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Asidinimarruecoscompleto nº Monografías electrónicas S.E.A. ISSN: 2386-5318 33 Los Asidini marroquíes. Ensayo monográfico sobre la tribu Asidini (Coleoptera, Tenebrionidae) en el Reino de Marruecos Francisco Pérez-Vera & José Miguel Ávila SEociedad ntomológica A ragonesa www.sea-entomologia.org Zaragoza, 31-12-2012 Monografías electrónicas S.E.A.,3 Los Asidini marroquíes. Ensayo monográfico sobre la tribu Asidini (Coleoptera, Tenebrionidae) en el Reino de Marruecos Francisco Pérez-Vera & José Miguel Ávila SEociedad ntomológica A ragonesa www.sea-entomologia.org Zaragoza, 31-12-2012 Monografías electrónicas S.E.A.,3 Los Asidini marroquíes. Ensayo monográfico sobre la tribu Asidini (Coleoptera, Tenebrionidae) en el Reino de Marruecos Francisco Pérez-Vera & José Miguel Ávila SEociedad ntomológica A ragonesa www.sea-entomologia.org Zaragoza, 31-12-2012 Los Asidini marroquíes. F. Pérez-Vera & J. M. Ávila. Monografías electrónicas SEA, 3 (2012) 1 Sumario Préface. Fabien Soldati ................................................................... 3 Agradecimientos ............................................................................. 4 Introducción ..................................................................................... 5 Mapas de distribución de los Asidini paleárticos .......................................... 9 Mapas de distribución de los Asidini marroquíes ......................................... 9 Comentarios sobre la distribución geográfica del subgénero Glabrasida en Marruecos ............................................................ 10 Comentarios sobre la distribución geográfica de los subgéneros Machlasida y Pseudoelongasida en Marruecos ..................... 12 Comentarios sobre la distribución geográfica del género Asida en Marruecos ........................................................................... 14 El órgano genital masculino de los Asidini marroquíes y su significación en la Sistemática ......................................... 17 Sistemática de los Asidini de Marruecos .................................... 31 Género Alphasida: Subgénero Glabrasida .......................................................................... 33 Subgénero Machlasida ............................................................................. 53 Subgénero Pseudoelongasida ................................................................ 54 Género Asida: Subgénero Peltasida................................................................................. 54 Subgénero Globasida ............................................................................... 54 Subgénero Planasida ................................................................................ 55 Descriptiva de los Asidini de Marruecos ..................................... 59 Género Alphasida: Subgénero Glabrasida .......................................................................... 59 Subgénero Machlasida ........................................................................... 125 Subgénero Pseudoelongasida .............................................................. 156 Género Asida: Subgénero Peltasida............................................................................... 159 Subgénero Globasida ............................................................................. 162 Subgénero Planasida .............................................................................. 164 2 Los Asidini marroquíes. F. Pérez-Vera & J. M. Ávila. Monografías electrónicas SEA, 3 (2012) Apéndice: La tribu Asidini en el mundo .................................... 191 Índice de los taxones tratados en este trabajo ......................... 197 Bibliografía ................................................................................... 200 Planchas ............................................................................. I-CXXXVI Género Alphasida: Subgénero Glabrasida ............................................................................. I Subgénero Machlasida ...................................................................... LXXVI Subgénero Pseudoelongasida ............................................................... CX Género Asida: Subgénero Peltasida.............................................................................. CXII Subgénero Globasida .......................................................................... CXIV Subgénero Planasida .......................................................................... CXVII Los Asidini marroquíes. F. Pérez-Vera & J. M. Ávila. Monografías electrónicas SEA, 3 (2012) 3 Préface Les Tenebrionidae constituent une des familles de Coléoptères parmi les plus diversifiées au niveau mondial, avec environ 19.000 espèces actuellement décrites. En réalité, beaucoup nous