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Home , Belgrandiella, Daudebardia, Eupulmonata, Ferussaciidae, Gyraulus rossmaessleri, Lindholmiola

Vol. 15(2): 49–58

CHECKLIST OF THE MOLLUSCS () OF THE SLOVAK REPUBLIC

TOMÁŠ ÈEJKA*, LIBOR DVOØÁK, MICHAL HORSÁK, JOZEF ŠTEFFEK

*Correspondence: Institute of Zoology, Slovak Academy of Sciences, Dúbravská cesta 9, SK-84506 Bratislava, Slovak Republic (e-mail: [email protected]) ABSTRACT: The checklist of 245 mollusc known so far from the Slovak Republic is presented, plus 11 species limited to greenhouses or thermal waters. Critical comments on species erroneously mentioned in re- cent publications from Slovakia are included. KEY WORDS: Mollusca, checklist, Slovak Republic

INTRODUCTION

Research of Slovak molluscs started at the begin- cal evaluation of the previously published checklists ning of the 20th century (CSIKI 1918). In the first half (BANK et al. 2001, ŠTEFFEK &GREGO 2002). We deci- of the 20th century J. F. BABOR and later also his col- ded to use the monograph Molluscs of Slovakia (LI- league J. PETRBOK worked on the Slovak malaco- SICKÝ 1991) as the most suitable baseline because it fauna. Unfortunately their publications were not sys- contains the most recent reliable list of Slovak tematic and especially not critical enough, resulting molluscs. Therefore the original literature sources in erroneous records of some mollusc species in Slo- are given for all the species first recorded in the Slo- vakia (LISICKÝ 1991). The situation changed after vak Republic after 1982. World War II. The work of the new generation of The checklist of Slovak molluscs published by ŠTEF- malacologists resulted in a reliable knowledge about FEK &GREGO (2002) has several shortcomings. The the fauna. The entire research was dominated by the authors uncritically adopted many taxa from the work of V. LOZEK, who critically and systematically CLECOM List (BANK et al. 2001), though several of the evaluated Slovak molluscan fauna for the first time species do not occur in Slovakia and/or their occur- (LOZEK 1956). He also stimulated and indirectly man- rence has not been confirmed, revised or their records aged the work of other Czech (J. BRABENEC, S. MÁ- have not been published yet (see next chapter). On CHA, I. FLASAR, V. HUDEC) and upcoming Slovak the other hand, the following Slovak species were miss- malacozoologists (L. KALAŠ, M. J. LISICKÝ, J. ŠTEFFEK, ing from their list: Theodoxus fluviatilis (L.), V. LUÈIVJANSKÁ-KROUPOVÁ, T. ÈEJKA). The Slovak ma- panormitanum (Lessona et Pollonera), Sphaerium lacological research, initially focused on faunistics, solidum (Normand) and Corbicula fluminea (Müll.). The now extended to include zoocoenological surveys or authors did not consider the taxonomic revisions of monographs (KROUPOVÁ 1977, ŠTEFFEK 1978). LISIC- the Alzoniella (BERAN &HORSÁK 2001) and the KÝ (1991) was the first to compile a synthetic mono- family , and thus used junior synonyms graph of the Slovak molluscan fauna. When examin- (FALKNER et al. 2002). Furthermore, they listed green- ing several collections he realised that for the sake of house alien species and those living in the wild without reliability, it would be advisable to limit the study to a clear distinction. Non-native, exclusively greenhouse post-war collections. He verified and processed al- immigrants are usually listed separately, in order to most all published and collection data from prevent misrepresentation, for example when compar- 1945–1982. ing species spectra of the regional checklists. The The aims of this checklist are: (1) updating and authors listed subspecies which is often redundant and supplementing the list of Slovak molluscs and (2) criti- misleading. Finally, they included also problematic 50 Tomáš Èejka, Libor Dvoøák, Michal Horsák, Jozef Šteffek taxa in their list, which is not a commonly accepted diaphana (Draparnaud, 1805) – a single practice, especially when they are not classified at the specimen was found by V. LOZEK (LOZEK &ŠTEF- species level, e.g. Hauffenia sp., Bythiospeum cf. FEK 1989). Despite repeated visits to the locality, oshanovae (L. Pintér, 1968) etc. Some comments on no other individuals have been found; it is likely such species are provided below. that LOZEK’S specimen came from another locality repentina Hudec, 1960 – based on the re- and was misplaced accidentally. sults of allozyme analysis C. repentina is not a separ- declivis Sterki, 