarpment ironting the FortescueValley. Around Wittenoom, short rivers have cut deep, narrow gorges into lhe Plateau. When present, soils on the ranges are rocky and skeletal. Further south and east the topography is different and comprises broken country of impressive ranges separated by broad alluvial plains of deep, earthy loams. The highest eminence in Western Australia, Mt Meharry (1 235 m), occurs in this area.

The vegetation of the jasperlite and dolomite ranges is characteristically a tree steppe dominated by Eucalyptus leucophloia (snappy gum) wiLh a Triodia wiseana (spinifex) ground cover. On basaltic hills, as in the Tom Price-Paraburdoo area, a vegetation mosaic of Acacia aneura (mulga) low woodland and A. inaequilatera- ?riodia shrub steppe is found. Most of the valleys carry A. aneura low woodland.

Phytogeography of Acacia in the Hamersley Range area Distribution maps of all speciesmentioned in this discussion are given in Maslin and Pedley (1982).

As discussed by Hopper and Maslin (1978) the Pilbara is one of the secondary centres of species richness for Acacia in Western Australia. Of the 54 species recorded lbr the Pilbara (Maslin and Hnatiuk, unpublished) 46 occur in the Hamersley Range area (Table 1). Only,1 species,A. daweana,A. effusa, A. ex rs and A. hamerslevensisare endemic. "fhe Acacia flora of the Hamersley Range area is dominated by sections Juliflorae (25 species)and Phyllodineae (14 species)which together comprise 84 per cent of the total. The remaining seven speciesare contained in sections Acacia, Lycopodiilbliae and Plurinerves (Table 2).

The main geographical afTinities of the Hamersley Range area Acacla speciesare with arid zone and subtropical regions. This is evident from Tables 1 and 2 where it is seen that 33 species (729") and 32 species (70',rn) are shared with the adjacent Central Eremaean and South Kimberley Areas respectively (see Figure l for FFF>>tsF>ts$>tsFF>F>>F>F > >tstsi' 6rt $ E$ !o $ $ $ *gi.YFil fir€:+eii$i sE$u=E [$ XE: :3-+EF:Y;5SH|! l i 3*E : t{ { ^-:.

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aq} ur errPry'uIIseI g g s9 eEuBU.{alsraur€H l l- 1""-l,i.J;^]i' the genus contribute to these arid zone/subtropical affinities but most species are contained in the sections Juliflorae and Phyllodineae. For example, A. aneura, A. cuthbertsonii, A. kempeana, and A. stowardri (all section Juliflorae) and A. dictyophleba, A. inaequilatera, A. maitlandii, A. pruinocarpa and,A. tetragonophylla (all section Phyllodineae) are common in the central and southern parts of the Australian arid zone. Most of these speciesfind their western or north-western limit of distribution in the region of the Hamersley Range. The subtropical affinities are attributed mainly to species distributed in both the northern arid zone and in subtropical parts of northern Australia, e.g. A. acradenia, A. ancistrocarpa, A. hilliana, A. fenuissima, A. tumida (all section Juliflorae), A. coriacea, A. moiticola, A. retivenia (all section Plurinerves) and A. "victoriae(section Phyllodineae).

Based on current taxonomic knowledge, 44 of the 46 Hamersley Range area Acacra specieshave at least one known closerelative and are therefore classified as beine of recent origin according to the criteria adopted by Stebbins and Major {196si. O;ly A. claweana and A. tetragonoplylla seem to qualify as possible relict taxa becausl neither is known to have taxonomically close relatives. The speciesrichness of the Hamersley Acacia llora is therefore the result of an accumulation of recentlv derived taxa rather than through the persistenceof numerous relict taxa. This accords with results shown for other areasof Western Australia (Hopper and Maslin, 1978).

Of the recently derived species,six are either restricted to the Hamerslev Ranee area or extend only to nearby regions. These six speciesare listed here toget-herwiih their closest relative(s) and it is to be noted that in every case, at least one of the closerelatives also occurs within the Hamersley Range area.

A. arrecta/A. abbreviata, A. arida, A. hilliana, A. minutifolia, A. orthocarpa;

2. A. atkinsiana/A. adsurgeas, A. duriuscula, A. kempeana, A. nelsonii, A. rhodophloia, A. stowardii;

3. A. effusa/A. chisholmii, A. gracillima, A. lysiphloia, A. trachycarpa;

,1 A. exilis/A. tenuissima;

5. A. hamersleyensis/A. citrinoviridis, A. tumida, A. xiphophylla;

6. A. marramamba/A. inaequilatera, A. pyrifolia, A. strongylophylla.

Bowler (in press) proposed that areas peripheral to the central arid zone were climatically the most unpredictable regions of Australia during the euaternary. According to Maslin and Hopper (in press), the resultant siresses'placed on sedentary organisms in such regions created situations ideal for localized sneciation. From the data given above it is evident that the Hamersley Range area has been one such region of speciation. It is intriguing to speculate why a number of the recently_ evolved Hamersley- Range species are confined to the nearby area. perhaps, as suggested by Randell and Symon (19??), ihe formation of extensive sand- dune systems some 30 000 years B.P. have played an important role as an isolating ,,isolated,, mechanism. Certainly the Hamersieys are in the sense that thev are bounded to the north and east by the Great Sandy Desert and the Little Sandy Desert and to the south by the Ashburton River sedimentary basrn. 'seare 'qtroN (.{ur)IaqtEarll puEIEr'tuaJ 0t pue(r.,\s) puE qtnos jiurMoqs(UMS) uor;au qlnos :01 seaJeuPaEtuatg (as) qlnos pu€ IEts€oJ lsa,^A 6 lse]!\ 't'(aMu)6 S (a,$s) qlnos !(e)) r{uoN 1s3l }saruqtJoN't Bur^\oqs(ug) uortau tsan 'sParBI lsrlual 9'(au) I'(e^\) uBaEuarg :8 t Ialraqury (Is) qtnos Z puB ({u) q?roN I 8ur^'\oqs(Ey) uorBeu .{elreqrrry :Z I 's[ar pu8.q I x oI uo paseq(paqsrtqndun) urtsew plrp {nrtEuH ,{q peurJapsBarv Enerv ua1 aqt Sulaoqs EItErtsnV uretsar!\ Jo dPt{ I arntlJ

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Taxonomy

Key to the Acacia species of the Hamersley Range area (Numbered species are described and/or discussed in the text. Descriptions lbr speciesmarked with an asterisk (*) are in Maslin (1981)).

1. Foliagebipinnate; stipules spiny. ...*A. farnesiana Foliagereduced to phyllodes...... 2 2. Phvllodes with very sharp, spiny, needle-like tips; flower-headsglobular...... 3 Phvllodes not sharp and spiny (although sometimes the tips may be coarsely pungent, especiallyon terete ph:'llodes); flower-heads globular to cylindrical.. .8 3. Phyllodes 1-4 cm wide, elliptic but often asymmetrically so; stipules spiny, sometimesdeciduous;inflorescencesracemoseorpaniculate ...... 4 Phyllodes less than 1cm wide, linear to narrowly oblong-elliptic or narrowly obovate; inflorescencessirnple and axillary (neither racemosenor paniculate). .6 .1. Phyllodes symmetric with the rnidrib centrally situated, often glaucous; inflorescenceaxes yellowish or with a pale purplish tinge...... 10. A. pyrifolia Phyllodesasymmetricwiththernidribsituatednearthelowermargin...... 5 5. Branchlet apices white-pruinose; phyllodes 2.5-7 cm long, 1.5 3.5 cm wide, often grey-green,reticulum prominent; inflorescenceaxes purple; bark corky; gnarled treesto c.5 m tall...... *A. inaequilatera Branchlet apices not pruinose; phyllodes 2-4 cm long, 1-2 cm wide, pale green, reticulum obscure; inflorescenceaxes red-brown; bark fibrous; shrubby trees to 3(5)m tall ...9. A. matamamba 6. Phyllodes fasciculate (solitary on new shoots), acicular, c. 1mm wide; diffuse shrubs or trees to 5 m tall. .... *4. tetragonophylla Phyllodesscattered (not fasciculate),more than l mmwide...... 7 7. Stipules spiny (although absent from some nodes); phyllodes normally 3 6 cm long; flower heads pale yellow; bushy, non-resinous shrubs or trees 3-7 m tall...... 4.4. cuspidifolia Stipules not spiny, persistent, c. 1mm long; phyllodes 0.8 2.5 cm long; flower heads golden yellow; resinous, rather spindly and open shrubs 2-3 m .* A. maitlandii = lerete: legumessticky: shrubs dense. spreading,to lmtall...... 9 Phyllodesscattered, normally more than 1cm long...... 10 9. Phyllodes less than 5 mm long, ascending,grey; branchlets densely white-pilose; petalsstriate...... ,. , , *A. adoxavar, adoxa Phyllodes more than 5mm long, spreading (at maturity), bright green; Branchlets not conspicuously hairy; petals obscurely 1-nerved * A. spondylophylla 10. Phyllodes minutely punctulate i.e. with minute, circular, whitish dots which are observable at x10 mag., nerveless or very obscurely nerved; flower-heads cylindrical; legumes woody, basally narrowed; seeds oblique with straight, narrowlyturbinate funicle/arils...... 11 Phyllodes not punctulate, nerves often conspicuous; flower-heads globular to cvlindrical ...... 12 11.Phyllodesflat...... A. arida Phyllodes terete (very closelyallied to the above) ...... *A. orthocarpa 72. Phyllodesterete,subtereteorquadrangular ...... 13 Phylkrdes distinctly tlattened (although occasionallyonly 1-2 mm wide)...... 21 13.Longestphyllode less than 7 cm...... 14 Longestphyllode more than ? cm...... 16 ''" ur I uEql 'reauq aJoru squqs IIeI:aôqE uBql Ja^'\oJrBupue .ra8uol ,{1no.r.reusapollfqd '' vsnlla 9' v 'pauorlrsod 'aln)E ul I ol sqnJrls Surpeards-epr,n :oJcnlu elnD€ .{ll€Jalel u€ rllr^\ 'apr,,$ '3uo1 'f,Ildllla papunor xadB urlu 1-g urur 9I-6 dllpculotutu,{sB sapoll,{qd 8A " " ' t2" ..IqclIU IuuItrAJ,,lou IrEq :rBlnqo[8spBaq ra,$old ' ' 'alElnJrlar 'alBurJ tZ'' snoursal + ) saurn5al:pua qJ€a ruoq,{lasrorlar IJnJ qcrqarsFur,reqs âorrBu ur SurlBtloJxapuB qslppo|a'l IuultrÎ,, ,,lqrl'Iu {JBq:(srBrs .ZZ ra,rou ioJ alcB!.docoJ lcac{sur sueurrcads 3ur1rnr3 uo) IEcIrpuII^) sp€aq-.I4,'!1old " " '(suaurrcads 62" parJp uo pa^Jasqolsaq padolaâp {Irood ulaql Jo auo qlrd\ 6 ol pecnpal so^Jau ?soua^rq y ur :pacBds dlapl'!'\ ',\\al 'snoJaulnu pu€ snoncrdsuoc saurrlauos lnq raqla8ol asolc pup ouIJ {JaA dllEuuou so^Jau) ar€J qJ€a uo a^rau IBurpnll8uol I upql arou qll^\ sapollfqd " " ' " ' ZZ (drp uaq,L\pal{urft ^laurJ uaql sapoll^qd aqtt'aenop!^ 'V. 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' . . 'snorqE[8 LI. sdB^\lB s]elqcu€rq aql q]l.^{ .raqla8ol 'IIB 'JElnqolE ur pa^rau-i sapol^qd :saruoJeJ^Jel|x€ ur paBuBJrB spEarl-Ja^'\old9I 'V*' 'llel '8uo1qo '{>1ct1s ernauy uI I uBql arou soarl Jo squqs :apr,$ulur gI-t 1 '^Joded 'alqpuB^ lou saurn:al luaaJg,{ar8 sapoll^qd :8uol ulr I ()1 salJunpod 'V 'snoulsar BUz1M * IIBI ur I ol sqn.rqsButpeards :paua{rll{l 'apr,,'\ 'J 'pa^\oJrBu '(8uno.{ sur8JBtu ulur t ,{ll€sEq lnq r€aurl uoq,\\ lsBal '{poo,ld 'Buol 'gI lE) Îcrls saLunFal:uoar8 urc a o1 sepollfqd i3uo1ure g-1 salDunpod ' ' gI""' IeJupurl{JspEaq-Ja,\rold 'I ' '' 'llE1 eJrarrvv 'prcsr,r '8uol rlr I ol sqnJr{s SurpBards {11q31ys :spJ€d\dn pa^rnc luc I 01 'apr,^^ 'pan\orJ€u '8uno,{ sapo11/{qd:poua{)lql 1ou surS.reru rrrur g-l l{ll€sEq uaq^\ ',{poo,,$ 'fflnqol8 ?socluraA sauln8al :8uol urc } g I solDunpad uo spBaq ra,\\old ]I (dlcr"rEH.qr,', p..,p.,t' dBUEU url-rA U B a