sont encore inconnues, probablement au moins autant. Ils font partie des familles dont l'étude est encore très fragmentaire, nécessitant la réunion d'une bibliographie importante et souvent ancienne. Un des problèmes majeurs pour étudier les Tenebrionidae est la quasi absence de monographies récentes. A cela s'ajoute leur grande diversité liée en partie à leur ancienneté, à l'aptérisme de la plupart de leurs représentants, à leur rapide capacité d'évolution et à leur grande aptitude à s'adapter à des niches écologiques très variées. On rencontre ainsi des Tenebrionidae sous toutes les latitudes, depuis les zones littorales jusqu'à près de 5.000 mètres d'altitude dans les Andes ou en Asie centrale himalayenne. Ils abondent surtout dans les régions chaudes du globe et l'endémisme qu'ils présentent est particulièrement élevé, en particulier dans les montagnes. A ce phénomène, les Asidini ne font pas exception, bien au contraire. On ne compte pas moins de 331 espèces pour les seuls deux genres Asida et Alphasida en région paléarc- tique, dont 85 espèces pour le seul pays du Maroc, le deuxième plus riche derrière l'Es- pagne. En dépit de leur proximité, ces deux pays n'ont seulement qu'une espèce en commun malgré le paléo-épisode de la Bétique ! En zone paléarctique, les Asidini n'occupent essen- tiellement que la partie occidentale du Bassin méditerranéen avec pour zones les plus diver- sifiées le Maghreb, la Péninsule Ibérique, les Baléares, l'Italie et ses îles ainsi que les Bal- kans. Comme beaucoup de groupes très diversifiés, les Asidini nécessitent une révision globale qui représente un travail considérable. Comment en effet pouvoir étudier simulta- nément autant de taxons, retrouver tous les types, confronter d'innombrables spécimens de collections institutionnelles et de particuliers. Et pouvoir étudier sur le terrain, quand cela est possible, certaines populations afin de pouvoir mieux définir leur statut taxonomique et avoir un aperçu de l’écologie des espèces considérées, ces insectes étant le plus souvent rares avec des périodes d'activité en hiver ou au début du printemps. Un tel ensemble s'en- treprend par groupe phylétique ou sur une zone géographique bien délimitée. C’est justement de cette manière que les deux auteurs, MM. F. PEREZ-VERA et J. M. AVILA, ont décidé d’étudier les Asidini, en commençant par le Maroc, qui totalise tout de même 135 taxons, en comptant les sous-espèces, dont de nombreux endémiques et on peut à ce titre saluer leur courage. Dans le cadre de ce travail, ils ont pu étudier la plupart des types et syntypes des taxons actuellement connus et nous livrent ici une synthèse richement commentée et iconographiée, accompagnée de toutes les descriptions originales, le plus souvent traduites en langue espagnole. Ce travail constitue un point de départ pour la future grande révision des Asidini marocains qu’ils envisagent et ont déjà sous presse plusieurs travaux d’importance majeure. En plus d'être d'une grande utilité, leur présente œuvre est également très agréable à consulter et constitue une remarquable source d'information. Fabien SOLDATI, Quillan, le 26/09/2011 4 Los Asidini marroquíes. F. Pérez-Vera & J. M. Ávila. Monografías electrónicas SEA, 3 (2012) Agradecimientos La realización de esta Monografía sobre los Asidini de Marruecos ha sido posible gracias al concurso de numerosos factores. Hemos contado desde el primer día con el estímulo del equipo del Departamento de Zoología de la Facultad de Ciencias de Granada, en el que destacamos al Profesor Francisco Sánchez Piñero, que puso a nuestra disposición su colec- ción de Tenebrionidae marroquíes y con el que compartimos frecuentemente sus conoci- mientos en esta fauna entomológica. Debemos resaltar la inapreciable ayuda de los respon- sables de las diferentes instituciones nacionales y extranjeras, que guardan las más impor- tantes colecciones de estos insectos y que accedieron a ponerlas a nuestra disposición. Un lugar preeminente ocupa la Dra. Mercedes Paris, encargada de la colección entomológica del MNCN de Madrid, que nos permitió el estudio de las importantes colecciones, de Esca- lera y de Cobos entre otras, que se encuentran conservadas en este Museo. Al mismo tiem- po expresamos nuestro agradecimiento por la amable disponibilidad de Dª Mercedes Hitado y de la Dra. Amparo Blay, encargadas del Laboratorio de Entomología. De forma muy es- pecial reconocemos el apoyo y los consejos siempre oportunos del Dr. Miguel Ángel Alon- so Zarazaga, Científico titular del departamento de Biodiversidad y Biología Evolutiva del mismo centro. El Dr. Ottó Merkl, del Hungarian Natural History Museum of Budapest (Hungría), nos ha comunicado el material que le solicitamos de la colección Reitter, algún espécimen preciado de Koch y varios ejemplares
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