1893 – V. LOZEK found only ate species, but a form of C. lubrica (ARMBRUSTER one recent specimen at the bank of the Danube &SCHLEGEL 1994). river; it is possible that the specimen was washed Bythiospeum cf. oshanovae –ŠTEFFEK (1979) found it in down from the upper stretch of the river. Its occur- flood debris of the Danube river as a single empty rence in Slovakia is very unlikely. shell; live individuals have not been found to date. On the basis of a critical revision from 2006, the Hauffenia sp. – the genus is taxonomically problematic; list of Slovak molluscs comprises 245 native and natu- there exists material of recent, preserved individ- ralised species known as living populations from the uals in private collections (V. KOŠEL, J. ŠTEFFEK), territory of Slovakia (excluding non-native species though the species has not been described yet. restricted to greenhouses and species living in ther- dilatatus (Gould, 1841) – the species has not mal waters); of these 217 species (88.5%) are gastro- been recorded from Slovakia to date, and its ap- pods (52 freshwater) and 28 (11.5%) are bivalves. pearance in this country in the future is unlikely. The native and naturalised molluscs of Slovakia were Holandriana holandrii (C. Pfeffer, 1828) – one speci- divided into 10 main groups and 33 subgroups on men was found by V. LUÈIVJANSKÁ in 1991 (see the basis of their ecological requirements (LISICKÝ ŠTEFFEK 1996). The species is excluded from the 1991). The majority of terrestrial gastropods are present checklist, considering its accidental intro- woodland species sensu lato (55%), followed by duction. Subsequent malacological surveys in euryoecious and steppe species s. l. (both 10%), the 1994–2006 did not confirm its presence. remaining species represent other ecological groups corcyrensis (Férussac, 1839) – ŠTEFFEK (see Fig. 1). (1982) found an old empty shell in the flood de- bris of the Danube river between the Kravany n. Dunajom and Moèa. The species is excluded from the present list for the same reasons as Holandriana holandrii (see above). dunkeri (L. Pfeiffer, 1865) – the species has been synonymised with O. elegans (FALKNER et al. 2002). bielzii Seibert, 1874 – the species does not oc- cur in Slovakia. All records of J. BRABENEC and V. HUDEC have been revised by A. WIKTOR (Wroc- ³aw), and all have turned out to be Limax cinereo- niger (Wolf) (WIKTOR, pers. comm.). Moreover, the taxon may not be a valid species. hortensis A. Férussac, 1819 – the species was for- merly believed to occur in all of Europe, then based on genital characters turned out to be a complex of three species of which sensu stricto is distributed only in Western Europe. In Central Europe (including Slovakia) only (Mabille) is found (DE WINTER 1984, BACKELJAU 1987). Fig. 1. Ecological groups of land gastropods in Slovakia (ac- Normand, 1852 – the species does cording to LISICKÝ 1991, slightly modified by the not occur in Slovakia, the nearest records are in authors): EU – euryoecious species, HG – hygrophilous Silesia, Czech Republic (LISICKÝ 1991). species, PE – rock-dwelling species, PT – praticolous fuscus (C. Pfeiffer, 1821) and S. occultus (heliophilous) species, SIe – euryoecious woodland spe- (Jackiewicz, 1959) – both are West-European spe- cies, SIh – hygrophilous woodland species, SIs – cies, and their occurrence in Slovakia is unlikely. stenoecious woodland species, SIp – rock-dwelling heterostropha (Say, 1817) – is a of P. woodland species, SIth – shrub-dwelling species (thamnophilous), ST – steppe species, STp – steppe acuta (DILLON et al. 2002, WETHINGTON et al. rock-dwelling species, RP – polyhygropilous (ripico- 2003). lous) species, XC – xerophile species Checklist of the molluscs of the Slovak Republic 51

Fig. 2. Ecological groups of aquatic molluscs in Slovakia Fig. 3. Zoogeographical groups of molluscs in Slovakia (ac- (LISICKÝ 1991): Et – euryoecious species, FN – fonti- cording to LISICKÝ 1991) colous species, PD – paludicolous species, RV – rivi- colous (rheophilic) species, SG – stagnicolous species

The freshwater species of Slovakia were divided into euryoecious (11%) and fonticolous species (8%) (Fig. five main groups: limnophilous (stagnicolous) species 2). From the zoogeographical point of view Palaearctic constitute the greatest proportion (36%), followed by (14%), European (13%), and Holarctic (11%) species rivicolous (rheophilous) (30%), paludicolous (15%), predominate (for details see Fig. 3).