70 NuytsiaVol.4, No. I (1982)

Phyll"de, all lessthan 5 cm long. Phyllodes some or all more than 5 cm long. 25. Stipules spiny on young plants, often deciduous with age; branchlets not resinous, often pruinose; peduncles normally in pairs; phyllodes never spiny tipped, often finely wrinkled when dry; shrubs or small trees...... xA. victofiae Stipules not spiny, persistent, appressed, c. 1mm long; branchlets resinous, never pruinose; peduncles solitary in phyllode axils; phyllodes normally spiny- tipped;rather spindly,open shrubs2 3mtall...... *A. maitlandii 26. Phyllodes long and narrowly linear, less than 5 mm wide, normally terete but appearing flat when broad (seelead 17 above)...... A. sclerosperma Phyllodes more than 5 mm wide...... 2'7 27. Peduncles less than 1cm long; phyllodes less than 10 cm long; attractive, fragrant shrubs to 3 m tall, commonly forming groves along roadsides, trunks white... -.-....xA. validinervia Peduncles more than 1 cm long; phyllodes normally more than 10 cm long. . . .28 28. Legumes t chartaceous, 10-17mm wide; bracteole and sepal apices densely golden puberulous (best observed at x10 mag. on unexpanded heads); trees to 10 m tall, not confined to watercourses,llowering from October to December xA. (headsbright golden) ...... pruinocarpa Legumes I woody, c. 5 mm wide; bracteoles and sepals glabrous; bushy shrubs or trees confined to watercourses,flowering from May to August (headswhite or cream)...... *A.ampliceps 29. Phyllodes large (c. 10-20cm long and 1-2.5 cm v"ide) AND the minor nerves forming a loose, open reticulum between the 2-4 main longitudinal nerves; flower-headscylindrical ...... 30 Phyllodes shorter or narrower OR if not, then the minor nerves very fine, clc,se together and parallel (nor forming a loose, open reticulum); flower heads globulart., cyJindrical 30. Petals glabrous (best to observe unexpanded spikes); phyllodes :l symmetrically and gently falcate. . +A. cowleana Petals minutely sericeous; phyllodes t straight although normally curved near *A, their apices,somewhat asymmetric ...... holosericea 31. Flower-heads globular or obloid i.e. less than twice as long as wide (on fruiting specimensinspect receptacle fbr flowerscars)...... 32 Flower-headsdistinctly .cylindrical i.e. more than twice as long as wide...... 37 32.Phvllodes with numerous, very fine, parallel (never anastomosing) nerves which aresituated close together...... 33 Phyllodes with 2-5 widely spaced main longitudinal nerves between which the minor nervesfrequently anastomose and form a reticulum...... 3,1 33. Flower-heads globular; legumes rnoniliform, to 23 cm long, 6-13 mm wide; phyllodes t linear, 12-35 cm long, 1 7 mm wide, lax and olten pendulous; trees to 10m tall and normally confined to watercourses...... *A. coriacea Flower-heads obloid; legumes flat and broadly linear, to 10 cm long, 4-6 mm wide; phyllodes very narrowly elliptic to narrowly oblanceolate, 6-15cm long, (4)5-1C(12)mm wide, ascending; shrubs to 3.3 m tall, favouring open spinifex plains 34. Branchlets and phyllodes hairy (sometimes the hairs are very fine observe at x10 mag.); inflorescencesnormally not racemose. . "Minni Branchlets and phyllodes completely glabrous; bark never Ritchi" (s)-6s99t 'aII ' " "'PIIoJo:) tLpre,aolsY* oql Jo r{l8ual aql JI?I{ uPql ssal x,{lEr :oôq€ uPql asuap ssolpup naolla,{Jelrep E saIIds:J€auII ,{ln orJBu ^llBuors€tJolnq lIldITa {l^\orr?u '3pL!\ '3uo1 'alqBuE^ ^ra^ ro rBaurl ^lpPorq ualJo urru 8-I urc 9'61-9 7 sapollfqd ' sue4tnspe pII Bllorof, oql r{1Sualaql JIErl u€ql arour x.{lEc :,\\ollaf al€d Vx Jo '5uol 'asuap sa{rds :uaarEol€d'J€auII {lâorrEu'apl^l ulur t-Z luJ 0a-II sapoll,{qd 'snoacBlJBI{J 'snoulsal gl ' ' ' ' ' + 1ou saurnSal:aôqP sB 1ou sapoll{qd 'Ipoo,&qns'aplt\ edJe)oris2ue 'y*' ' '' urut AL'snoulsar 'saArau sauln8al:rlsl^\olla.{ puE olorJSIpalrau roulur'}uE}sIp passardurr leurBreur 'apl^\ 'aJntsqo ,{ra^ oql uPql luop-Iâ aJour sa^rJu tlsr^\olla,{t I urul II-A puB 'uaar8 'zv Suol ruc 8I-0T't'(rlsaJJ a^rasqo) ûlqs }Eq.\\auros aAITolq8uq sapo11dq4 " 'pallul saunBal zl" \ ^lsnol^qo lou ',{IalllxP '1q8rer1s :asoruatBJ lou sallds :oplt\ uIuI ZI oAoqE 1ou ualJo sapoll{qd 'snoulsar ',{rp uaq^\ pal{ulrt ypLLUni V*' lou ,tiieurpnlrauol ^lsnorlqo saurn8al lslooqs ^\au sB Jno ,to.rB flleur.rou qclq'r'r 'apl^\ ti-ainr ,t, pi3.,er.re so{rds :asoulnrd ue}Jo slalqcu€rq aql qll^\ raqla8o} 'aletlPJ-alBloacuPl sapoll^qd 'lv uIuI 0Z 0I puB Suol urJ 9I 0I lnoqB ^lsnon)Idsuor " " " " w ('8BIuoIX lB pa^rosqo-passardde pue alnurur ualJo sJIEq) aar8ap auros ol ^rter{ saurn8al Jo (slooqs ^\au uo fll€Icadsa) slalqcuErq Jo/puB sapoll^rld 'salcEldaJaJ ro salJunpod 8utJa,!\ou ral{llg " " ' '0? In (x,{lBr ]tal8au) sno.rqelEs1.red 11y '3uo1 gn....-. tucg ueq] ssalIIE sapolldr{d . . . . . '68 0n. 3uo1 ruc g u€ql aroul llB ro aluos sapolldqd " " " aôqBs€ 6t" Paulquoc lou srollpJEqr raqlo uaql (ar€J) os JI Jo luonuuoJ flles€q lou sa^rau apoll,{rld 'snolnJaqno e[uapeJcY'v;''''' ,{1a1nurur puE snoulsar slsltlJu€rq :asPq aql aôqE acu€lslp E JoJ ur5rPtq 'saJlde .rJ,ro1 aq1 qlrrd luanguoc .{lsnolAqo salrau auros'o1€tl€J lou al€ulrunJ€ 'apl^r 'D qlr.,r\ crldrTla ^lÂo Bu ,{lenbyqo urr t'I puB Suol ruJ 1I-9 sapoll^qd 88 "" gg ""' aôq€ sBpaulqurot lou sJalteJsqc '' edMctrq)e4 'V 'ZI' ' ' ses.rnoc.rale,rA8uol" punoJ.{llEuuou soarl IIBurs Jo sqnfl{s :puo qcEo urorJ ,{lasroJlar IJnc qtlq^\ sSuIAeI{s,t\ouEuuI Eull€llo]xa qr€a puE qsrppa|a l ,,lqJllU ruultr{,, {rEq:alElncrlar puB alBuIcrIJ sauln8al:aJEJ 'pl8lJ 'apIA 'JPaulT sapoll,{qd uo sa,L.rauIBuIpnlISuol t T lou uIEI U-I ^l,t\ollau lt " "' .'snoulsaJ eqayqdotlcrp V* 'snoocEl.reqc sqnrqs :apr,{\ urc A'I-I flur. g saurn8al :seAJou IEulpnll8uol g-Z qlla,!'snoacBuoJ'al€Incllar ^lsnoncldsuo, sapoll.{qd:esolllaJEr lou sacuaJsaJoUuI " esoua\tq 'Y '8' ' ' ' snoulsoJ 'fpoom sqnrqs :apr,r ruc I uBql ssol sarun8al :(padolaâp ,{lrood lou '.{rp ',{qsag sr asaql Jo auo lnq) so^rau leurpn|8uo1 Z qll'$ uaqd\ pal{uF^\ flaug 'alBlnJrlar,{lsnonJrdsuoc lou sapolldlld :{JBIIIXB SuIJPadde uaql sallunpad aql qlrn sloor{s ,{JEal sB lno .,r\orF^ eulrou sorua3BJlnq asouraJPJsacuocsaJouul 9t "' " ' ' eloluuourv; pua qcee urorJ dlasJorlarIrnJ qJlq,'r\sFur^sr{s 'llB1. rdorJ€u u,r SurlBrToJXapuE u^\oJq qslppal a'I r.rqcllu IuuItr tr,, {JBq lu I 'snolnJoqnd 01 saarl.Jo sqnJqs:aôqB ueql snonlrdsuoc ssal dlauIJ urnluaunpul 'apl,ld '8uol :passardur puE auu urnlncllar Jo soAJou urtrl gI-g IIIJ t-A I sapoll,{qd " "' ' '' ' ',,lqc1lu erua\tla|v,t, luuttrlr, lou{rBq 'll€1 ur I ol. sqnrqs uado la8rel speaq-.ra,tou :asoluauol ^losuap pu€ snonldsuot 'apt^\ '3uo1 urnl.uarunpur :alBlnJller fl5uorls urc 1-6 UIJ g'l t sapollfrld 9t

,{alsraruEH aql uI ErtErV'uIIsEt{ IL eBup8 U B I7 72 NuytsiaVol..1, No. I 0982) I 44. Legumes glabrous, to 21 cm long; spikes comprising somewhat widely spaced rather large flowers with short, cupular, t truncate, golden puberulous calvxes that are readily observable at x10 mag.; phyllodes coriaceous,rather rigid and normally 1 straight, extremely finely nerved, silvery-sericeous when intermediate aged but glabrous at maturity, very young new shoots golden; small, spreading often gnarled trees on alluvial flats ...... 13. A. xiphophylla Legumes hairy, hairs, yellow when young but turning white with age; spikes dense; calyx not as above; phyllodes often falcate, generally less rigid, less coriaceous and more obviously nerved (at x10 mag.) than above, normally 8- 14cm long and 5-15 mm wide...... 45 Legumes with a dense, soft appressed sericeous indumentum; flowering peduncles and spike axes (i.e. the receptacles) golden puberulous; phyllodes falcate, the new shoots citron-sericeous but hairs turning silvery-white on intermediate aged phyllodes, glabrous at maturity; graceful trees often with pendulous branches, normally found along watercourses. (Small phyllode variants, c. 5-6 cm Iong and c. 5 mm wide, are known frorn tops of rocky ranges)...... iA. citrinoviridis Legumes with spreading, pilose hairs; peduncles and receptacles sparsely resinous-papillose(not golden puberulous); very young new shoots pale citron- sericeous,quickly passing to glabrous phyllodes; spreading shrubby trees to 4 m, normally on ridges and upper slopesof ranges. . . .8. A, iamersleyerIsrs 46. Sepals free and linear-spathulate; legumes chartaceous, I glabrescent, loosely reticulate; phyllodes very variable in shape and size, grey green! together with branchlets minutely appressedpuberulous (at least when young); tall shrubs or treescommon on alluvialflats.... ,. , ,.*A. aneura Charactersnot combinedas above;sepals normally united...... -. -.. - -17 47. Phyllode nerves rather widely spaced (interstices readily observable and wider than diameter of the nerves); phyllodes about 3-5 cm long and 5-10 mm wide; legumeswoody and glabrous;uncommon shrubs.. .,...... 48 Phyllode nerves very fine and close together, either uniform or 1 or more slightly more evident than the rest; legumes chartaceous to coriaceous, hairy or glabrous...... 49 48. Phyllodes and brachlets glabrous; legumes not wrinkled, narrowed towards their base, 6-8 mm wide, to 6 cm long; phyllodes with numerous, prominent, longitudinalnerveswhichrarelyanastomose...... *A.ptychophylla Phyllodes and branchlets densely and minutely appressed sericeous; legumes coarsely wrinkled when dry, narrowly oblong, 1-2 cm wide, to 16 cm long; phyllodes with rather few, not prominent, widely spaced main longitudinal nerves with sparsely and openly anastomosing, longitudinally oriented lateral nervesbetween them . . , . . , ,,,.,.*A. cuthbertsonii 49. Branchlets and/or phyllodes (at least when young) and llowering peduncles hairy (observe at x10 mag.); legumes hairy or glabrous; bark never "Minni Ritchi"...... 50 "Minni Branchlets and phyllodes glabrous or if sparsely hairy then bark Ritchi" i.e. red and exfoliating in narrow shavings which curl retrorsely from each end; legumesglabrous ...... b1 50. Low shrubs to 1m tall; young legumes with sparse, short, spreading hairs, mature legumes not seen; stipules frequently persistent on branchlets, dark brown, 1-2 mm long; phyllodes (3.5) 4 5.5 cm long, 6-10 mrn wide, very sparsely reticulate; new shoots not golden hairy; spikes dense. . . .5. A. daweana Tall shrubs (exceeding 1m) or more comrnonly small trees; stipules early deciduous. . . .13. A. xiphophylla (or a short phyllode variant of A. citrinoviridis, see lead 44 above). 'poua{crql 'aJuacslqap |ou sutareLu :uorlJod IBcIdE uallor!\s € I{11,!\plolu8ls ,{l^\ollBI{s 'do1 'paArou s'u,ano11ojpa.\rnD soAIEA oql urorJ ,tllecllsela Butllllds .{11eu1pn1r3uo1 'snorqelS 'fpoor Bunol-'uiq,lr asocluJoA 'u,t\orq ol urdorq qsl!\ollo.4 IrBp reseq 'spaas '1urod apl^\ tuur g-t '8uol ru, q g-t aql spr€,1dol€ulrad€l aql lsapeb.rq lp 'pa^rau- 'snoJqElS ,t""^1aq p"1","1s.toc-l.ou'possaJduroJol alaJalqns'lcaJo saurnta? I 'ql8uol 'c '3uo1 rlaql E/z roJ aleuuoc urul I z slepd sa^rau uaa'|AJaqanssrl luacnlsueJl qlrr,l pi.trau-g ,tl8uorls aqnl x.{ler'soqol reln8uet.rl dlpeorq olul paplÎp dllroqs dra^ 'snoraur-g 'aBulursl 'e11oroc q13ua1 x/fal 'a1nJ3 dllunlq spnq sra^lold ;o Elz-z/I',lr1ol1 'snorqe13'8uol paiagur p.,e paua{Jrq} ut Sutpua pue JBauITsalell luul I l saloaPeJq 'slsaqluE -J 'pora^dou :pa8ue.r.re,{lasuap sr3^\ou 'snoulsar allnb }E yBIp uI tu UI-I '.relnqol8 '(8uol -q7-6g "ep1oe 1i13q speaq-ra^\olJI JualslSrad + puB rEInBuEIr-] 'c) ',{rB}ITos 'snorqEl8 'lsnqoJ urur B 0 IIBtus perq JelnJanpad Jesaq :snoursal fllqSITS '8uol 'lJara 'apou 'frBIIIxE saJUa)saroUUI urur 0t-gl salJunpad- Jad Z-I puB aldurls 'urElp 'luasqE a)glro luur U'0 c anssrl snoJaJllJau qsl.^dolladpasl€r EulslJdurot 'snu$1nd uo palPnlrs aql Jo pua I€lslp oql 1e apo11,4qdaql Jo uIErBur.raddn aq1 'arncs(o pua1,3:pa8raurqns 'a1qrsr,'i dle.req'8 sa^Jauisnolll^,{lasuap eaoorS'aseqreau 'pal{ulJ.^,\ '8uol flprxepa paAooJB 'aBuBJo qsr.'rro11atr dlasJaAsuBr} [laJnJsqo urur 9 0 'j-'i.rnisqti 'palooq snur lnd 1:uol ururI c ore l{rlq,rl slu,Iod u,t\oJq-a8uero1q5t1 'snoncouul 'alnJ€ 'asollBJ [11q3qs ro 1qi3re.r1s olul xade le pan\orJ€u ^lldnJqe :uaad -iai8;snu,,o1nd aql roJ ldaJxa snorqBlF'lcadsBSuIpuaJse cllslJoI)BJBI{D e arri o1 aseq 'utBtp '8uol 'EuI.{Jp aql rcau sp.re,,rrdnpa.t.rnc urul I urJ t-U (g'I) uodn passardutor 'snonplrap 'snoulsar 'uaar8 Illq8qs 1nq atroral sapoll,{qd sa1nd.r1g [11q8qs urnrpau 'slalqrueJq ol lq5rT s?ooqs ,.r.rar1-sno.rqe18 aql Jo spua aq1 sprEidol luaplâ lsour 'alaJol arp qcrqr\ sqlJ snoursal pal€lnuaJc qll,t\ saqJueJg saqcuPJq Jo spua sPJ€'|Aol 'parnssg u.nnorqrisrpp-Jr lnq.4oJt fllr{8ITS sI 1t oraq,lAsulals Jo aseq aruaJJxa18 ldacxa 'sorlJuBrq ()1 SuIpIAIp 'urBrp tu puE qloours rypg ul€ur 9 dn olur loAal punoJ8 ]€ A 'peddol-1eg 'Sulpeards 'asuap "$o'I 1le11 o1 sqn.rgs prcsr,r {11q311s + ro papunor '(Tgnl'y'ggy3 :osrlHJUgd :oloq) ,,'PUPrlIrq"Ire)V tqlr.al'xaflurds ul{€l) ûols 'pa^rnJ 'uaar8-.{ar8 o-"?i1 p"n i"l.ta,r."tog.tt :SulpuaJse trl]q8lls sapo11{qd 1""r'a- 'peddol-lBu :,{aI3 isaqruErq ulPur 9 olul 1a.ta1punorS 1e Sutptrtp + puE {rEq 'llErl 'ltgl Surpee.rds saurrlauos rJrI quqs papunoJ asua(,, utlsehl U g 086I Înf .Erlprlsny 'puBlpaH g ru)i ;sno^J zI uralsad\ lrod ol p€or al{l uo uloouollllA Jo II

'lraJJIpsrlBssPrJuI Jau slulurn3al snqrur8reur1a snqlrPlnqol; snqIIEJoU srlplrdBJ Enb E srulJJE uopEtr^tr "uErlf.l sDe)V pE luEllg UaJJlp sllP^rnl urnsrns 'snqr.relnqolB '".rqi1"r"1 'rue,p uru 'sriuol tuc 'sllelnlcund rau sllpollfqd 1 7-6ig'I) 'v' 'qluag snqrTBrou srplldec enb P SIuIJJB llantr l i edrnoqlro la eplre ercYJv (Z arn8rd) 'Aou ds'uIIsEIAlslcarJB BrcBcV I

" " " 'opl,\duflu eu7adaol Vx tII tl-t ollBJ qlpl^l 01 qlEual (lI)91 'JIldIIIa -? '€uol ruJ (6)9'L-8 .'{l,{rorrEu1 sapolfqd :salun8al aq} uI anbllqo .{lasra^supJ} ol asJaASu€Jl spaas :apln\ urur 9Z-0I 8uo1qo flpeorq saurn8al '0t-6 'apln\ '5uol upre,t\oJl y* aTI ' ' ' or1e.rqlpr.tr o1 q18ual turu 8-A urr (9 ZI idf) 'Drldrlla fl^\o.r.reu01 rBeuII'alqPuEA sapoll.'{qd:saurn5al aql uI anbllqo O-Sl '8uo1qo ,{11euipn13uo1 ol IBurpnlISuoI spaas :oph\ urru 1-7 ,{l,t,t.oJJ€usaun8aT Ag - - -...... Elloro? qFual3ql zg. aqlJo jpq ueqr ssal ro x,{l€? :a^oqe uuql asuap ssal ualJo sp€aq-ra,\\og :'{ar8 {rE€[ Jlerl 'leurpnlr8uol irciqaoioqt y* pff ' spaas:ap!,t\urut g r'Euolqo .t1,rio.r.reusaunSal ielloroc al{l Jo ql8ual orll JI€rl uBql arour ,{llq8lTs x,{lBr :asuap '3uo1 spPaq ra,\\ou :apl,|\ urur I 9 urr g 9-q ' sapo {qd :,,IqJ1IE IuuItrl,, {rBg Iq

a;upg,{alsraupH eql uI PrtErV ullsElr\l U B tL a

14 Nuvtsia\ ,,1.4, N,,.I (198:)

Seeds positioned obliquely in the legumes within pronounced depressions, rarher persistent following dehiscenceof legume, ellipsoid, 3 mm long, 1.5 mm wide, turgid but laterally compressed (2 mm thick), blackish and rather dull, marked with a darker coloured narrow peripheral band of lissue; pleurogram fine, open towards the hilum, bordered by pale coloured tissue; areole c. 1.7 mm long, 0.6 mm wide; funic?e all narrowly turbinate and straight.