CHECKLIST

Species living only in greenhouses arboreus (Say, 1816): FLASAR &KROUPOVÁ and thermal waters (1976)– it has been recorded in Hungary (KERNEY Melanoides tuberculatus (O. F. Müller, 1774): FLASAR & et al. 1983) and the Czech Republic (DVOØÁK & KROUPOVÁ (1976), MÁJSKY (2000), a prosperous KUPKA in press) and, like the previous species, may population of this tropical freshwater has spread outside greenhouses. been living for dozens of years in thermal waters in Pomacea bridgesii (Reeve, 1856) [= Ampullaria austra- Piešany (VARGA 1976) lis] – its yellow form was found by T. ÈEJKA pumilum (L. Pfeiffer, 1840) [=goodallii (Miller, (unpubl.) in a greenhouse of the botanic garden 1822)]: FLASAR &KROUPOVÁ (1976) in Bratislava. The species was probably brought io Verdcourt, 1974:FLASAR &KROUPOVÁ with imported aquatic plants or by aquarists. (1976) Native and naturalised species duryi (Wetherby, 1879) [in older literature incorrectly called trivolvis]:FLASAR & non-native species marked with an asterisk KROUPOVÁ (1976) Class: Pseudosuccinea (Say, 1817):FLASAR & Subclass: Prosobranchia KROUPOVÁ (1976) Superorder: Archaeogastropoda valentiana (A. Férussac, 1822): FLASAR & Order: Neritimorpha KROUPOVÁ (1976) Superfamily: Neritoidea (Lessona et Pollonera, 1882): Family: Neritidae DVOØÁK et al. (2003) – the species will probably be Theodoxus Montfort, 1810 found outside greenhouses in the future (see danubialis (C. Pfeiffer, 1828) HORSÁK &DVOØÁK 2003). fluviatilis (Linnaeus, 1758) *) 1) transversalis (C. Pfeiffer, 1828) 52 Tomáš Èejka, Libor Dvoøák, Michal Horsák, Jozef Šteffek

Superorder: Valvata O. F. Müller, 1774 Order: Architaenioglossa cristata O. F. Müller, 1774 Superfamily: Ampullarioidea macrostoma Mörch, 1864 4) Family: Viviparidae =V. pulchella auct. nec Studer, 1820 Viviparus Montfort, 1810 piscinalis (O. F. Müller, 1774) acerosus (Bourguignat, 1862) Borysthenia Lindholm, 1913 contectus (Millet, 1813) naticina (Menke, 1845) Order: Neotaenioglossa Subclass: Superfamily: Cerithioidea Superorder: Order: Family: Thiaridae Superfamily: Acroloxoidea Subfamily: Thiarinae Family: Acroloxidae Esperiana Bourguignat, 1877 Acroloxus Beck, 1838 esperi (A. Férussac, 1823) lacustris (Linnaeus, 1758) daudebartii (Prevost, 1821) Superfamily: Family: Family: Subfamily: Tateinae Galba Schrank, 1803 Potamopyrgus Stimpson, 1865 truncatula (O. F. Müller, 1774) antipodarum (Gray, 1843) *) 2) Stagnicola Jeffreys, 1830 =P. jenkinsi (E. A. Smith, 1889) corvus (Gmelin, 1791) Subfamily: Horatiinae = palustris (O. F. Müller, 1774) part. 4) Alzoniella Giusti et Bodon, 1984 palustris (O. F. Müller, 1774) 9) slovenica (Lozek et Brtek, 1964) 3) =turricula (Held, 1836) =Belgrandiella slovenica komenskyi Hudec, 1972 Radix Montfort, 1810 =Belgrandiella kalasi Lozek et Brtek, 1964 ampla (Hartmann, 1821) =Belgrandiella alticola Lozek et Brtek, 1964 auricularia (Linnaeus, 1758) =Belgrandiella bojnicensis Lozek et Brtek, 1964 ovata (Draparnaud, 1805) Subfamily: Amnicolinae =balthica (Linnaeus, 1758) Bythinella Moquin-Tandon, 1856 peregra (O. F. Müller, 1774) austriaca (von Frauenfeld, 1857) agg. =labiata (Rossmässler, 1835) hungarica Hazay, 1881 4) Lymnaea Lamarck, 1799 metarubra (Falniowski, 1987) 5) stagnalis (Linnaeus, 1758) pannonica (Frauenfeld,1865) Subfamily: Lithoglyphinae Superfamily: Physoidea Lithoglyphus Hartmann, 1821 Family: naticoides (C. Pfeiffer, 1828) Fleming, 1820 hypnorum (Linnaeus, 1758) Family: Bithyniidae Physa Draparnaud, 1801 Bithynia Leach, 1818 fontinalis (Linnaeus, 1758) leachii (Sheppard, 1823) Physella Haldemann, 1843 troschelii (Paasch, 1842) 6–8) acuta (Draparnaud, 1805) *) 10–11) =transsilvanica (E.A. Bielz, 1853) tentaculata (Linnaeus, 1758) Superfamily: Family: Superfamily: Aciculoidea O. F. Müller, 1774 Family: Aciculidae carinatus O. F. Müller, 1774 Acicula Hartmann, 1821 planorbis (Linnaeus, 1758) parcelineata (Clessin, 1911) Anisus Studer, 1820 Platyla Moquin-Tandon, 1856 leucostoma (Millet, 1813) polita (Hartmann, 1840) =septemgyratus (Rossmässler, 1835) Superorder: Heterostropha septemgyratus (Rossmässler, 1835) Order: Ectobranchia =calculiformis (Sandberger, 1874) Superfamily: Valvatoidea spirorbis (Linnaeus, 1758) Family: Valvatidae vortex (Linnaeus, 1758) vorticulus (Troschel, 1834) Charpentier, 1837 contortus (Linnaeus, 1758) Checklist of the molluscs of the Slovak Republic 53

Gyraulus Charpentier, 1837 sterrii (Voith, 1840) acronicus (A. Férussac, 1807) triplicata (Studer, 1820) albus (O. F. Müller, 1774) Cossmann, 1889 crista (Linnaeus, 1758) bielzi (Rossmässler, 1859) laevis (Alder, 1838) riparius (Westerlund, 1865) Family: 12) Fitzinger, 1833 rossmaessleri (Auerswald, 1852) 15) Charpentier, 1837 pusilla (Vallot, 1801) complanatus (Linnaeus, 1758) =P. rupestris (Draparnaud, 1801) part. Fleming, 1818 Family: nitida (O. F. Müller, 1774) Subfamily: Spelaeodiscinae Froriep, 1806 Brusina, 1886 corneus (Linnaeus, 1758) triarius (Rossmässler, 1839) Ancylus O. F. Müller, 1774 fluviatilis O. F. Müller, 1774 Family: Ferrissia Walker, 1903 Subfamily: Valloniinae fragilis (Tryon, 1863)13) Vallonia Risso, 1826 =wautieri (Mirolli, 1960) *)14) costata (O. F. Müller, 1774) =clessiniana (Jickeli, 1882) enniensis (Gredler, 1856) excentrica Sterki, 1893 Superorder: pulchella (O. F. Müller, 1774) Order: Actophila Subfamily: Acanthinulinae Superfamily: Ellobioidea Beck, 1847 Family: Carychiidae aculeata (O. F. Müller, 1774) O. F. Müller, 1774 minimum O. F. Müller, 1774 Family: tridentatum (Risso, 1826) Subfamily: Truncatellininae Columella Westerlund, 1878 Order: aspera Waldén, 1966 16) Suborder: columella (Clessin, 1872) Superfamily: Cochlicopoidea edentula (Draparnaud, 1805) Family: Lowe, 1852 Cochlicopa A. Férussac, 1821 claustralis (Gredler, 1856) lubrica (O. F. Müller, 1774) costulata (Nilsson, 1823) =repentina (Hudec, 1960) cylindrica (A. Férussac, 1807) lubricella (Rossmässler, 1835) Subfamily: Vertigininae nitens (M. von Gallenstein, 1848) O. F. Müller, 1774 Superfamily: alpestris Alder, 1838 Family: angustior Jeffreys, 1830 Held, 1838 antivertigo (Draparnaud, 1801) dolium (Draparnaud, 1801) geyeri Lindholm, 1925 Clessin, 1876 modesta (Say, 1824) pagodula (Des Moulins, 1830) =arctica (Wallenberg, 1858) Charpentier, 1837 moulinsiana (Dupuy, 1849) doliolum (Bruguière, 1792) pusilla O. F. Müller, 1774 pygmaea (Draparnaud, 1801) Family: substriata (Jeffreys, 1833) Turton, 1831 secale (Drapanaud, 1801) Superfamily: Buliminoidea Held, 1838 Family: Buliminidae frumentum (Drapanaud, 1801) Beck, 1837 Reichenbach, 1828 tridens (O. F. Müller, 1774) clienta (Westerlund, 1883) Turton, 1831 tatrica Lozek, 1948 montana (Draparnaud, 1801) Held, 1838 Family: obscura (O. F. Müller, 1774) Fleming, 1828 Held, 1838 alpicola (Charpentier, 1837) detrita (O. F. Müller, 1774) muscorum (Linnaeus, 1758) 54 Tomáš Èejka, Libor Dvoøák, Michal Horsák, Jozef Šteffek