5mm ,\T ffiffi "W "W

2mm /---n

ffin lli\\ W ftl ,,...-.' ,W ffio''"- t\

Figure 2. Acacia atrecta. A Portion of branch. B and C Seed (B-plane view; C side view). D Legumes held erect on receptacle (note pe$istent seeds following dehiscence). E-Flower. F Phyllodewith €nlargementsshowing apex and transversesectional shape (note positionof the 8

A fron I. L. Lethbriclges.n.; B l) from M. L H. Brooker 2089a;E from K. S/ewart's.n.; F from B. R. Maslrl .1614(the tvp€). Other collectictnsexatnined. WESTERN AUSTRALIA: 15mi (24km) E of Nullagine,J. S. Beard 4603 (KP); Millstream,6 Dec. 1974and 10 Oct. 1g7b,R. F. Black s.n. (PERTH); Southerntributary of the FortescueRiver jusl E of Gregory Gorge,30 Jan. 1975,R. F. Black s.n. (NSW, PERTH); About 0.5mi (t krn) SE of Millstream Slalion homestead,M.I.H. Brooker 2089a(CANB, K, MEL, PERTH); 2 mi (3.4km) E of Yampire Gorge, 22'20'5, 118'30'E, H. Demarz 4416 (Kp, PERTH); WittenoomGorge, Hamersley Range, A. S. George1068 (PERTH); 10mi (16km) SE of Wittenoomon Drillers Hill, July and Aug. I97I, I. L. Lethbridges.n. (BRI, PERTH); ,10km E of Wittenoom, A. A. Mitchell ll48 (PERTH, WAIT); WittenoomGorge, Sept. 195?, I{. Stewaris.n. (PERTH).

Distribution. (Figure 10) North-westWestern Australia in the FortescueBotanical District (1;250000 mapsl'50-6, 7, 1,1,12; F51-5). Ranging from Millstream (21.85,S, 117"04'E)east to near Nullagine (21'53'S, 120'0?'E) and extendinssouth to the northern part of the Hamersley Range around Wittenoom Goree (22.14,S. 118"20',E).

l 'alqBuBA 'guol rrlc g'?T-g 'alBloaJuElqo ^l,\\oJIEu o1 cr1dr11e{1,r,rorreu dra^ sapolL[qd- 'snonptcap ',{lpixepe paue{Jrq1 '8uol urul q t 'reln8ueul salndrls o3B rlll/'r dar8 0 'sno.rqe18 S.tiuiociq 1nq u,rlo.rqlq8q ro u,$orq qslppal puB snoulSoJlBtlr\auros ^ll€JldB 'p€AJau dlornrsqo .{raA 'alaral s?aJqJu?JS sacuaJsarouul qlIM pslBlcosse uaql AJellIxB 'fJp 'snoulsal s?ooqs,,|!aN Jr .ro slalrlruErq uo lBuruJol Jal{lla uaq^\ qsluaoJq {JBp 'qloours qlBur 'qlEaurapun u.norq r{srppoJ lnq dll€uJ3lxa far8 asl/$Jaqlo surols Jo asEq 'suo-rlBnlrs 'lJeds€ lE paJnsslJ ylpg. ql,!\oJ8or ur fqsnq SuluroJaq flpulds E l{1li\ pu,E paururals-i13urJ fllBuoISBJJo 'la^al punor8 ruo.r; Surstre surals uIPtu ltara-SuIpEaJds 'c 'turpEards 'uad0 9 o1 dn qly puB IIel uI t t o1 sqnrqs rEInqIpunJuI ro papunoJ '(AN 'y 'gNyC :osr :HJUgd :oloq) ,,'slua:raura erclqdornal snld.tlerng q]I^l pUBISSE.I; '.{uo}s guI^\oJC Molla^ lcour*nq euEasr,\\elporrJ);3a1 do111rquo IIos u^loJq-pal uI 'uaar8 'u,to.rq s.rearolg llnp sapolldqd sruals 1s33rBI Jo aseq 18 snorqlJ pue qBnor 'q}oorus 'llBl r{srppar {rEg llq€q Sutpeards urZ'A qsng,, 86tA uaEpnry wloJ[eIN '086I 'BIlErtsnV 'g,69.rII 'S,88.22 'p€or ()1ocnrg dEI I 0I uratsalA uloouallllA lI I aq1 urorJ (aJlrd uroJ ltr\l sprB^\ol) peo.r aurtadl4 ancsalrod qlnos SuolE ru1 1 :snd'f.tr 'sIlBI urur g-t snqrururn8al13'(sIlBI tuur 81o s13uo1urur 0I J rsaqluB qns) sleprolqo snqrlBrol+ srInlIdBJ :lraJJrp auoll€ulqwoJ uI srpuncas snqlJolterEq) ',rqJlru eDecv rIIallo ruultr l, uou sloJautt acllJoc enb E SIuIJJEuITSEtr l ErolqdopoLlr

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'aBueu {alsJourBH aq1 JoJ papro)er 'V 'epLJe ,{lluaululoJd IIE aJE auarJliq pu? edJyJoqiro V ErratY su-I8rBu pauarlJrql s1t ^q paqslntunslp 1ee1 qarqar saurn8al slt dq oslE puB sp€aq-ra.Âou .re1nqo13 .r,eie sr lnq uaprel4tr euvryUq V salquasar salcads aql ^Soloqdrout apo fqd 'ralroqs sepo11,{qd alElcund I€raua8 pu€ llq€q uI Jo ra,\\orrpu Jaqlla arE qcrq,t uou slr puE speoq-ra,t\ou (l€JupulT^r lou) r€lnqol8 sll fq paqsln8ullslp ,{Ilp€ar 'y sr elcarre I/ lnq IIanI l I edrecoqlro puB qluag aprB y sarJads polElar ,{losulJ aq1 esoql sB surPs aql fllBlluossa arB saJnleal lsJISolodJBJ asaqJ IIJe-alJIunJ Jo 'pa^\or.rBu alpurqJnl ,{l^\oJIPu 'lqEtel1s s1t pue saurn8al pa^rau d eutpnllEuol {llEs€q 'a1a.ra1 'lJara 'lJara 'a8re1 1 ^poo,$ sll plaq oJB qclq,,'\ sp€aq roddou .re1nqo13 altnb 'lleurs slr 'sapoll^qd Surpuacs€ 'alaroJ sll tlllM sarJads J,^-IlJullsIp e s vlJarJY e:JeJV 'ôlpad ' ,{q pazru5ocar lou sarJasIErnlBuun uE qluag ('qluag) SaIUJoJIUIBIBCsalJas uI rnJco plno,lr sorcadsaql uorlBrurssElJ (iggl) s.ur€qluag uI aq)]ag la uapIBI l ('qluag) 'V 'uoIlEcIJISSel) i5,r"aur.rn14uorlaas uI parBld sI elJa.rre (8/6I) s,âlpad EqISn

'saurn8alSuIdola^ap oslB puB (sISoqluB lp puE pnq ur) sra^\ou qloq ssassod,{IluanborJ Jaqualdas pue ,{lnf uaa^llaq pa}rallot suaurrJads Jaquraldos olel ur palcalloJ uaaq aAEq spaas aJnJ€ur 1111,$sa1unia'I 'lsn8nv puE raquaca0 lnoqe ruorJ sI pouad urBur aql lnq rBa^ aI{] Jo stlluour lsotll Suunp luasaJcl 3rE sra,\\oll aulos qFnoql sE srBadd€ 11'pouad 3u4rn4 puv SuLJa'$oli '( ds arpouJ ) {lJor ,xaJrurds, .{q pelEuFuop eddals-qnJl{s ut sleg duols pa}EIJossEpuB slllq n oI srnoAEJsalJads aql'pu€q o1 suollJallor ,ldoJdlaAll€lar aql ruoq Eutapf.f leiIqeH

pr.,pJ!'iurtcel{ rSueU (rlsrrrupH dqr ur 8 B 76 NuytsiaVol. 1. No. 1 (1982)

(4)5-10(12)mm wide, length to width ratio (8.5)10-20(26),slightly curved, ascending, coriaceous but not particularly rigid, glabrous, characteristically pale grey-green, reflecting sunlight in a I silvery fashion, resinous (resin layer very thin and not sticky); apex possessinga brown, obtuse, knob-like callosity which is often slightly recurved, callosity very pronounced on young phyllodes; pulvrnus 3-5 mm long, transversely wrinkled, yellow to brown; gland situated on upper margin of the phyllode at the distal end of the pulvinus, lamina swollen about the gland, submerged, comprising a circular orifice c. 0.3 mm diam. and a very indistinct rim; longitudinal neryes numerous, very fine and close together, not anastomosing, the central nerve and normally one on either side of it slightly more evident than the rest, margins yellowish but not thickened. Inllorescences axillary, 1 2 per node, normally arising at extreme base of a rudimentary shoot which usually grows out. Peduncles 8-15 mm long, rather stout, glabrous; basal peduncular bract deciduous, triangular, c. 1-1.5mm long, concave.FLower-heads obloid, c. 10 mm long and 8 mm wide at anthesis, somewhat resinous, c. 90-flowered, flowers densely arranged; bracteoles linear-spathulate, c. 1,1.5 mm long, lamina inflexed and resin-papillose (papillae white). Flowers predominantly 5-merous although a few 6-merous flowers are present in some heads. Calyx slightly exceeding half the length of the corolla, gamosepalous,stout, very shallowly divided into broadly triangular resin-papillose (papillae white) lobes which are visible in inflorescencebuds, calyx tube I obscurely 5-nerved and glabrous. Peta,lsc. 2-2.5 mm long, glabrous, 1-nerved. Legumes broadiy linear, to 10 cm long, 4-6 mm wide, slightly curved, t cartilaginous to firmly chartaceous,wrinkled when young, flat, barely raised over seeds,glabrous, slightly greyish brown, somewhat resinous but not sticky, abruptly narrowed at apex, basal stipe c. 5 mm long; marglrs slightly thickened, not constricted between seeds.Seeds longitudinal to very slightly obliquely placed in the legume, narrowly obloid or sometimes narrowly elliptic, narrowed at hilar end, 4.5 5 mm long, 2 mm wide, somewhat compressed (1 mm thick), dark brown but areolar area yellowish, not mottled, shiny; pleurogram obscure, open towards the hilum, surrounded by yellowish tissue; areole less than 0.5 mm long; funicle c.3 mm long, flattened and membranous, gradually expanded into a convoluted creamy white ariJ which extends down c. 1/3 the length of the seed. 'I'om Selected specimens. WESTERN AUSTRALIA: About 1 km S of Price airport. K. Atkins 1209 (CANB, PERTH) and 1213 (K, PERTH); Marandoo, just S of Mount Bruce, Hamersley Range, t 22'40'S, 118.09'E , C. Dawe M154 (CBG, pERTH): Rio Tinto Gorge, Hamersley Station, H. Demarz DbTbB (KP, PERTH); 6b km W of Millstream, H. Demarz D7084 (PERTH, TLF); ?5 mi (120 km) from Wittenoom towards Roebourne, B. R. Maslin 2733 (AD, MEL, NSW, PERTH); 14 mi (22.5 km) S of Robe River crossing on North West Coastal Highway, B. R. Maslin 2761 (B, BM, G, NSW, P, PERTH); Hamersley Range National Park, on the Wi enoom to Juna Downs road 7.2 mi (11.5 km) S of turn-off to Mount Bruce, M. E. Trudgen 2b0B and G. Marney (BRI, PERTH).

Distribution. (Figure 10) North west Western Australia in the Fortescue Botanical District (1:250000 maps F50-6, 7, 10, 11, 16). Extending from the vicinity of Mount Bruce in the Hamersley Range (22'36'5, 118'08'E) north west to the North West Coastal Highway around Yarraloola Station (21.34'S, 115o53'E).

Habitat. Favours rocky loam on spinifex (Triodia pungens, T. wiseana) plains and is associated with a variety of species including Acacia ancistrocarpa, A. aneura, A. pruinocarpa, C.assiaspp., Eucalyptus leucophloia, etc. Acacia atkinsiana often forms dense regrowth populations in disturbed sites such as along road vergesand in burnt areas. .IILZ ullsslN g trroJ4qlggLg(IzisuaT'H lttor! J'g'laurew'C pue 60qUl,atPn{ 3'WurorJV E 'q"o",q 'auBas"ltp 'ra,!rold-(I poes-C "-tta"1-g 1o uoluod-V soexv 8 ern8l{

t. lE *ffiv

eBuBU,{elsrrujeH eql ur BTTBJV'ullselU g iJ":;l,:::;i::', May to July, however, some have also been collected between December and March. From anthesis it takes about 3 rnonths for seed to mature. Fruits with mature seeds have been collected in September and October.

Acacia atkinsiana belongs to section Juliflorae (Benth.) Maiden et Betche and is a member of the A. stowardii Maiden group of species (see speciesno. 11 below). The new species is most closely related to A. rhodophloia Maslin from which it is readily distinguished by its grey bark (not red and exfoliating in narrow strips which cuil 'Minni retrorsely from each end i.e. Ritchi'). Acacia rhodophloia is a variable species (Maslin, 1980) and throughout most of its range has cylindrical flower-heads, a character further distinguishing it from A. atkinsiana. However, in the reeion of the Murchison River (which is about 600 km south of the main area of distribution of A, atkinsiana\ A. rhodophloia has flower-heads which are globular or obloid and phyllodes which are 3-8 mm wide. Flowering specimens from these populations may therefore resemble those of A. atkinsiana, but in addition to its grey bark and iti more northerly distribution, the new species is recognized by its generally longer phyllodes (4 8(9) cm in the Murchison River variant of A. rhctdophloia), iti lack of minute, ferruginous resin-hairs on its new shoots, peduncles, calyx lobes and bracteole laminae, and by its broader legumes (2 mm wide on the Murchison River varrant).

The specimen, H. Demarz D7084, 65 km W of Millstream, previously cited under A. rhodophloia (Maslin, 1980) is A. atkinsiana.

The speciesis named in honour of Mr Ken Atkins who provided much valuable field data and many specimensof Acacra from the Tom Price Paraburdoo area.

3. Acacia bivenosa DC., Prod. 2:452 (1825)

The Hamersley Range populations of this variable species are characterized as follows. Dense, bushy shrubs to 2.5 m tall, with up to 6 spreading-erect branches arising from ground level, occasionally single-stemmed and whispy (see pendulous variant referred to below). Phyllodes obovate to narrowly elliptic, 2-b.b cm long, g 15 mm wide, length to width ratio 2.5-6, bright green on new growth, fleshy and subglaxcous to distinctly glaucescentwhen mature, finely wrinkled upon drying, 2- nerved (the principal longitudinal nerve is t centrally situated and a minor second nelve occurs on the adaxial side of it). Inflorescencesracemose but racemesnormallv grow out as leafy shoots, the peduncles then appearing axillary, conilorescencl acropetalous,llower-heads rich golden, buds bright green. Legumes woody.