Suborder: Superfamily: Superfamily: Family: Family: A. Férussac, 1814 Subfamily: Alopiinae acicula (O. F. Müller, 1774) Alopia petitiana (Benoit, 1862) 19) bielzii (E. A. Bielz, 1856) A. Férussac, 1821 Superfamily: cerata (Rossmässler, 1836) Family: fimbriata remota Lozek 1952 Subfamily: Punctinae laminata (Montagu, 1803) Morse, 1864 orthostoma (Menke, 1828) pygmaeum (Draparnaud, 1801) Subfamily: Clausiliinae Subfamily: Helicodiscinae Lindholm, 1924 Lowe, 1852 filograna (Rossmässler, 1836) =Hebetodiscus (Baker, 1929) 16) Pseudofusulus H. Nordsieck, 1977 singleyana (Pilsbry, 1890) *) varians (C. Pfeiffer, 1828) =inermis (Baker, 1929) Hartmann, 1841 Family: latestriata (A. Schmidt, 1857) Fitzinger, 1833 17) =M. borealis (O. Boettger, 1878) perspectivus (M. von Mühlfeld, 1816) plicatula (Draparnaud, 1801) rotundatus (O. F. Müller, 1774) tumida (Rossmässler, 1836) ruderatus (A. Férussac, 1821) ventricosa (Draparnaud, 1801) Draparnaud, 1805 Superfamily: Vitrinoidea cruciata (Studer, 1820) Family: dubia Draparnaud, 1805 Zonitoides Lehmann, 1862 parvula Férussac, 1807 nitidus (O. F. Müller, 1774) =C. rugosa parvula Férussac, 1807 18) pumila C. Pfeiffer, 1828 Family: Subfamily: Baleinae Reinhardt, 1833 Hartmann, 1844 fulvus (O. F. Müller, 1774) plicata (Draparnaud, 1801) praticola (Reinhardt, 1883) H. & A. Adams, 1855 =alderi (Gray, 1840) biplicata (Montagu, 1803) Family: Pseudalinda O. Boettger, 1877 Draparnaud, 1801 stabilis (L. Pfeiffer, 1847) pellucida (O. F. Müller, 1774) Balea Gray, 1824 Eucobresia H. B. Baker, 1929 perversa (Linnaeus, 1758) nivalis (Dumont et Mortillet, 1854) P. Hesse, 1916 Agassiz, 1845 elata (Rossmässler, 1836) kotulae (Westerlund, 1883) gulo (E. A. Bielz, 1859) semilimax (J. Férussac, 1802) turgida (Rossmässler, 1836) O. Boettger, 1877 Family: cana (Held, 1836) Subfamily: Vitreinae Fitzinger, 1833 Superfamily: contracta (Westerlund, 1871) Family: Succineidae crystallina (O. F. Müller, 1774) Mabille, 1871 diaphana (Studer, 1820) oblonga (Draparnaud, 1801) subrimata (Reinhardt, 1871) Draparnaud, 1801 transsylvanica (Clessin, 1877) putris (Linnaeus, 1758) Subfamily: Zonitinae Oxyloma Westerlund, 1885 Lindholm, 1927 elegans (Risso, 1826) epipedostoma (Fagot, 1879) 21) =Succinea pfeifferi Rossmässler, 1835 minor (Stabile, 1864) =S. dunkeri L. Pfeiffer, 1865 nitens (Michaud, 1831) Quickella C. Boettger, 1939 pura (Alder, 1830) arenaria (Bouchard-Chantereaux, 1837) Checklist of the molluscs of the Slovak Republic 55

Perpolita H. B. Baker, 1928 Superfamily: hammonis (Ström, 1765) Family: =P. radiatula (Alder, 1830) Arion A. Férussac, 1819 petronella (L. Pfeiffer, 1853) circumscriptus Johnston, 1828 28) Subfamily: Oxychilinae distinctus Mabille, 1868 29) Fitzinger, 1833 =A. hortensis Férussac, 1819 part. cellarius (O. F. Müller, 1774) fasciatus (Nilsson, 1823) depressus (Sterki, 1880) fuscus (O. F. Müller, 1774) orientalis (Clessin, 1877) =A. subfuscus (Draparnaud, 1805) part. =Schistophallus orientalis (Clessin, 1877) lusitanicus Mabille, 1868 *) 30) =Celariopsis deubeli (A. J. Wagner, 1914) rufus (Linnaeus, 1758) draparnaudi (Beck, 1837) silvaticus Lohmander, 1937 glaber (Rossmässler, 1835) hydatinus (Rossmässler, 1838) 22) Superfamily: inopinatus (Ulièný, 1887) Family: Bradybaenidae Held, 1838 Family: Daudebardiidae fruticum (O. F. Müller, 1774) Hartmann, 1821 brevipes (Draparnaud, 1805) Family: rufa (Draparnaud, 1805) Subfamily: Helicodontinae A. J. Wagner, 1895 A. Férussac, 1821 calophana (Westerlund, 1881) obvoluta (O. F. Müller, 1774) Subfamily: Monachinae Family: Westerlund, 1889 Lessona et Pollonera, 1882 strigella (Draparnaud, 1801) budapestensis (Hazay, 1881) 23) Fitzinger, 1833 = gracilis (Leydig, 1876) cartusiana (O. F. Müller, 1774) rustica (Millet, 1843) Subfamily: Chemnitz, 1786 Superfamily: =Trichia Hartmann, 1840 Family: hispidus (Linnaeus, 1758) Bielzia Clessin, 1887 striolatus (C. Pfeiffer, 1828) coerulans (M. Bielz, 1851) villosulus (Rossmässler, 1838) Limax Linnaeus, 1758 lubomirskii (Œlósarski, 1881) cinereoniger Wolf, 1803 Beck, 1847 maximus Linnaeus, 1758 b¹kowski (Poliñski, 1924) Lehmann, 1864 bielzi (E.A. Bielz, 1860) flavus (Linnaeus, 1758) 24) filicina (L. Pfeiffer, 1841) Malm, 1868 unidentata (Draparnaud, 1805) tenellus (O. F. Müller, 1774) Fitzinger, 1833 Lehmannia Heynemann, 1863 striata (O. F. Müller, 1774) macroflagellata Grossu et Lupu, 1962 Kobelt, 1871 marginata (O. F. Müller, 1774) unifasciata (Poiret, 1801) 31) nyctelia (Bourguignat, 1861) =soosiana (Wagner, 1933) Family: Xerolenta Monterosato, 1892 Deroceras Rafinesque, 1820 obvia (Menke, 1828) agreste (Linnaeus, 1758) = candicans (L. Pfeiffer, 1841) fatrense Mácha, 1981 25-26) Schlüter, 1838 laeve (O. F. Müller, 1774) bidentata (Gmelin, 1791) praecox Wiktor, 1966 25) =P. bidens (Chemnitz, 1786) reticulatum (O. F. Müller, 1774) dibothrion (M. von Kimakowicz, 1884) rodnae Grossu et Lupu, 1965 25) Gude et Woodward, 1921 sturanyi (Simroth, 1894) incarnatus (O. F. Müller, 1774) turcicum (Simroth, 1894) *) 27) vicinus (Rossmässler, 1842) Pseudotrichia Likharev, 1949 Family: Boettgerillidae rubiginosa (Rossmässler, 1838) Simroth, 1910 Lindholm, 1927 pallens Simroth, 1912 *) umbrosus (C. Pfeiffer, 1828) =B. vermiformis Wiktor, 1959 56 Tomáš Èejka, Libor Dvoøák, Michal Horsák, Jozef Šteffek