As discussedby Pedley (7977) A. bivenosais a member of a complex species-group whose centre of diversity is Western Australia. The Hamersley Range populations of A. bivenosa often have phyllodes slightly more elongate than normal fbrlhis species which according to Pedley are usually less than 3.5 times as long as wide. Seemingly when initiated the inflorescencesare always racemes of which most, but usuallv iot all, grow out as leafy shoots. This explains why on a single plant both racemes and simple axillary heads can be observed. No plants definitely attributable to the sand- loving species A. ligulata A. Cunn. ex Benth. have been observed from the Hamerslev Range area. This species is very closely related to A. bivenosa and differs in its generally more elongate, l,nerved phyllodes and in its racemes which normallv do (1.c.) lo1 qrojv out. Pedley treats A. ligulata as a subspeciesof A. bivenosa,however, I feel that until a thorough analysis of these two species and their allies has been 'ssalo^rau 'snoJqEIS osr.draqlo asollrd€d-alrq,$ {la1nurur ,{UEurFJBur fllP3IdB 'Buol 'snoluaqnd 'Suolqo 'peal{-rad\oU rulu g a r slelad {1e1nu1ut dllBcld€ {l,t\orrEu alSuIS 'q13ue1 'J € ur uaAaJuasard uorlErrE^ Jo a8uEr slql r1aq1Z,/1 toJ pallun flqBIJ€AJo aa$ 'sFlad .qlaual 'snorq€lt 'alBIotTIt Jo Z/tt/I spdas snoraur-g sra,alold' s^\€lJ aEu[uBI '8uol 'alplnqlpds -&aurI '^\olla,{ 'rBlnqol: lrlur I-9 0 saloal.Jerg para^\ou Zt-tZ al€d 'srsar{Ju€ 'snolnJaqnd speaq-]a^\oll 1€ alBal)BJqa asBq saulllaulos ro sno.rqe18 '8uol IIIru0Z-gI sapunpad 'srsaqluE qr€ar sp€aq aql alull aql Iq {llrnleru qJEar 'slooqs sapollfqd Furpualqns asaql ,uau IBurrurol 3ur,no.r8 .{1a,rr1re8uo[B sapolldqd rapou 8uno,{ Jo slrxB urql-r,!\ palpll.IuI seJuacsaJoUura l snolsladoJcB sacuaJSoJoUuoJ '^rEIIrxB Jod 6(I) puE eldrurs soJUaxsarolJul oranu IBcIdB al{l ,,!\olaqulSreul raddn aql 'luaultlloJd '3uo1 uo luesard ^llaurJou puElS (JallBurs) IEuollrppE uB 1ou dt1 urur I-g'0 ':uolqo '1-r ol JEInJJTc o^oqB urur g ol dn Jo snur^lnd aql Jo pua Flsrp aql lB apoll{qd aq1 ur8reur Jaddn aql uo palenlls puBJ.8 :ulnlnctlal uado asJ€ds € SulurroJ puE 1o 'luauluroJd (fJp uaq,rl lseel 1B luar€ddB flrpPal la^) aurJ sula^ IBralEl qtlprar :8uol urul g 0 > puB arnDsqo snurapd 13uo1urur 6 r slurod IBJrdp u,roJq lq8rl lua8und 'luaJsaJqel8 ,{ldrEqs laf aurJ olur saJrdE lB pal.tBr}uoJ ,{l}dnJq€ + 01 snoJqEl; '8uno^ 'pa^rn) uaq,r uaaJ8lq8rl lq8uq'aJnlEru uaq^\ uaar8 aAITorunlpau {llprurou '(gA)qI-g rapl1l\ '8uol 'atqBIrB^ .'{llqBrTSo? tq8r€rrs = M/-I uulg-Z rur(gl)g-t(Z) JaqlEJ azrs 'alB^oqo fl,$oJJBu o1 Suolqo {l,t{oJJEu sapo '[qd sepou auros luoJl '.{urds 'alEJnpur 'lno1s '3uol luasq€'paAJncaJ,{llqEIIs puE SurpBaJds rutu t-e salndrls 'qsru^,!oJqol uaar5-daJg'snolnJoqnd dlasreds saurtleurosro snorqylS 'parrau flaut; 'alarc1, 's{unJ} slalqcLrerg urBur uo parnssrJ pu€ snoJqIJ {JaE.IIB1 ur I t saaJJpalJeuts 'urBlp rfqsnq 'asua0 ssal ro aJou o1 8u1,r,ro.r8 ur t c ol sqnJqs paqJuBJq qtntrl

'(IN 'y 'SNVC :osr:HJllgd :oloq) 'aeuoJ)t^ ,:esojuaurol euae4qcug puu e|etawo|E eDUaL'\e1 aIJaty qlln uoIJ€IJosse 'll€1 '1961 ur punou alorJlrs ^\ol B uo 8ur,$orc ru g t aarl-qns,, su1;f?Y Y 086I 'AoN rooprnqEred 8A ErlErlsnv uralsoM (aulul) lsBg t IIIH Jo qlnos ttl{ z :snd,ff .lraJJlp srueld srrpoll.{qd enb B sruIIIB eleput; lp.rep.td vtrvrv pE ruPllg lraJJIp snqIrBIIIx€ Jadlrlas srlluaJsarouur 'snqrlua8und srrpoll.{qd Bnb B srurJJ€ qlua[ aBIJolc[^ epe)V

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'uaI€lrapun 'y sr dnorB "soua^rq aql Jo IBSr€rddE IIEJoô palr€lap B Irlun IuBIJEA slql ol {uEJ 'slalqcuBrq lpurJoJ SurlnqrJltB uroJJ paurB4ar a^pq I snolnpuad .{llEJrlsualJ€Jeq) 'olBJrlap sBq puB Iaâl punorF 1e par{JuBrq dlSurreds Jo paluluals alSuIS sI luBrJEA ar{J ,{dsrq^\ roqler puB uado sr qJrq,!\ (uaas uaaq }ou a^€q spaas pue sarun.Sal 'y qSnoqllB) llqBq sll ur ^luo psoua^rq urorJ sraJJrp .{lgulwaos lI pa^rasqo leJrd,{l 'quou uaaq seq Bsoua^rq y Jo luErJE^ aATIJBJJ1E^Ja^ E aql o1 ru{ 002 auos lsEoJ aql uo JardurEq puE aFUEU-,{alsraurBH aql uI aclrd uroJ uaa,t\laq saJeld snoIrBA uI 'ardqMasld pdqsrlqnddq lJr,\\srlnqdJ drll pue uo{ElJapun Suraq flluarrnc are rpJ os paraloJsrp spuq,{r{ olqrssod aql Jo sos{leue 'dnor.3-sarcads uallod pue IBurosouroJqC srql ulqll,ld pa^rasqo sarlrxaldtlloJ aql ol 'suorle^Jasqo 3ur1nqr.r1uor ur rolJBJ IBSnBc€ aq dlqrssod ,{pur dlrpuq,{q lEr{1 suraes lr plarJ pelrurll uo pasPq acuoprAa pclSoloqdrour urorl gur3pnf sar)ads f,lqdrou^lod 'a18urs 'V ,{1q3rq E Jo sarcadsqnsparaprsuoc aq o1 a^€q plno,/r{'lpur1 euDtles pue 'y 'Tlanr\l -y urlsBtr{ sdarrldr{e d BLuJadsoJalJS uoâ sdBqrad puB urqan-I ruos,{? 'y 'qluag eurrlluex 'y ''qlueg EralllaFory sE qcns sarcadsror{lo uaqt 'palldd8 sr {uBJ JrJlcodsqnslI'sarrads lcurlslp sB tuaql lEaJl o} alqBraJaJdsI }I ual€lJapun

ojlueu ialsraruEHaqt ur EDpry'urlsEl,\I u g 80 NuytsiaVot.4, No. 1 (1982)

,"I L D't B.R.A. tq8t F$""M Figure .{. Acacra cuspidifolia. A-Portion of branch (note recurved spiny stipules and twinned peduncles). B-Phyllode apex showing spiny apical mucro with a small sland (s) at its base. C Flow€r. D Bracteole. E-Legume. F and G Seedshowing strap like funicle (F side view; G pJaneview). A,B from H. Dematz 2778;C, D from A. M. Asiby 4,1911EC from B. R. MasLin2,77I. alppqlJ lp.re{rrd V'rl1 palBlar dlasotr aq ol salJadsasaql Moqs osle puB dIqsuollelaJ aeLrol)r^ V etplLpdsnJ y asolt E lroddns ('uuroJ sJad) lauln0 r{d Iq salpnls qallod 'pa8u?rre [lasouraJpr Ja^ou aJE qcrq4r sacuaJsarogurs1r [q puB sapoll,{qd 'y lua8und slr [q avuolJr\ uroq par{srn;ur}slp ^llp€al sI arlojr.prdsn, PrrPJy 'sauln8al rlEU 'aEJEI 'sra.,r\ou snoa)BlJEqr Jraql pu€ sledas aarJ + Jlaql lo Jaqunu r€lrwrs € SurrPoq sp€oq-ra,uou ,uolla^ alBd 'rPlnqolS rroql 'saJuaJsaroljurpauul/tl [ll€ruJou rraql 'sapol[,{qd pa^rau-I rlBU Jraql salndrls ^urds Jlaql apnlJul salJads o,$l aql ^q paJEqs sarnlEal lecrSoloqdrour sso,rBluecr;ru8rs ar{J uoll)unf au,{o)sEC pue uoAJEuJBc uaa,,'\laq '5 a seJEId ur .{llB?rJ1€dur'{s,\\o.rg ol pa^Jasqo uoaq a^Pq 'pBaJdsaplM sor?adso^\l asaqJ qluag aeuolct^ V saltads alqErrB^ aql rllltr\ s?lllulJJe ]sasolJ slr sBrI puE CO aBaurpoll^qd trcos lo raqurau B sI BrJ.)IprdsnJslJsry 'aunf pu€ lrrdv ur palJalloJ uaaq sBq peas a.rnlBur :raquraJac ol raqolro urog sranaold 'pouad.3ur1rn41 pue 8uua,{ol[

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'uor8el eq1;o erog p/JE V.ar{l Jo luoruela rS,UIotZ snonDrdsuocB Jou sr pup (9,0'.LII oopJnqBJEd punorE) sfalsrourEH at{l Jo ,{lrulaJlx? uJ?qlnos aql ol ,{luo spualxa sarJads aqJ oguEu ,{alsrauBH ar{l 'S,SS.9Z) 'S,0g.gZ ot r{lJou puE (S,08.8II prrBqtE{aal I rBau oJ JSpa(g.iII r) .{€g '9 'S 'I-OSC '8I-0gd {rBqS lo,{lrut'r^ aql urors Surpualxa (8 :8I-Ig.{ :gI :9I-6nd sdBru 'uo^r€urPJ 000 0SA:I) slcrJlsr(I IBJruPlog oncsalJod uJaqlnos pu€ urlsnv uJaqJJou 'uolrnqqsv aql ur ellerlsnv uralsai^ lsaM-rJuoN (0I arn8rd) uor?nqrr?src.

'(HJESd) ZI}qullsetrN'U g'BrrPqlE{aa] I 01 p€or aq} uo uollcunf 'uortBlc 'uoIlB}S au{o?sBC Jo U uI{ 0ZI t{grnquol0 :(HJESd) t009 &.IISP.tr{U g BuEr,r\oI ol pBor aql uo uolJ?unf au,{of,sB0 Jo S uI{ S tZ :(HJUSd) 9661 urIsPJ,1l .U ,ppor g, uorlcunf ou-,{orsBCuo^.r€ur€C arll uo ^B^\q8lH I€ls€oC lsal& q}JoN 'B' 'ooprnqPrEd Jo g (rI{ S'Ii :(HJUSd) tggi urlssrll H' :(HJuSd) ILLZ ultsehl A '9.',{EiaqBrH IBts€oJ lsalA quoN Jo uoAreurpOJo S uII qeI :(HJUUd) 80Z9.ralrpr"J 'y 'ra^ru 'aeEepu€,{& rsraÎu J e.'tlrurJ^l :(HJEUd) ZgIg nupreyv' J B.{lIuIII puB 'q8ralurlral{ 'U'uo^reurBC -d ou,'to)ssCuaa,$teq :(HJUgd) 8V lla]retrto Jo (lu{ 992) 'uorlpls 'raÎlf rur 09I {oarC {rrBO :(HJUSd) Z99L z.reuaQ .17 uopu.{'I Jo N ru{ It ,d>I) 'euBrllE{oal 'dX) :(HJUSd ]LLZ zrewae H I Jo N (ur{ g'q8) Iu S SS:(HJEgd 'uBur.{\aN IIg zr?waI i? tunotrAl ol. rpur^'\rpuntrl urorJ pEor SuoIP uI{ 88 l(HJHSd) ruorlcunf 'dX) 'a{e'I ggEl preog s ? au.{orsBC:(HJUSd 0I9t pr"ag S T llES Jo ',{p,$qErH tsortA :(HJUgd) lg,AI sull1v l,I urarllroN JEarC uo err€qle{aal,\l Jo N uIrI '.ISI 'y'SNVC) 0gI lnoqv :(HJUS.I I Zf6I ludv qI luEld stuBs oql rqorJ sd{lllqd '.D '116I rI6il 'V .'{q p?]J?llor stlnrJ raCI AI palrallor suaulrads ra,{\ou {qqsy IN 'JJo uJnl uor'l€ls BSuupA uI{ g t 'uoAJPuJBCpuB uo}PIeJaC uJa,\\}aq.{€{qEIH Jo S 'paulutexa IPIsBQJ lsalA qlroN :VITVUJSOV NAUJSS,U su('IJxallor raqlo 'IrJp !unlrq uB olur popuEdxalou aql ()1asolc paploJ-acuo lnq lq8r€Jls {ll€ruJou 'a{rT-dPrls puB {crql ,{la^rl€lal alrrunjr :.{}un}Eur ol rorJd paas aql lo lsal uEr{] 'J Jnoloc JarlJEpB'apl^,r urul I-g'0 8uol urlu g I aloaJe:umlrq aql spJp,\\oluado'adpr{s 'u,uoJq '({Jrq} 'apr,r -,,n,, pole3uola ue utetSotnald :,{urqslou lttur Z-g I) passaJdruof, 'proôqo 'aurn8ol urur 9-gtEuol uIuI g'1,-g ,{lpBorq ot prosdlTla,{lpBorq aqt ur {lanbllqo 'rncco ol ,{[osJeASuEr]pauorlrsod spaali op suor]cuJsuoJ daop uropuBr lnq spaes aql uae^\]aq palJrrlsuoJ ,{llq5lls ^ra^ .{luo {llBurrou 'qsrarole{ 'pauo{rrql .{loJeq 'olEln)rlal 'u,ldoJq strr.SJBEr:spoasJaAo eJ€uoqun ^llqFrTS {luodo ^lasJa^Su€Jl lq8rT 'snorqEl8 'lBU 'snoarBlrErlJ 'apr,i\ '8uol '8uo1qo rrJ A-I ure 6-9 tr ,{l,!\orr€u saunEaT

(dl-r,LUpH I8 dllreu dtlr ur E/,r.,t' rrlcql^l U A t2 NuytsiaVol.4, No. 1(1982)

(which is restricted to the extreme north-east of South Australia and adjacent Northern Territory)- Acacia pickardii is distinguished from these two species,and indeed from all other members of the A. victo-nae group by its terete, pungent phyllodes.

The specific epithet alludes to the spiny phyllode apices which are characteristic for this taxon.

5. Acacia daweana Maslin, sp. nov. (Figure 5)

Acacia hammondii Maiden et A. malloclada Maiden et Blakelv affinis sed combinatione characterum secundarum distinsuenda: fruticibus ad 1m altis phyllodiis anguste ellipticis (3.5) 4 5.5 cm longis, 6 10 mm latis, nervis tenuibus numerosisque nervo medio (et plerumque nervo uterque latis) plus prominentibus, parce reticulatis; spicis densis;calyce breviter tomentoso. lypus: 6 km N of Marandoo (which is just south of Mount Bruce) on lhe road to Tom Price, 22'35'S, 118'05'E, Western Australia. 16 July 1980. B. R. Maslin 4682. "Shrub 1m tall, stem 2-branched at base; phyllodes dull green; flower-heads bright golden. Low rocky rise in spinifex at base of rocky range." (holo: PERTH, iso: CANB,K, NY).

Low, spreading, often more or less flat-topped, dense s-hrubsto 1m tall, dividing near ground level into a few to many spreading erect, slender branches. Bark on main stems grey but near their base peeling to reveal a reddish undersurface. BranchleLs terete, very finely nerved, very slightly resinous, apically yellow but becoming reddish-brown with age, shortly tomentose or shortly antrorsely strigose but becoming glabrous with age. Srrpules frequently persistent on branchlets, triangular, 1 2 mm long, 0.6-0.8mm wide at base, dark brown, 3-nerved (nerves sometimes obscure), adaxially appressed hairy or sometimes glabrous. Phyllodes narrowly elliptic with the upper margin slightly more convex than the lower margin, (3.5)4-5.5cm long, 6-10mm wide, length to width ratio 4.5-8, straight or more normally slightly curved towards the apex, rather spreading, glabrous or glabrescent (indumentum as on branchlets), medium olive green, not shiny; apical mucro thickened, acute but not pungent, normally slightly curved, brown; pulrinus 1 2 mm long, transversely wrinkled, yellowish; neryes fine with the central one (and normally one on either side of it) more evident than the rest, very sparsely anastomosing, nerves close together but the interstices broader than the width of the nerve, marginal nerve discrete and yellow; gland situated on upper margin of the phyllode at the distal end of the pulvinus (or 1-3 mm above it), obscure, 0.3 0.5 mm long, lip narrow and yellowish and not significantly thickened, normally with a discrete circular or oblong central orifice. Inflorescences sirnple and axillary but sometimes appearing falsely racemose due to the loss of the subtending phyllode, I per node, somewhat clustered towards the ends of the branchlets. Peduncles 5-10 mm lone. indtlmentum as on branchlets, base ebracteate at anthesis. Sprkes light golden, flowers densely arranged, 25-40 mm long and 5-6 mm wide when dry; bracteoles linear-spathulate, minute (< 1mm long), light or dark brown, sparsely hairy abaxially, lamina inflexed and acute. Flowers 5-merous. Calyx c. 1/3 the length of the corolla, cupular, gamosepalous,shortly divided into broadly triangular lobes, shortly tomentose. Petals c. L'7 mm long, connate for c. 7/2 their length, glabrous or sparsely shortly tomentose on lower half, obscurely 1 nerved. Ovary papillose. /-egumes (immature, few seen) to 40 mm long and 7 mm wide, prominently raised over seeds, 'r?oBo srq:lur papnpur sarrads aql Jo fuBur uo ElEp plau alqBnle^ Surprô.rd pue sueuraads iiurlcaloc ur dloq srq io uerlBrcorddB ur a,,!\ECsrJqC Jo Jnouoq ur pauEu sr sJl.rJds dqJ 'aAJau Il]JJqac pacunouoJd aJour € aABq lou op r{Jrr{^\ pue alBrJls {losJBoc aJoru a.lE qJrq,r sapoll[qd s]r ,{q (aôq€ palou {pEarlE sJalJprBqJ aql ()1 uorlrpp€ ur) eueo^\ep V uroJJpaqsrn8urlsrp aq UBJll (asroJlue puP.{{lrs ssal ro aJou srrBq) slalr{Juerq ,{rreq ,{lasuop etrnb seq epelrolew y qtnoqtlv IIB1 ur Z ol sqnrqs Surpeards oJB .{trtBcol srql ]E slupld aql ('ululo) srad) arqJouoretr{ uqof ol Surprorcv (JN) frolural 'ra^ru urorllroN qIPg Jo N (rn{ i) Iru 9 z urog palrallor LlSl elepuaddrr{J ! sr aur ol ur\oul uaurrJadsJaqlo fluo aql (,,BrlErlsny uJaglJoN,, palElouuB {ldrurs sr qJrq,,!\) ad,{l aql saprsaq pu€ sarrads uourruoc .{laa E lou sr {l}uarBddB epepoueur ETJ"JV' JsaJ oql u€rll luoprâ aJour dpqFrIS aq ol a^rau l€rluar eql ^\oqs sapoll{qd s}r sEq puB IIEI ru I ol. qnJqs B sr'saJrdBLaIr{Juerq aso:}uoulol-l.roqs .{lsnonJldsuoJ ssq puprl raqlo oqJ uo r?u"a/r"p "rJEry surEJpur slr lo r{)Ba pue apoll,{qd aql Jo aJluac aql uaad\}aq ,{E,rprul ssal ro arour palBnJrsarE asaq} puB }sal ar{l uBrll paJunouoJd aloru.{11q8qs so^rau o^\J a^pq sapoll{qd sll (99I :816I ,{alpad) IIE1ur } :}noq€aarl E ot sd\or8pup s?olqruprq luaf,sarqElS ro snorqelS seii Lrpuouweq BrrBry x,{lpJ {rreq aroru s1r pue nâ{rdsJosuap slr'sapoll,{qd a}Blnrrlar ,{18urreds'rapeoJq slr.{q sertads asaql qloq 'asolJ urorJ posrnFurlsrps eLrea^\epaiJaJy fFelnorlled lou sr d[{suor]pla.r aql lnq (JN),{la{Blg to uaprE]I epepolew-y puE (plb JN VM) uapr€tr1 rrpuor{r{rreg 'y 'x,{lef, ieau paJpld ]saq suaas snol€dasoue8 s1r pue saJuaf,sarol}uraleJrds'8uol 'sopoll.{qd 'Dr}drlle 'salndrls slr o}BrJlsrllnlll dl,rorrBu slr luelsrsJod s:U Io Junocr€ uo puE aqclag la uaprB] I ( qluog) a€JourTnfuorJcos qr pacpld sr evea!\\eperJeJV '( tutrloc srad'a,$p0 slJqC) las sEn^JrnrI alllll fra^ uorlelndod aJnrg lunow arll tE Suuo^\ou JrJrToJdolrdsaO Jaquroldas al€l ur palcalloJ s€.r puB spaasarnlBurrlrr passassod uaas uaurrcadsSurlrn{ auo aqJ {lnf prul ur paJarll€t arau pueq ol suaurrJads Suua^\ol] IIy paururralap uaaq lou s€r{ a8ue.r FurlrnrJ 'pouad -;urra,rou IInJ aql suorlJalloJ Jo flr)nBd aql ot an(I EuLfLnJ.lpue Euuenoyg '(.r\olaq aas) esn.l.la'Vro.l sV iellqeH 'drull tudsdJddql tBdpeLu 'a.roJaraL{l'pal.,.lllo.t aq ]ouu€J snlBls uorlE^rasuoJ Jraql Jo luarussassealEJnclE u€ 'Jeâd\oq 'suor:}nqu}srp ,{lJood sr EaJE aSuBU ,{alsraurpH aql Jo qcnu palJrJlsar aêq sarJads asaql qloq lrTql arB suorlEJrpur luaJJnC (^\ol?q aas) ssn/Ja I/ qlr.{,\ 'S,tgoZe) ,{ e)ulpd!u.{s srncco "uvar1"p y sarlrleJolasoql qloq lV (g,6A"gII uorlB}S sua\oG Bunf Jo r{trou ru{gI lnoqE lB lsBa-qlnos aqt ol ru{09 osl€ pu€ (g/g0ogll 'S,gt.ZZ) acnJg lunotrAlJo lsa./t\uI{ I lnoqP IIIq,\\ol P uIoII palJalloJ uaaq sBq}I ?raq^\ {rEd lEuorlpN a8upE ,{alsraur€H or{1ruorJ,{[uo u,1ôuy (II-09d :000092:I) lrrrlsr0 IeJruBlog onrsalroJ aql ul Erlprlsny uJalsal^ lsa^\-qFoN (II aJnErd) uot)nqulsre '(HJUSd) '8' 'uroouallrla 87r9Vullsehl U o1 p€or oql .J .arnrg uo su,/r\oceunf Jo N trl{ 9I :(HJUgd) OIZ aaee , tI I Jo M uD16 FurFl Ilrrl l€cruoc B Jo aprs ls€g :vI.IvEJSnv NUEJSSIA pautwexa suo4ralln ftqlo 'Iurrls 'qsrnaolla^ '{qsaU lEqMauros pelnloûoJ puE poualcrql lBq,\\aulos fJB-alJrunir :{urqs 'u,\\orq 'apri!\ '8uol 'prosdrTla 'ouln8a[ lou lq8rT ruru 6 ulru t + aq? ur paJBId [lanbrlqo (arnlBururr) spaas'spaas aq1 uaa,rlaq palJrrlsuoJ 'qsr,ro11ef sur5.rarlr:srreq,{\olla,{ puB 'prcsr^ alrq,/r\Jo aJnlxrur E q1r,$ asolrd .{ltroqs ,{lasJEds {l1r{3rTS'alBlncrlal FIasra^SuEJJ ,{IarnJsqoraqlpr'u,{oJq qsr[oJ8 BaJEIBU Surua^Ja]uraql puB u,\\orq IrBp oqurn oql prpry rurtsptr'{ a:uE€ ,{alsrarueH aqt ur U g NuytsiaVol.4, No. I (1982)

t- IF WWfu'l c* l*

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Figure 5. Acacia daweana.A Portion of branch. B Phyllode with €nlargementshowing apex, sparsely anastomosingnerves in middle of lamina and baseshowing stipules (s). C-Flower. A from B. ll. Masiin 4682(the type); B, C from B. R. Maslin 4628. 'urnlrq oql rBou alEJnpur puE ra{JEp sauroJaqLnq snouBJquaur puE uBarJ sr IrrE aql'li lo aprs auo u,r\op [EA\ JI€q lnoq€ Surpualxa ualJo puE paas aql Jo dol uo pauorlrsodIIl? paploJ e8lel e olur papuBdxa ';uol lruJoJrTrJ luul I c alrrunJr :apr^\ ruru t'0 c sr qJrq^\ anssrl r{sr,rollad Jo pu"q 'urBrp 'paas alruuap E dq papunorrns urur g'0 aql Jo alpprtu aql ur palpnlrs aloaJa 'relntrrJ '({crql 'apr,r lsnonurluoJ urpJtoJnald :uel {J€p luru I) passaJduroJ ruur g Z '8uol 'prosdrTlo'euln8al 'qsr,raolla,{'paas rulu t aql ur pauorlrsod dlanbrtqo spaas aql uea.,!1laqpaJ?rrlsuoc (^llq8rls ro) lou'paua{J[{l sur'J"ur :alBInJrlaJ^lleurpn1r.3uo1 'u,r\orq 'snoJqE[.8'snoursal 'spaas dlaJnJsqo + {JEp aql ralo pasre.r ,,{11q3r1s 'snoorErrof, 'opr,r\ '(qlSuol 'alEurJJrt ?nq 1BIJ {Plrrqt luur 8-, papuEdxa) Suol urr q 8-g 'snorqBlS sawnEa.I (saJrdBlBlad lE paua{Jrql ,{llqFrTSsa^rau) paAJau-I ^laJncsqo 'srsar{luB 1E SurpEardssle?ad'^\ollad IIIe?rdB aJ€ teql seqol otur qt8ual sll EIZ-ZII 'elloroJ 'luacserqBl5 'snouBrqurou papr^rp orll Jo ql8uol aql t/I lnoqE rar{ler x.qpJ 'sa{rds aql urqlrd\ pasradsralur sJa^\ol} snoJaur-} ^lal B .{lFuorsBJco lnq snoJaur g srarrlo/d-luocsarqEl8'o1€lnJrde'paxallur.{llqBrTS pue paualcrql dllqErTSaBurulel'Buol 'a|ssesqns '(lrnrJ ururg0 r sapafJqrq :pa:ueJJe {lasuap {lJelncrlJ€d lou sJa,roU ur uar{nAurur gZ ol dn Jo srsaqJuB}€ (uur gI 0I) salJunpad aql s€ Suol s€ aJr,\\} lnoqe 'uaplo8 'solf,unpad {llEuirou lq8uq sayrds ;o ;Ieq raddn uo pet€ntrs Jcprq snorrprs 'slsaqluB u,rorq passarddp (8uol urut I-9 0) IIBrus .{JaA p r{lr^\ ualJo lB alPal)BJqa 'snolnJaqnd '8uo[ 'soqruerq oseq urur 0T L sapunpad aql spua aq] spJEd\ol 'apou ',{rBIIrxB Jo palBJJuacuoJ .rad I puB aldulls saJuaJsaJoLIUI Barp IEJluaJ u,roJq {JEp 'luaururoJd '5uol ') 'pale8uolo e qI^\ sourlauros lou drl lllur t'0 [1lq31s .ro lelnurc 'puBI5 'snurÎnd aql lnoqE paluapur ,{I,,r\ollEqssourrlauros ur8Jpur aql aôqe urur g't -Z 'luauruord opolllqd eql Jo urSJBru Jaddn ar{1 uo pelEnlrs 1ou pua1.3 :qsr,L\olla^ '8uol ,,{lleurpnlrSuol Iuru g'0 c snur lnd :(qsorJ a^Josqo) dllpoul saurrleruossa^rau 'urFJBru Surpuarl pue Sursourr'4seu€sa^Jau IBrolBI lBrxpp€ paua{Jrql oq} o1 luaJBfpB Suruun.ra,rrau IBurpnlrEuol puotes E puB a^Jau IEJJuaJauo qI,,!\ :uoar8 a^rlo 'slalqDuBrq 'SurpuaJsB 'i ,epr,n uo s€ urnluaurnpqr + A orl€J qlpl.r o1 q13ua1 uru I '8uol '(orJnur 'lq8rBJls 'luo8undqns -8 uIIu gI-6 u,$oJq ,alnJp ,pouorlrsod,{ll€JalBI E ur Surpua pu" papunor.a.r) sarrd€ uroJrrlsor q1r^,\JrldrTla IIIEJulauur^s€ sopollfqd 'u,rorq 'snorrBJs'8uol '.reln8uerJl'a8E {rEp wur6-1 qlr,1^snonptJap lnq slalqtuPrq ,(palrn) ,lua.lEd,aur.l uo Jualsrsradsalndr?fi oEEqlr,r snorqplt Sunuo3aq Jo lqSrEJls + dreA srrEq) snolnraqnd .{la}nurur 'pa^rau ,{larnJsqo ,alora} s?alqrup.Jg. ..ra,{€l Japun p par np IEa aJ oJ puo qJEa uroq ^lasroJlal IJnJ ol pual qcrqn\ s5ur,\€qs.,l\oJJ€u uI SurlBrlolxo puB (sar{Ju€Jqra8unof) par r{Sr,{ar8Jo (s{unJl plo) âJE xa}rot.rolno ',.rqJlru ,paddot-l€U ruurtrl,, {re[J Iaâl punor.S?B SurqruErq surals ro pauop rarl]la "uerp ,prJSrA 'asuap ,.ro.I tll g-Z puB IJpl ru I ol sqnJqs SurpEards-opl,t\ ?Bq,,!\awos '('ISru'y'SNVC .uaploE :osr :HJ,llgd ;oloq) ,, o8uEr,{{Jor lo osEqlB xaJrurdsur osr.rd{Jor,no1 lq5uq 'uaaJ8 sa{Ids :ellq,ld {lEaur uorlBua^ a^rlo sapoll,{qd :,rqJlrE ruurtr{, {JEq :ttl g-A x 'Surppards ,asuap ',r,\o.L,.Iggi .096I ur I o1 qnrqs paddol IBU saurrJauos u.Vsel/4[U g '€rTprlsnv 'g,g0oglI 'S,St.AZ 'aEuBU .{lnf 9I ura?sa^\ dalsraur€H ,aJrrd ruoJ ot.pBor aql uo (ocnrg lunotr l Jo qlnos lsnf palpnlrs sr qcrq,r) oopuBrEtr^lJo N (uI g ;snd.t

'EpuangurlsrpsrlPI 'srFuol 'snqrluJoJulsor Iulu tr-t uru gI-6 alrdp pE srcrldrla acr.rleultu.{sesrrpo1l,{qd :ollB ur I pB errlrnJJ pas unrurJJE urnrarJads la llonl l I elolqdls{I V' alPlrurJJB xg ',rou (9 arn8rd) ds'ur1seryesnJJa BrcBaV 9

a8ue1],{alsJauBH aql ur errery'urtspll U g NuytsiaVol.4, No. I (1982)

4mm g.n.h.

Figure 6. Acacra e.ffusa.A Podion of branch. B Legume. C Node showing phyllode subtended by stipul€ (gland position arrowed)-D Flower. E and F Seed(E side view; F-plane view). A, C, D from B. R. MaslJrr4681 (the type);B, E, F from C G. Dawe 212.

Otber collections examined. WESTERN AUSTRALIA: Just west of the base of Mount Bruce, C. G. Dawe 008 (PERTH); East side of conicalhill, 3 km W of Mt. Bruce, 22'35'5, 1i8'05'E, C. G. Dawe 212 (BRI, CANB, K, PERTH); 15 km N of Juna Downs on the road to Wittenoom,B. R. Maslin 462? (PERTH); North east slopeof Mount Bruce,R. Pul.len10.940 (AD); Gully at baseof south side of Mount Bruce,Hamersley Range National Park,22'36'S, 118'07'E, M. E. Trudgen319 (BRI, PERTH); HamersleyRange National Park, westernfoot of Mount Bruce, 17 Aug. 1974,J. H. Wi?rss.n. (MEL 503364,PERTH).