Hygromia Risso, 1826 Pseudanodonta Bourguignat, 1877 transsylvanica (Westerlund, 1876) complanata (Rossmässler, 1835) Sinanodonta Modell, 1945 Family: woodiana (Lea, 1834) *) 32) Subfamily: Ariantinae Turton, 1831 Subclass: Heterodonta arbustorum (Linnaeus, 1758) Order: Veneroida Faustina Kobelt, 1904 Superfamily: Sphaerioidea faustina (Rossmässler, 1835) Family: Corbiculidae rossmässleri (L. Pfeiffer, 1842) Corbicula M. von Mühlfeld, 1811 Fitzinger, 1833 fluminea (O. F. Müller, 1774) *) 33) cingulella (L. Pfeiffer, 1842) Family: Sphaeriidae Fitzinger, 1833 Sphaerium Scopoli, 1777 isognomostomos (Schröter, 1784) corneum (Linnaeus, 1758) =I. personatum (Lamarck, 1792) rivicola (Lamarck, 1818) Schileyko, 1971 solidum (Normand, 1844) 1) holosericea (Studer, 1820) nucleus (Studer, 1820) 34) Subfamily: Helicinae Musculium Link, 1807 Held, 1838 lacustre (O. F. Müller, 1774) hortensis (O. F. Müller, 1774) Pisidium C. Pfeiffer, 1821 vindobonensis (A. Férussac, 1821) amnicum (O. F. Müller, 1774) Linnaeus, 1758 casertanum (Poli, 1791) lutescens Rossmässler, 1837 henslowanum (Sheppard, 1823) pomatia Linnaeus, 1758 hibernicum Westerlund, 1894 35) 36) Class: Bivalvia globulare Clessin, 1873 Subclass: Palaeoheterodonta milium Held, 1836 Order: Unionoida moitessierianum Paladilhe, 1866 Superfamily: Unionoidea nitidum Jenyns, 1832 Family: Unionidae obtusale (Lamarck, 1818) Subfamily: Unioninae personatum Malm, 1855 Unio Philipsson, 1788 pseudosphaerium Favre, 1927 crassus Philipsson, 1788 subtruncatum Malm, 1855 pictorum (Linnaeus, 1758) supinum A. Schmidt, 1851 tumidus Philipsson, 1788 tenuilineatum Stelfox, 1918 Subfamily: Anodontinae Superfamily: Dreissenoidea Anodonta Lamarck, 1799 Family: Dreissenidae anatina (Linnaeus, 1758) Dreissena van Beneden, 1835 cygnea (Linnaeus, 1758) polymorpha (Pallas, 1771) *)

ORIGINAL SOURCES FOR SPECIES RECORDED AFTER 1982; COMMENTS

1) ÈEJKA T., HORSÁK M. 2002. First records of Theodoxus 6) GLÖER P. 2002: Bithynia leachii troschelii (Paasch, 1842) – fluviatilis and Sphaerium solidum (Mollusca) from Slo- die östliche Rasse von Bithynia leachii (Sheppard, 1823) vakia. Biologia 57: 561–562. (Gastropoda. : Bithyniidae). Arch. 2) ÈEJKA T. 1994. First record of the New Zealand mollusc Moll. 130: 259–265. Potamopyrgus antipodarum (Gray, 1843), (Gastropoda, Hy- 7) NESEMANN H. 1994. Zum Vorkommen von Bithynia drobiidae) from the Slovak section of the Dunaj river. leachii (Sheppard, 1823) und B. transsilvanica (Bielz, Biologia 49: 657–658. 1853) im Donaugebiet (Gastropoda: Prosobranchia: 3) BERAN L., HORSÁK M. 2001. Taxonomic revision of the Bithyniidae). Nachr. bl. vorarlb. malakol. Ges. 2: 5–13. genus Alzoniella (Mollusca, Gastropoda) in the Czech Re- 8) FALNIOWSKI A., GLÖER P., S ZAROWSKA M. 2004. Bithynia public and Slovakia. Biologia 56: 141–148. troschelii (Paasch, 1842), a giant of unknown origin? Folia 4) GLÖER P. 2002. Süßwassergastropoden Nord- und Mitte- Malacol. 12: 137–139. leuropas. ConchBooks, Hackenheim. 9) BARGUES M. D., VIGO. M., HORAK P., D VORAK J., 5) FALNIOWSKI A. 1979. Hydrobioidea of Poland (Proso- PATZNER R. A., POINTIER J. P., JACKIEWICZ M., branchia: Gastropoda). Folia Malacol. 1: 1–119. MEIER-BROOK C., MAS-COMA S. 2001. European Lymnae- idae (Mollusca: Gastropoda), intermediate hosts of tre- Checklist of the molluscs of the Slovak Republic 57