Distribution. (Figure 11) North-westWestern Australia in the FortescueBotanical District (1:250000;F50-11). Known only from the HamersleyRange National Park where it has been collected from ar.oundthe base of Mount Bruce (22'36'5, 118'08'E) and also 30km to the south-eastat about 15km north of Juna Downs Station(22'53'S, 118'29'E). According to the collector'snotes on C. G. Dawe008 the speciesis commonbetween Mount Bruceand WittenoomGorge (22"14'S, 118'20'E). At the localities from where collectionshave been made,A. elfusa is locally abundant and occurs sympatrically with A. daweana(see above). Current indications are that both these specieshave restricted distributions. However, much of the Hamersley Range area is poorly collected,therefore an accurate assessmentof the conservation statusof thesespecies cannot be madeat the presenttime. (r)-6s991 '(ssaJd ul'raddoH pue uIlsBI I aas) esuas lBtlSololEulllr E uI eraq pasn sI iBcldorlqns * 'snourluadJas 'snornEl3qnsalBd .{l^\ollBt{sfJa^ ol pa^JnJ dllq8llS'pI3IJ }ou'BulpuaJSB 'alaJal ''urBrp '8uol 'uuoJITIJ urur I c urr SI-tI sapoJl,{qd snonplJ?p salndrls '8uno,{ 'snorq"l8 'paqqu-ulsal 'pa^Jau ^ra^ uaq^\ snoursal Eooqs ^taN snorqBl8 lou 'urdorq flaJncsqo ,{ra^ 'alaJal splqJaerg pal Jo u$oJq 1rl3rl aJB r{clq^\ slalqcuBJq 'slolqcuBrq uo ldarxa .{aJ8 pu" saqcuErq uo rlloours lnq s{unJl ul€t[ uo paJnsslJ 'l)adsB ,{IlEurpnlr8uol pue snorqlJ {r"g ^qsnq puE asuap .raqle.r flleutrou su,lrorJ SurpuaJSB up qlr,r\ sar{Juerq pup s{unJl ul€ur 'lalal punoJ8 urorl Sulstle s1unr1 'llel urEru g I o1 dn qll^\ ro paruurals-a18uls raqlla wr l-8 saa4 JIPrus Jo sqnJqs

'(,\N "IsW 'X 'SNYJ ;osl :HJugd :oloq) ,, (Barp olarclEc) ruEol u^\orq lqSll IroS'BrcBJy puB aallBur (ZIIZ Jgl\l) Psoalo 'Jc snldflecng qltar puelsse.r8 elpoul :8oL Uos alrnb sapoll^qd ,li\olla^ sra,'{old 'a8uero lslaIIJuBJq ',{ar8 s31a,r1 raddn uo rlsrppar {JBg surals lEra^es q1L/$ IIP1 ur 9 Z qsnq gItA ua?pnrl w[orletrN'086T dBI I I €ITErlsnVura]sal\'lood ulddoC V,, 'Ired sp.re,ro1 s8ugdg lllll l llll1lrg urorJ tu{ I8 IEuoIIEN aFu€E ,{alsraueH :snd,t 'lr4JrpoprqlE ollrrP-ulnJrunl'srlel 'sl8uol uIIug-t srulurn8al '(srarne slSEur) stpr^Eurad st1.ra;uocsnuM slclds rucSI 'srJ€JsoJoursar 'llanntr -iI srrpoll,{qd uou sllnruBJ enb B stuIJJB ,{ eurlsstnual eIJe)V

(Z arngrd) ,rou ds'ut1se14islllxa €rcecv l,

'saIJads srql Jo lrqpq qpdor8 Surpeards apl^1 JrlsrralcErBqc aql ol sraJal laqltda crlrceds aqtr '(sserd ur) raddoH pue urlsEtrl pue (816I) uIIsBl\l puE raddoH uI passncslp3rB'dnor8 'etceJv e1o1qd1st1 y aql Surpnlcur ar uorleltads Jo slf,adsv a8ueU falsraureH aq1 ur .rnrco arlrec,{gre4 'v pue e[oJ?uoaI y qlog spEaq-ra,$ou (alef,rds lou) fflnqolS 'Uoqs dllErurou sl.r puB (aplt\ urur gT-9 pue 3uo1 urur 0g-UI) sapoll.,{qd pBorq sll 'y {q pezruSoca.rdllpear sr lnq dnor8 elotqdlsq oqt urorJ paôurar dll.uelslp ool lou 'IN sr sarcads ruurl raqlouE sl epcquour 7IJBJy (sarJrdsraqlo l€ql !.rr{JlrU I' {DBIS f 'llErus aql ur ffau-rl puB Suol III€uuou) sapol[qd tlldrlla fllBJulauru,{sB sll {q puB '^\ol lrqBr{ asnJJrp s}r ^q dnol8 "roJqdrs,{J y arll Jo sraqurau raqlo ulorJ peqsrn5utlstp 'anssll .{Ipear sl esnLla e?ecv parnoloc alEd Jo puEq ,!\orJ€u e ,tq papunouns 'pauolllsod aJ" salooJe snonurluoc qlpd spaos paJnoloJ .{lanbrTqo 1€rl1 Il€urs IrEp 'sapoll,{qd pup'surSJBur paue{crql qll,\^ sarungal alEuIJrIt ualJo olElntllar lEU plcsl^ 'salndtls '(pua al?InuoJJnru pa^Jau-^r€J lualsrsJad qD€a ulorJ flasroJlar IJnJ qJIq^\ 'a s8ur^"qs ,$oJJEu ur SutlEIToJxapu€ I{srppal I) {rBq ..lqrllu IuuII L, aABq IIP salJads 'Y *IBcrdorlqns-auoz prrE asarlJ (lazlud g edreJtqrsrl puE alepurl ewt\DerE 'y 'ôlrEg 'a I l d IlrIrloqslqt y I) sarTIBsl.I pu€ IIanIAI l eloqdtsq y o1 palelar sr pue ar{rlag }a uaprBlÎ ('qluag) aEroUITnf uollJas ol siuolaq esnlla er)e)V

'raquraldas a}€l uI palsallor uaaq aA€r{ spaas aJnJeunul .{llqBrTSqlriu sarun8aT lsnFny o1 fetr l lnoqe uloq spuolxa Polrad 5uua,\rolli eql puBq lB suaurrtads aql ruoq Sul8pnf'Po]Jad EuqInr! pue Euuauolg 's^alp^ pBorq olul unr slaorc aql aJaq.trs1€U IBt.Lnl1e aq1 3uo1e rPJ pBaJds lr saop rou sadols ^{JoJ ar{l dn rpJ puolxa lou saop dl8ulluaas esnJJa "rJ?rY'slllq ar{] a^€al sasrnocralPd\ asaql araq^\ saull Suop relnellled ul puB sa8u€J ,^Aol {aarl '?"?rq?]? lo sadols aaJcs JaA\ol 3q1 uo ruPol pal dlJor uI ('ds Pipoiu ) xaJlulds 8uoluv

a:uEU,{alsretrleHaql ur Jery'urrsPl I U g NuytsiaVo1.4, No. 1 (1982)

glabrousexcept on upper side of the pulvinus at its extremebase which is densely minutely villous, uniformly longitudinally multistriate; narrowed at apr.cesintl normallydelicately hooked, light_brown, non-pungent points; pulvrrrus c. 3-mm long, orange, wrinkled; gland situated on upper surface of phyliode at distal end oi pulvinus,very indistinct,subrnerged, phyllode lamina,,o.-"lly very slightlyswollen about the gland,.Inflorescences normally twinned and with a ne* ihooit developing Irom within the axil, the subtending phyllode often deciduousby the fruiting stagei Spites light golden, 1-2 cm long, flowers somewhat distant; biactectlesmiriute ic. 0.3_mmlong), puberulous, claws oblong, laminae inflexed. peduncles 10_13 mm long, glabrous, base ebracteateat anthesis. Flowers S-merous.Ca_lyx cupular, t/+_7l} tie length of the corolla, membranous, sinuate-toothed with widely triangular norr_ thickened lobes,white-villous. Petals 1.5-2mm long, glabrous,very obiscurely1_ nerved.Zegumes (slightly immature) narrowly oblong,1o T cm long, 4_5mm wide, lirmly- chartaceous,gently curved, flat but slightly raised over the ieeds, glabrous, greyish brown; margns constricted between the seeds.Seeds (slightly iri-at,_r.*j longitudinally positionedin the legume, obloid ellipsoid, 4-4.bmt ltng, 2.bmm wide, compressed, dark brown but with a yellow "u"-shaped, central portion, ,r"ot .hir.,y; pl.eyrogyam open towards the hilum; areo.lec. 0.6 mm long and 0.,1mm wide; funicle much convoluted, flattened, membranous, whitish, imperceivably passinginto the all

Other collections examined. WESTERN AUSTRALIA: Tom price, 0.8 km from minesite entrance,K. Atkins 1221 (CANB, K, pERTH); Marra Mamba area, l-ol!919u" Sy"I,Iamersley Range,J. V. Blockley 291 (pERTH); Hamersley Ranges, 22%0'5, 117'43'E,M. ColeWA509T (PERTH); 2? km from Tom price orr the .laj to Paraburdoo,B. R. Maslin 4666 (BRI, CANB, K, MEL, Ny, PERTH, WAIT); 10 \+ I rt Tom Price-Dampierrail crossingon the Wittenoom_Nanutarraroad, _d. Maslin 4672(PERTH); HamersleyRange National park, \. .t.M, 4.8km from Milli MiUi S_prings-towards Coppin Pool, M. E. Trudgen 2401 G. NT, p, pERTHt; Hamersley Range National Park, 1.4 km soulh-west of hut at Cnppi" eoot, M E' Trudgen2548 (AD, BRI, NSW, PERTH). Distribution. (Figure 11) North-west western Australia in the Fortescue Botanical District. (1:250000 map F50-11). Known only from a restricted area of the HamersleyRange from HamersleyStation (22.17,S,i17.41,E) south-eastto Coppin Pool(22'53'S. 118'08'E).

Habitat. Low, undulating, rocky hills in ferruginous soil derived from the Marra Mamba and Brockman Iron Formations(de la Hunty, 1965).Some of the species recorded in association with A. exilis include A. aneura, A. maitjandii, A, spondylophylla, Burtonia polyzga, Calytrix longifolia and Goodenia scaevolina. ,spinifex, Tbe groundcover is dominatedby speciesof (?riodra sp.) Flowering and fruiting periods. Flowers from May to July; legumeswith near_mature seedshave been collected in late September

Acaciaexilis is placedin sectionJuliflorae (Benth.) Maiden et Betcheand is most ctosely a_l,liedto the widespread and zone-subtropicalx species A. tenui.ssimaF. (W.A., Muell. N.T., QId.).Although both speciesoccur in the HamersleyRange they have.not been observedto grow sympatri caliry-Acacia exrl.is is distinguished flom A tenurssr'maby its branchlet apices which are not resin-ribbed, its norma y longer phyllodes(14-.18 cm comparedwith6-1b cm), its longerspikes it0_20 mm "o-p"r-"d with 5-10 mm) which are a light golden colour (paler yeilow in A. tenurssr.maland + Subtropicalis usedhere in a ctimatologicat";;G;;NGJG;;J;G;; 't60qv.4[ aao, '_l,tl 'C 'U 'A 'f luorJ g :999t rrJsBJ,lJ g urorJ 3 :(adh elq t) tIrZ uoapn+ g W uJo4 A'.162 faqrolB uto4 V '(ernleurul ^IlqBIIs)ewn8e'l-g PaaS C qruerq Jo uor?rod C re,}lold-g'(d) snul^Fd Pue (3) PUBISSul,toqs eseqepolldqd V srlrxaerrBry l embtd

o t@ I Fq) ffiH h p t q? tr I p# d

^alsrauBH aql ur prrpty'ullsBl 'U'g 68 o8ueu 90 NuytsiaVol_ ,1. No. 1 (1982)

which possessflowers that are less densely arranged, its broader legumes (4-b mm compared with 2-3 mm) and its whitish funicle-aril (funicle-aril drying yellow in A. tenuissima). Although A. tenuissima can reach 4m tall (Pedley, 1978), in the Hamersley Range it is generally a shrub not exceeding about 2 m in height. Acacia exrlrs on the other hand is a taller shrub or small tree up to 4 m high. In the field, flowering plants of A. exrls may superficially resemble A. aneura F. Muell. ex Benth. (the two speciesoccur sympatrically in places), however, the new speciesis readily distinguished by its longer, lighter green phyllodes (grey-greenin A. aneura), by its lessdense flower-heads and by its long, narrow legumes.

The specific epithet refers to the slender, lax phyllodes.

8. Acacia hamersleyensis Maslin, sp. nov. (Figure 8)

Acacia tumida F. Muell. ex Benth., A. citrinoviridis Tindale et Maslin et A. xiphophylla E. Pritzel affinis sed combinatione characterum secundarum distinguenda: phyllodiis plerumque leviter falcatis 8 1,1cm longis, 1-1.5(2)cm latis, demum glabris, manifeste subtiliter nervatis; inflorescentiis axillaribus (non racemosis);spicis densis,3-6crn longis,6-8mm latis; leguminibus breviter pilosis, planis, non rugosis differt.

Typus; Rhodes Ridge, Hamersley Range, t 23'06'5, 119"2b'E, Western Australia. 10 "A Aug. 1973. M. E. Trudgen 391. bush 2.5 m tall. Bark on lower stems silver-grev and sornewhat fissured, higher up bark is grey-brown. Older phyllodes slightly glaucous, others medium green. Quite common on stony ground just above srnail breakaway on the south side of a ridge (forming a thicket at top of ridge).,, (holo: PERTH: iso: CANB. K).

Spreading, open shrubby trees 3.5-4 m tall, often rather slender and with a whisov aspect when young, single-stemmed or sparingly branched at ground level twith up to 6 spreading-erectstems when young). Bark grey, normally fibrous and fissured on main stems especially near their base, smooth on branches. New sfioofs verv oale citron-sericeouswhen very yottng, Brunchlets terete but angular towards the apices, finely ribbed, marked with distant scars of fallen phyllodes, glabrous o, so-"ii-"s glabrescent towards the apices, sometimes pruinose. Stlpules deciduous. piyi.lodes normally slightly falcate and tapered equally at both ends, slightly asymmetric with the upper margin more convex than the lower margin, 814cm long, 1-1.5(2)cm wide, lenglh to width ratio 6-12, coriaceous,normally speading,glabroui except when young, subglaucousto distinctly glaucescent,light green when young, the acuminate apices narrowing to blunt callose points which are slightly hooked; longitudinal rerves numerous, very fine and close together, neither anaslomosing nor basallv confluent, c. 3 nerves yellowish and slightly more evident than the rest, mareinal nerve- discrete and yellowish; pulvinus distinct, 5-10mm k,ng. ctnspicu,,usJy wrinkled, normally glabrous, light orange, depressedand stightly dltated atihe base; g.landsituated on the upper margin of the phyllode at the distai end of the pulvinus, lamina slightly swollen about the gland, submerged, comprising a circular o, oblong pore (0.3-0-6 mm long) and a very obscure rim. Inflorescencessimple and axillary bui often falsely racemoseat the ends of the branches due to phyllode reduction, 1--2per node, clustered at ends of branches. Ped,uncles4-10mm long, thick, longitudinaliy wrinkled, I sparsely puberulous, base ebracteate at anthesis. R"""ptu"L spu.r"ti resinous-papillose.Sprkes bright medium golden, 3-6 cm long, 6 g mm wide, howers srcuatrelsJaureqy sa8pu q3lq aql Suolv 'saEuBJaql urorJ Sulppal sasJnoJJale^\ 'salor8 FuolB{llErcadso surrog^luouuocun lou lI so8uErJo sadolsraddn aq1 pue se8plr aql uo s,i\org,{llprurou lr eJeq.$slros snour8n;rragol palJrrlsor sulaas ?E?.rgElI '(g,8I.lI r 'S,BZ 'S,AI.tZ) "ZZ) uBrrDlcorg}unotr tr Jo dtrurrrA aql 01 rluou puB (g,Ot.lII 'S,9I.82) ooprnqBred ol lsa^\ (g,Ft06II uBur.t\aNlunotrAl urorJ spualxa lr araq^r€arB '(9I 'AI 'II-0S.{ aBUEEfalsrourBH ar{l ruo.rJ^luo u^\oux sdEru000 0gA:I) lrlrlsltl 'uot)nqHlstI IBrruBlogancsalJod ar{l ur €rT€JlsnYuJalsal\ lsa,$-qlJoN(II Jrn8rd) '(99et09 'H 'f 'tL6I "1lp Tgt{ Jo dnp-HIUSd) u's s..JIIl.l cnv lI U 009 t 'c '{rpd 1Eacnrg lunotrl Jo salll^llraplas IEuorl€Na8u€E r{alsratllBH :(HJusd) sII 'y 'g,gt.6Tl 'S,6totz 'aFueu n eA| prluleqtqdo :(HJUSd) ,6 puB (HJUgd) 96 '(HJUgd '.USN '.ISI 'y 'uEru,raN'{c€qalEr{iA I'CgC) 96 rolleft tunotr{ :(HJUgd 'ANYC) 'g .J;!.',,qcln0 'eprs tgZZ uaEpnrl acpurrr0,,Jo lsBa ur 002 r{lnos uo (acnrg r{rBd 1I,\IJo qlnos s8pu lsrrJ) a8pru oopuBrBII IEuorlENaFupU âlsrauBH :(HJUSd '.IUW) 'g,60.BII 'S,Ot"ZZ 'o8uBU 'a8plu TSTI uaapnrf g ,{ + dalsraurEH '.{lln8 'r{ 'g,60.gII 'S,ot.eU oopuBrEtrl ,,osaupguBlL, :(HIUSd) glOI uaapnrL a 'o'uBU 'a8ptu 'r{ 'S,00"8II + âlsraru€H oopuErBI\l:(H&UUd) ZtjI qaapnrf A 'S,9Z"ZZ 'oFuBU + folsraru€Heq1 ur eSro8paueuun uv :(HJUSd) 169ua8prua'g ',tr 'o8usu 'agpru 'r{ 'S,f0.8II 'S,ttoAZ'a8uBU dals.raureg sapoqu: LZE uaEpnlL 0. 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'su,rlo0 'a;plE 609l- ulFeIN U €unf 01 peor aql uo ueur^laN Jo l N ru{ g 99 'HJusd) '8"uelu,r1aN'{r€qalEqla sapoqu :(JIyi 989, ull;ehl u lunot\l :(Hlusd lgNyJ) 'aJnrg 8IZ a.,!1eQ, , lunol I Jo lsaM ^lalerpaururrpue oopu€r€I l Jo N ru{ i 'J'peor 'dTJ 'HJUSd) aJrrd uroJ-oopuBrplÎururu uo SurssorcIaaJC :(JMA AIInI 'aFu€U,{alsraur€H'(arnrg a,.llpQJ 1W Jo S a8pu lsJrJ)a8pru oopu€retrl :(HJUSd) 'hl '9,w"Ifi 'S,0t.ZZ 'saEuBU 9009VlA aIoJ dalsraurEH:(dX) BtS tratttrotg 'as€rrrEls '.Isl It r's,Oz.8IT's,gloaz s.|â0 aôqY:(HJ,utrd l) otzI puE(HJugd '>I) 'acud qZZI suyllv'y uroJ'ssolaurENlW Jo sadolsraddn :(HJEgd) gtt sultpv '14-'alrsu^rol acrrd uroJ :vI.IYuJSnv NusJSsl pauluexa suollralloc ]aqlo 'paas ar{l Jo do1uo pauotltsodLIr" paploJ 'urroJrTIJ 'alnulul ^qsau ,,$olla.{alBd ,{ra^ B olur papu€dxa aIJrunJ:apl!\ uIuI g 0-2 0 aloar" :unlrq aql spJBdloluodo pu€ padBqs-,,n,,Jo snonulluoJ 'aJnJsqouraJtolnald :,{ulqs {lqEuosEa.r'oloale aql SurpunoJrnsanssrl painoloc elEd qllt\ u^,!oJqI{sL{arB '({clql 'apl'r '3uo1 'plosdlTla lq8ll IuIu Z J) passarduocfl}qFITS urrug 8-g A ururg'1-7 ol prolqo 'aurn8al aql ur poc€ld ,{lonbrlqo spaas spaos aql uaanaloqpalJlrlsuoJ 'ualo.rq 'pauo>lcIq} (.{l}q8rTssaullauros ro) lou 1q3r1 Jo r{sl,\\ollaf sarS,reuri3uo1 'un\orq '({1t.rn1eut urur/-t adrls J€ paôJr€u,{1ldnlqe urnlparu 1Brlslllq^\ lpseq'xadB 'alPlnJllar 'spaas lnq Sunod uaq,,$uaplo8 olBd srlBt{) asolld dl}roqs -{larnJsqodra^ 'snoursoJ 'apl|A ar{l raô pasrBr dllq8rls'alBlnpun.{llq8lls'snoacElJ€qJ{luulJ ulul S -g '8uol ruJ g otr 'Suolqo ,{li oJJBu'salcBldeceJ uo paJal.snlJualJo saurnta? peAJau-I 'luacsarqBIS 'qlSual 'c 'Euol ,{lornrsqo ro snorqBl8 rlar{l Z/I roJalBuuoc Luut9 Z slepd '(osuapualJo urnluaulnpur) snollr^-alrq^\ 'qsru,,r\oJqfleurrou puE pe^Jeu-g,{laJnJsqo aqnl xflec 'qsr,{olla{ pue paua{Jrql dllecrd€arB qclq^\ soqol 8uo1qoolut q13ua1 'Blloroc st: E/Z-EII roJ papr^rp Jo q$:]ol tlZ-Z/I x,{I?J snoraur-gsra,1lold paxouul 'snollr^ '8uol 'alBInI{lBds-r€aulT eururBl ^lesuop urul I c saloalJeJq:paBuEIJB dlasuap 'e8ueu i6 ,(elsretueHaqt ur prte V'urtspl^l U B Nu lia Vul.4,Nu. 1 (1982)

'2mm

IL t. .ffiI' \"%

Figure 8. Acacrahamersieyensri. A-Portion of branch. B Flower. C Seed.D Legume (side and plane views).A from M. E ?rudgen 391 (the type); B frcm B. R. Maslin 4630;C, D from C. G. Dawe 218. 'earEaBUEU {a1s.raure11 aql ruo.r; IIuo u^\ou{ sr salJads aql }Eql tr€J aql ol sraJar laqllda clJlrads aqJ

'luaJsaJnBIS dllcurlsrp Jaôu puB snoatB,Irocerotu oq ol puol sapo11,{qdarnleur aql 'aulrl alqEJoplsuo, B roJ slslsJad pu€ JoAIISsuJnl unluaulnpul snoatlJas arll pu€q 'lua)sacnpl8 raqlo aql vo elL{qdoqdx V'uI dllcllrlsrp 01 sno)nEl8qns auocaq urnl ur qcrq^,\ sapo11.{qd8uno,{ uaar8 lqBrrq snorqelS ol ssBd dl{rlnb asaq} srssa,{alsYaule{ 'y 'palBrlrur ur lsJrJ uaq,!\ slooqs ,\\au snoatlros ,t!o a[ aêq sal)ads qloq 'pauallBU 'aplt\ 'Fuol q8noqllv ,(llrullslp oIP puP r€lnotqJo lsourlp uIuI 0I-l ruul II 'apl,t 6 aJ€ spaas pa8uErrE .'{llBurpn}rEuolslr urJg'I-I aJE puE Suol urcle rlJeaJ uBc puE snorqslE eJB sarun8ol ar11elfqdoqdtx'y uI 'spaasonbllqo 'passaJduro)ssal 'Joll€urs s1r puE seulnFal asolrd'JallEurs slt 'snolnJaqnd uaploB Jaâu sI qJlq,{\ x,{[BJ 'roSrBI 'salrds papr^rp ^ldaap arour s1r asuap sll'(FEur OIX lE a^rasqo) sapollf,qd pa^Jau Îsnor^qo arolu sl.r ^q v tqdoqdlx y uloJJ paqsln8ullslp sI srsua,{alsJaurBv y d1ecr8o1oqdro141dr€nrqad puB raqualdaS uaa,tlaq IIurcuI srar'!\oupu€ s1€U IBI^nllE 'azrs 8ur.{l ,!\ol uo sJnJJo eylqdoqdtx enecv pu€ adBqs rellurls B Jo sopoll^qd qll^\ puE saJuocsalou:u1alecrds'3uo1 qll^\ (salcadsJaurroJ al{l ut palrBuE oJour q8noqll€) 'V' saorl Surp€oJds IIBurs ar€ srcuatralsJaureq puB (,,poo,!\o{Eus,,) erutqdoqdLx eowv '(lualBd 'souln8alSuno^ snooJrrasdra^lrs ro uorlrJ lou) passaJdde[lasuap pue sapBldaJar pu€ selcunpad snolnJaqnd uaplo8 ^lasuap sBq 1I'(srsua,{alsvaar"tl 'y ur lsol ^l{crnb .{raA unluaurnpul) o3€ 3l€rpauualul lllun urnluaunput 'ra,$oJJeu ,{JaAIrs ro uoJl.rt 'passarddE uE aAeq qtlq,!\ sopoll,{qd alEJIEl arour dllBrouas sErl srp.rJ..ôurJ?rJ"rtEtv' ^lnf puE IudV uao^lloq ,{lutPul sra,rrrogflSururaas 'rall"l 'y qDIt{,!\ (lu g ol) aarl InJaceJg oJoru s sI lI srsua,{alsrau{Bg rlllra parBdruoC 'sasJnocrolB^\ ol palJrJlsal .{llEulJou oslP sI '(,,ulet JaAtJ,,) slplJ] ot[JlLJ eoeov

'satunFJl JqoLrd ^llJoqs 'poplurra-uou '(asoruacBJ 'sapollf,r{d slr dq pup sa{rds rapBoJq lou) .{J€llrx€ slr alBDIpJ ^lsnoncrdsuoc ssol 'Jo,!\oJreu ualJo s1r ^q epLwwy uroJ1 paqslnEullslp sI s6uatralsJoareq vDery spuo qloq 1E ^IIEnba paradEl aJB qcrq,!\ sapo {r{d alerJlsrllnur 'luoJSacnBIS serurloruos 'sno.rqe13 'a8Jel qlld\ puB 'slalqcuEJq arnleur asournrd 'snorqPl€ 'saauacsalouur saurrlauros qlu^ al€crds Suol qtl^,\ saall IIBurs ro sqnJqs 'sasJnoJJale,/r\ aJ€ s/sua,{alsJaureq V puB salcads slt{l qlog ol paulJuoc ^lulelu IlBl 'puE sr srsua,fals.raur"q 'y a{rTun uotltuocun .{laAIlBIaJ sI 1I EoJB oSuEE .{olsJaureH aql ur lnq J N pue y &\ I"Jrdorlqns/leJldoJl ur uoulruoJ sr epILunJ eIJeJV

'Iazlrrd 'V" g erutqdoqdlx V pu€ urTsetrl la alBpurl srprrr ouu?IJ qlueg xa IIantrAI ';4eptwnl'v. sorrads palnqrJlsrp dlapl$ aJour aql qll^\ saIlIulJJBaAEq cl1 sJeadde puB arlJlag le uaprE] I ('rllusg) aBroullnr uollcas I]1 sSuolaq s.rsua,fals.raweqeDeJv

'raquraAoN IlJea (]1 Jaquraldas alBI urorJ palcafloc uaaq a^eq spaas alnlBur qlltt saurn8a] lsn8nv alel 'potnd lnoqE Irlun spuslxa puB,{lnr ur sacuaurruoJ Suua^\old Eu\tnry pue Suuaaoli

'BrolqdouoJqJlp'g puv BrolqdoJnal 'E 'el'{qdowq8 sryd/]leJng S a salJads snldtlezng p.cB Irpuepreu] 'V 'edJeJouInrd 'V 'ernouy y 3'a sar3ads elxa)V J€qlo qll\\ uollpltossB uI "u"asr,,'1 .J puP 'J, 'sua8und 'Ltzurrlzs ercEruel etpouL auqce4ce[d 3 o frlunoc ,,xaJlulds,, ur s,,rro.r8 'a8roC sarcads aqI {Is aql tsurBBE pallanoqls puBq ddsrqa B urroJ ol tuoouallllA 'uBru,r\aN lunow S a saculd ur uaas aq uBc pu€ lrqpr{ dlpurds € soulnssB ^lluanbaJJ 'a;u€U 'uqsel €6 ,{elsreurEHerll ur erc?ry I u g 94 Nultsia Vnl.4,No. 1 (1982)

9. Acacia marramarnba Maslin, sp. nov. (Figure 9)

Acacia inaequilatera Domin affinis a qua cortice fibrosa; remulis non-pruinosis, phyllodiis 2 ,1cm longis, 1-2 cm latis, vix undulatis, pallide viridis vel subglaucis, obscure reticulatis, axibus racemorum pedunculisque rubiginosis, legumis leviter arcuatis, funiculo brevissimo differt.

?ypus: Marandoo Ridge (first ridge south of Mount Bruce), Harnersley Range, t 22'40'3, 118'09'E, Western Australia. 24 Jttne, 1975. Malcolm ?rudgen 1338. ,,A dense bush 2-21: metres high, fairly spreading. Bark grey-brown, smooth, dull. Branchlets red, flower stalks red, leaves dull whitish green. Growing with ?riodra and other Acacra speciesin stony red loam." (holo: PERTH; iso: CANB, K).

Spreading shrubby trees2-3 (5) m tall, becoming t gnarled and straggly with age, trunks sparingly divided near ground level. Bark not corky, fibrous and slightly fissured at base of main trunks otherwise smooth, grey on trunks and branches but red-brown, light brown or orange on branchlets. Brcnchlets terete, finely ribbed, marked with raised scars of fallen phyllodes, glabrous, not pruinose. Strpu.les indurate, spinescent, 2 4(7) mm long, patent, straight, brown, often absent from some nodes especially with age. Phyllodes asymmetrically elliptic, very unequal sided with the upper margin longer and much more convex than the lower, 2 4 cm long, 1 2 cm wide, length to width ratio 1.2-3, coriaceous,barely undulate, glabrous, pale green to subglaucous,not conspicuously pruinose, :l abruptly acuminate and ending in a straight sharp brown point 1-3 mm long, narrowed towards the base, margins yellowish; principal nerve situated near lower margin, raised, yellowish, lateral nerves obscure and tardily anastomosing;pulrrnus c. 1mm long, wrinkled, slightly dilated at base, brownish; gland situated on upper margin of phyllode at distal end of the pulvinus or up to 1mm above it, oblong, c. 0.8 mm long, lip yellow and very slightly raised, lacking a distinct central orifice. Racemes acropetalous.1-2 per node, greatly exceeding the phyllodes, 6-15 cm long, concentrated towards the ends of the branches, the axes together with the peduncles glabrous and red-brown. Peduncles twinned along raceme axes. 1-2 cm long, base ebracteate and very slightly dilated. I'1ower-.heads globular, light to medium golden, the red-brown peials 6ften visible through the stamens in the centre of the heads at anthesis, 25-35-flowered: bracteoles linear spathulate, c. 1mm long, glabrescent. Flowers S-merous. Sepals united at their extreme base, linear-spathulate, c. l/2 length of petals, glabrous, apically light brown otherwise normally colourless.Petals 1.b-2.5mm lons. connate for, c. 2/il their length, the free portion red-brown, glabrous. Legumes-narrowly oblon_g,to 7.5 cm long, 7-8 mm wide, * chartaceous, slightly iurved, slightly undulate, raised over seeds (umbo wrinkled), glabrous, slighlly shiny, light biown (yellow brown prior to maturity); margins narrow, barely constricted between the se_eds.Seeds mostly oblique in the legume although some may be longitudinal, widely ellipsoid to almost orbicular, 4.5-5 mm long, c. 4 mm wide, slighlly .o-p.".."d, black, dull; pleurogram very obscure, represented by a shallol,ly curved line c. 0.5 mm long which is situated near the top of the seed; lirnrcle thickly filiform, expanded into a clavate anl which extends down about 1/4 the length of the seed.

Other collections examined. WESTERN AUSTRALIA: Northern end of the Mount Beasley group of mountains, S of Mt. Beasley itself, between Meekatharra and Mount Labouchere on the road to Mount Augustus, 2 May 1981, M. I. Btackwell I0 (PERTH); Hamersley Range, (Mt.) Brockman, 22.32'5, 117.14'E, M. Cole WA5068 (PERTH);Rhodes Ridge, 53.5 km NW of Newman on the road to Juna Downs. B. E. Maslin 4610 (K, MEL, NY, PERTH, WAIT); Paraburdoo, B. R. Masln 4660 nrorJd-C :(ad.{r aql) s Etr ualpn; ? ,.1truro4 J :rsgzua Epnq g tN utorA:gLg" ""tpfll:;1iry"{tJv 'aunge.I (.\er^ eprs g :Mer^suEld () peas g pue ( re^\old 'rlru€rq d 3 Jo uourod-g aseqapolltrqd puB salndrtsIurds 8ur^\oqsapoN y equeuerreur ErrErF 6 arn8rJ

'waatSbt ,M c --- .-. Lu. e s /sJ ffi-1

- (uur a

aBuB! delsraureHaqt ur prrery'urlseI\I U g 96 NuytsiaVot. 4, No. I (1982) (PERTH); 16 km S of Kumarina Hotel on Great Northern Highway, B. R. Maslin 4965 (PERTH); 50 km N of Mt Vernon Hst. and S of Mininer Stn.; A. A. Mitchell 761186(PERTH); 50 km N of Tangadie Station, A. A. Mitchelt 269 (CANB. PERTH); 20 km E of Prairie Downs Hst., Upper Ashburton, A. A. Mitchell 341 (|E-RI{)-. Yq*ldoo Ridge,Hamersley Range , t 22" 40,5,118.09,E,M. E. Trudgen 1049(PERTH); HamersleyRange National Park, 4.6 mi (7.5km) south on tracli to Turee Creek from the Juna Downs to Coppin Pool track, M. E. Trudgen 2b?6 (NSW, PERTH); HamersleyRange National Park, small unnamedhill on the north side oi Mount Trevarton , M. E. Trudgen 2631 (BM, BRI, PERTH).

Distributictn. (Figure 11) North-west Western Australia in the Fortescue and northern Ashburton BotanicalDistricts (1:250000maps FbO11, 15, 16; GbO4,8). Most collections are from the Hamersley Range area from about Mount Brockman (22"28'5,117o18'E) east to RhodesRidge (23.05'S,119.22,E), however, the species does extend south into the Ashburton River basin. Although not forming locally- densepopulations, the speciesis not uncommonthroughout its areaof distribution.

Habitat. In the HamersleyRange area the speciesgrows on low hills the rocks of which-normally belongto the Marra Mamba Iron Formation (Macleod,1966). Soils are skeletal and comprise highly ferruginous red-brown loam. The associated vegetation is Acacia shrubland e-g. A. aneura, A. pruinocarpa, A. maitlandii, A. stowardii, with a ?riodla wiseanaground cover.

Flowering and fruiting perrod. Flowers from May to July. Legumes with immature seedshave been collected in late August.It is probablethat theseseeds would have reachedmaturity by about late September.

Acacia_manamambabelongs to Sect. Phyllodineae DC. and is mostly closely allied to the Western Australia-Northern Territory arid zone speciesA. inaeqiilatera Domin. Both thesetaxa are small trees with spiny stipules,distinctly asymmetric, pungentphyllodes and long terminal racemeswith brightly colouredaxes.'Although -A. the two speciessometimes occur sympatricatly in the Hamersley Range, manamambais readily distinguishedby its smoothor fibrous (not coiky) ba"rk,its nor pruinose branchlets, its normally smaller, Iess undulate, pale green to subglaucousphyllodes with their very obscure lateral venation (piyllodies in A. inaequilateraare 2.5-7 cm long, 1.5-3.5cm wide, often grey-green,and alwayswith a prominent reticulum), its red-brownraceme axes and peduncles(not puiple), its slightly curved legumes (not circinate) and its funicle *hich does not enciicle the seeds.

The specilicepithet refers to the Marra Mamba lron Formation on which the speciesis frequentlyfound.

10.. Ac-acia pyrifolia DC., Prodr. 2: 482 (1928). ?ype; Nouvelle Hollande, cote Orient,Mus. de Paris 1821(holo: G DC: iso:K. P).

Acaciaclementii Domin, Biblioth. Bot. 89:812 (1926),non Maiden et Blakelv (192g) syn. nov. Lectotype(here selected): N.W. Australia.Between the Ashburtonand De Grey rivers.E. Clenent s.n. (PR52?7b8-right hand specimenon the sheet;iso: K, PR left. hand specimen on sheet no. SZZZ58).Lecbparatypes: 'Clemint (1) Western Australia: Between the Ashburton and yule Rivers. E (K_vegetative specimenson the sheet are A. pyrifolia, seedsare those of A. inaequilatera);\2) Bay oI Rest,Dampier's Archipelago, Feb. 1818,A. Cunnrngiams.n. (K). 'u.\loulun saurn8aT EIloJoJ aql Jo qlSual aql 'snolEdasourB3 ',!\ollaf, JIer{ uBql arour X,{IBC pa;UB B flasuap sra,rolt aled sa1rds 'so^Jou JaqJo aql uEql Juapr^a aJou {l}q8rls o^Jau IBJluoJ arl} qlr^,\ alErJlsrllnur 'snoJnEl8qns 'apr,r '5uol 'JBaurl ,{laurJ al€d trlur t Z urc 0Z II ^I,!\oJJEusapoll,{qd :aJB sarDadssrql l)1 paJJoJaJsuaurrJads a8ueu {alsJaurBH aql Jo sarnlBal luarlBs arlJ

,{lorl€lg la uapr€tr1 suo8rnspe'V €II

'dno.I3slql ult{]lu salllxalduoJ clurouox€J oql eAIosaJ ,{llnJ ol rapJo uI paJlnbal are salpnls ,{Jo}BJoqBI puP plalJ aÎsualul aJour ,,{lpalqnopull '€ars a8upu .{alsralu€H ar{l urorJ sorJads toJ uouBrlJlluapl ol ple ue se puE uorlBrrB^ aql 01 aprn8 e sE paprôrd aJE salou 8ur,$olloJ aq1 suoseal asaql rod slaaqs urnrJBqrarl uo pa^\atA uaq,!\ ,{llBrcadsa 'JaqlouE auo alqurasal dlIEIOIJ.Iadns '^ll€uorlrppv sar)ads aql Jo ûBur saJnlBal lEclSoloqdJou asaql lo ulBlrac ur slsrxa uorlErJEAolqeraprsuoJ 1Eql salpnls unIrBqJoq pue plall qloq uIoJJ}uaplâ 'Jaô,,!\oH sr lr strlsualJBrEqJ p€aq-ra,\\ou pu€ {rBq llaq]. puB uoll€JualJo paas .rraql 'suorsuarurpaun3al puE apoll^qd JIaql ^q {llBdltulrd paqsrn8urlstpa.IB solJads oqJ'(186I) dalpad puE (086I) urtsel! ur osl€ pup aôqB "uErsuiT?e'V'rapun palou uaaq oABq dnor8 r.rpre,,r,to1sy aql Jo sJaquolu UIE1.raJSulqsln8ullsrp uI paJelunoJua 'Suolqo 'lEU sorllnf,rJJr(I sarunlie1 snoulSelrlrBJ + o1 snoaJ€lrBqc 01 reaurl rlor{} puB sax,{lBc palJassrp dpJoqs 'snolBdasoure8 ;rraql 'spBaq-Ja,\\ou (JElnqol8 {laleJ) I€rrJpurl^J ro prolqo rraql'Jar{]o8o} asolJ IraA sa^Jau qJI^\ sapoll,{qd alelJ:}slllnru 'SulsourolsBuE-uou ,{1aur; Jraql ^q pazrJolteJ€qt arB dnoJ; slq} Jo sJaquaur aqJ 'earB 'y a3upu ,{alsraluEHeql Jol paprotar arp fiuoslau'y puB Blntsn_unp lnq IIe puE '(ssard ',{alpad ur puB uIISeI,{) ExBl auoz pIJE 3ur?uer-apr,trarB sallods asaql Jo JIEr{ uBql arotr l (.{alpad Blotr rl|y ufs) uoplBl,{ LIpreA\oJSy pu€ UIISEtr^leolqdopoqr 'y '.urISEtr 'V '(arootr\l 'l '3z1ld l rLuospu S erLrlqls V ufs) Ilonl,{ eueadual y 'n rurTSEW 'y 'dla{Blg 'z\^ IA plnrsnrrnp y eue8ulrl1e to uaplBtr^l suaErnspe V dnoJS-saI'adsxalduroc flertruouoxpl srql ol PaqIrcsE{l}uaJJnJ aJEselDads uaâS dnor8 rrpr?,llols BIJBaV 'II

'EJaje[tnboeLqy 'sarJads,$auu,{\o slq 'Y JO Spaas pu€ e!pJ!,r'ld Io sra,\\ou puE aSBIIoJpoqlJJsap urulofl lr?uat{Ia/, Y aureu 'JJo-ua{orq aqt rapun lBrll sr pauaddBq sErl t€q.^ {lluarPddv uaaq sBq alJlunJ aq} 'uaql lsour uo qgnoqll€ 8u11cJtrua^lalaldtuoc salJrunJ a^€r{ r{)lq^\ }a{tBd pauolluaut 'elloJytd a^oqp aql ur spaas auros aJe aJaqJ y ur JalJoqs qcnu sI lnq e.Ialellnbaeut 'y ul ploJ alqnop E ur psas aql salDrrcua{lalalduroJ qslqr\ alclunJ aql Jo ql6ual 'v arl? uo {lrrBuflJd pasEq orB eralelrnbaeut pue erloJr d Y uaa,|Alaq ssJuaraJJlp 'uruoq pass aqJ era?Blrnbaeu|Y o1 alqBJaJar arB qJlqd\ Jo auros sapoll,{qd 'y asool arE 'sa^lastllaql spaasaql saptseq'1er1cedpaas aql uI 'e!IoJ! d JoJ uollP-u€^ Jo oSuEJIBJnlEu eql ulqllid sllBJ^lrp€al laaqs slql uo IEIJaJBUTaAl]€JaFJ^ aqJ sJa^lJ alnl pua uoFnqqsy aql uoa,!\laq uro4 palcalloc suolurf,ads (elrrals) luauralc oql qll,\\ la{cBd B ur papnlrur arp pue ^\a){ Iv ale lLluaualJ y rapun poqIJJSapululo(I qJlq,t\ spaos aqJ ralceJeqc Jrurouox€l Jr1sou8plp .ro luecr;ru8rs e lou sr slql (sraqlo UEI{I snoacprrocarour oq u€c auos a l) {IIIIqEIJ€A I€rnlxal apoll^qd etuos llqlqxa saop 'v. (ururoO ruosi our apnlrur plno$ I araq) asuas peorq a\1\ q elloJlrtrd y qSnoqllV 'uraql uaa,/$laqsaJueraJJrp lPrluElsqns ou lJalap uB) puP rr?uaL{JLl v'lo sJd,{}u^slle oslB puE "rlojr/J,,fd-yJo ad^loloq aql paulruBxa aAPrlI spsas eql alJ.lrJuJ^LalJldtuuJ qJrq,r solJrunJ slr -'tq puE sopoll.{qd snoa)BtJo, s}L{q s/Jolr,{d y tuoq parlsln8ullslp sP,\\ sarJads aqJ suollcalloJ aanll uo pasPq se]li\ ululocl rr?uaualJ y aurBu aqJ

a8ueu,{elsreltleHeqt ur BrrBr}''urtsel{ U g 98 NuytsiaVol. 4, No. i (1982)

_ The characters given above accord with those of A. aclsurgens(see Maslin, 19g1), however, legumesfrom the Hamersley Range populations are required to confirm the identification.

This species is superficially very similar to the long, linear phyllode forms of A. stowardii (see below) but can be distinguished by its generally longer and slightly broader phyllodes with a slightly pronounced central nerve, its denser, paler ylltow spikes and its slightly longer caryx.

11b. A. atkinsiana Maslin

See speciesno. 2 above.

1lc. A. kempeana F. Muell. (syn. A. sibirica)

This rvidespread . arid zone species is not common in the Hamersley Range area having been recorded only from Marandoo which is situated near the base of"Mount Bruce. Acacia kempeana is most closely allied to A. stowardii from which it is distinguished mainly by its broader phyllodes and legumes and its transverse to obliquely transverse seeds (see key to species above). pedley (19?g) reports that where the ranges of the two species adjoin in north-western eueensland intermediates sometimes occur. Judging from some specimens seen, it seems probable that intermediates between A. kempeana and,A. itowardii also occur within the Hamersley Range area. More intensive field studies (in particular to gather fruiting material) are required in order to gain a clearer understandlng ;f the relationship between these two speciesin the Hamersleys.

11d. A. rhodophloia Maslin

previously (Maslin, _,As noted 1980) A. rhodophloia is a variable species. The Hamersley Range populations of this species possessphyllodes that are narrowly elliptic.(often linear elsewhere) and which "ange from 4.6 6.5cm long and b-gmm wide_with a length to width ratio of6-13. Sometimes A. slowardiJ (sieebelow) has phyllodes similar to this but A. rhodophloia is readily distinguished by its ,,Minni Ritchi" bark (i.e. red and exfoliating in narrow shavings whici curl retiorsely from each end) and also by its flower-heads which are br&der and with floweri more densely arra nged. _Additionally, the calyx on A. rhoclophloia slightly exceedshalf the length of the corolla whereas on A. stowardii it is as long u", or-.hoit"" than, half the corolla-. Flowering specimens of A. rhodophloia from the Murchison River area may superficially resembleA. atkinsiana (seespecies no. 2 above).

11e. A. stowardii Maiden (syn. A. clivrcola pedley)

This species.is .. recognized by its grey bark, its finely multistriate phyllodes, its golden light spikes with flowers not densely arranged, iis gamosepalouscaly, which does not.exceed c. U2 the length ofthe corolla, i"ts "h"rt"a""o.,s 1"g.,-". *hi"h.r" lrom 4 9(10) mm wide and its longitudinal to longitudinally oblique seeds.As noted by Pedley (1978) under A. clivicola (= A. stowaidii), the normai range in phyllode size for this species is from (2)3-6(7)cm long and (0.7)1_3(4)mm wide. Acacra stowardii in the Hamersley Range area is very variable witir respect to its phyllocles which .are often longer and/or broader than normal. In some populaiions the phyllodes are of a constant size but in others considerable variation has been observed. For example, on the collection B. R. L[aslin 4605 and 460bA (which is a

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13. Acacia xiphophylla U.Pritzel, Bot. Jb. 35: 305 (1904). Lectotype (here selected):Tree, 4 m in height, 25 km south of Roebourne,24 April 1901,L. Diels 2808(PERTH a singlephyllode only, ex MuseoBotanico Berolinensis). 'clementi'), Acaciaclementii Maiden et Blakely (as J. Roy. Soc.W. Aust. 13:26 t.19 ff. 1-4 (1928),nom. illeg.,non Domin (1926),syn. nov. Type:Between the Ashburton and Yule Rivers W.A., 1898, E. Clement (holo: NSW; iso: PERTH-fraqment ex NSW).

0 50 10Okm

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* A. cuspiditoria

POFT HEDLAN

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Environmental Services. Garry Marney is thanked for field assistance and Suzanne Curry for very competent technical help. The project was conducted at the Western Australian Herbarium (PERTH) with financial assistance provided under an Australian Biological Resources Study grant from the Bureau of Flora and Fauna.

Index to specimens studied for new species described herein This index is arranged alphabetically according to the name of the collector. Numbers in parentheses refer to the corresponding numbered species in the text. Unless otherwise indicated, the specimens cited are housed at the Western Australian Herbarium (PERTH). Abbreviations for herbaria are those siven in Index Herbariorum, Part 1, Edition 6 (19?4). In the case of Kings Park and Botanic Garden, Murdoch University and the Western Australian Institute of Technology (all in Perth), there are no formal abbreviations so KP. MURD and WAIT respectively are used inforrnally here.

Ashby,A. M. 4491(4 CANB,K, MEL, PERTH). Atkins,K 445(8), 1209(2-CANB, PERTH), 1213(2 K, PERTH), 1221(? CANB, K), 1225(8 K, PERTH),1240 (8-MEL, PERTH),1257 (4 Type:CANB, K, NY,PERTH), 1264 (4). Beard,t S.3510 (4-KP, PERTH),4366(4),4569 (2-KP),4603 (l-KP), 6148(2) B?ac-k,.R.L s.n. 10Oct. 19?5 (1), s.n. 30 Jan, 19?5(1 NSW,PERTH), s.n. 6 Dec.19?4(1). BlackwellM. 1.10(9\. Blockley,J. Y.291 (?), 3?8 (8 KP). Btooker,M. I. H.2089a(1 CANB,K, MEL, PERTH). Coie,M.5006 (8),5030 (2),506? (2) 5068 (9),5097 (7). Dawe,,Q. G. q0r8lqL?10(5) M112 (8-PERTH, TLF, WAIT), M154(2-CBc, PERTH),212 (6 BRI, CANB,K, PERTH),218 (8-CANB, PERTH). De4 B. s.n.? June1979 (2 MURD). Dernilzr_g-.61-_1_(1. 4P, BERIIH), 2?78(4 KP, PERTH), 4416 (l-KP, PERTH), D5?sb(2-Kp, PERTH),D7084 (2-TLF, PERTH),7652 (4) Gardner,C. A. 6152(4),6208 (4). George,A. S. 1068(1). Hastinss,B. S4356(2*KP). Lethbridge,I. L. s.n.July/Aug. 1971 (1 BRI, PERTH). Maloney,B. 73/53(2). Maslin,B. R.2733(2-AD, MEL,NSW, PERTH),2761 (2 B, BM, c, NSW,p, PERTH),2771 (4) 4586 (8 PERTH,WAIT),4609 (8),4610 (9 J{, MEL,Ny, PERTH,WAIT),4621 (8),4627 (6),462S (5), _4630-18),16,1{ (1 Type:q4l{P, K, MEL, PERTH),4646 (8), 4648 (2), 465i (4),4660 (9), 4666 (7.^..B_!]rCArjp,.4, UEL, N,y,PERTH, wAr"r), 4672\'/), 46'7'7(8), 4681 (6 Type:CANB, K, MEL, l-ER:ll!:,1-6q?,.(_5_. Type: CANB, K, NY, PERTH), 4683 (8 BRI, NSw,'Ny, pERfHj, 4685 (2 PERTH,WAIT),4746 (2),4965 (9),4996 (4),5003 (4),5012 (4). MitchelLA.4.76/186 (9),269 (9 CANB,PERTH),341 (9) 348 (1-PERTH, WAIT),967 (2). O'Farte. R. 48(4J. Pul.ien,fi. 10940(6-AD, CANB,PERTH). St'ewartl{. s.n.Sept. 1957 (1). Trudsen,M. E.319 (6-BRI, PERTH),323 (6), 327 (8), 391 (8-Type: CANB,K, PERTH),1042 (8) 1045 !E, Lq4 (9), 1151(8 MEL, PERTH),1299 (2 PERTH,WAIT), 1338l9-TYPE: CANB, K, IqBT4). 24 (8-CANB,PERTH),2407 (? BM, NT, G,P, PERTH),2A13(7 T'PE:CANE, K; y^EL:l. P_EjflJl.21s3,2T!'plqANB.K.Ny.PERTH'.2503,2 BRr.pERTH;.2s48,7 AD, BRI,NSW, PERTH),25?6 (9 NSW,PERTH),2631(9 BM,BRI, PERTH). Walkea.I{.95(8-CBG, MEL, NSW, PERTH),96 (8),97 (8), 113 (8). Weston,A. S.10842(2), r0844A (2). Willis,J. H. PERTH-Dup. of MEL 503363(2), MEL 503564(6 MEL, PERTH),s.n. 1? Aus. 1974 (8) (8)-6599!

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