matodiases, based on nuclear ribosomal DNA ITS-2 se- 24) ŠTEFFEK J. 2005. Spoloèenstvá suchozemských mäkkýšov quences. Infection, Genetics and Evolution 1: 85–107. severnej èasti Devínskej Kobyly (Malé Karpaty). Malaco- 10) BERAN L. 2005. Which Physella (Mollusca: Gastropoda) logica Bohemoslovaca 4: 26–38. Online at http:// lives in the Czech Republic? Acta Soc. Zool. Bohem. 68: mollusca.sav.sk, 18-November-2005. 241–243. 25) HUDEC V. 1970. Für die Tschechoslowakei neue Nackt- 11) DILLON R. T., WETHINGTON A. R., RHETT J. M., SMITH T. schneckenarten Arten (Pulmonata, Limacidae, Deroce- P. 2002. Populations of the European freshwater pul- ras). Biológia 25: 109–112. monate Physa acuta are not reproductively isolated from 26) A population of this taxon is currently being studied. At American Physa heterostropha or Physa integra. Inverte- present it is impossible to judge whether it should be brate Biology 121: 226–234. ranked as a species or just as an extreme local form of D. 12) ÈEJKA T. 2005. Record of living individual of the fresh- rodnae (REISE &HUTCHINSON, pers. comm.) water snail rossmaessleri (Auerswald, 1852) in 27) REISE H., HUTCHINSON J. M. C. 2001. Morphological Slovakia after thirty-eight years (Gastropoda: Planor- variation in terrestrial Deroceras turcicum (Simroth, bidae). Malacologica Bohemoslovaca 4: 1–2. Online at 1894) and a northern extension of its range in Central http://mollusca.sav.sk 20-May-2005. Europe. Folia Malacol. 9: 63–71. 13) WALTHER A. C., LEE T., BURCH J. B., O’FOIGHIL D. 2006. 28) The species probably lives as least in NW. Slovakia, but all Confirmation that the North American ancylid Ferrissia the material dissected to date failed to confirm its pres- fragilis (Tryon, 1863) is a cryptic invader of European ence. Earlier records based on colouration are irrelevant. and East Asian freshwater ecosystems. J. Moll. Stud. 72: The species is known from the Czech Republic from the 318–321. surroundings of Mariánské Láznì, Prague, Køivoklátsko 14) GLÖER P., Z ETTLER M. L. 2005. Kommentierte Artenliste region (DVOØÁK &TUÈKOVÁ 2004) and Silesian region. der Süsswassermollusken Deutschlands. Malak. Abh. 23: 29) BACKELJAU T. 1987. Electrophoretic distinction between 3–26. Arion hortensis, A. distinctus and A. owenii (Mollusca: Pul- 15) GITTENBERGER E., BANK R. A. 1996. A new start in monata). Zool. Anz. 219: 33–39. Pyramidula (Gastropoda Pulmonata: Pyramidulidae). 30) REISCHÜTZ P. L. 1994. Mabille 1868 in Basteria 60: 71–78. der Slowakei (Gastropoda: Stylommatophora: Ariona- 16) ÈEJKA T., DVOØÁK L., HORSÁK M. 2006. Malakologické cea). Nachrichtenblatt der Ersten Vorarlberger Malako- novinky na Slovensku v poslednom štvrstoroèí. Malako- logischen Gesellschaft 2: 21. logický bulletin. Online at http://mal-bull.blogspot.com, 31) PFENNINGER M., MAGNIN F. 2001. Phenotypic evolution 14-December-2006. and hidden speciation in ssp. (Heli- 17) NORDSIECK H. 1993. Beiträge zur Nomenklatur der eu- cellinae, Gastropoda) inferred by 16S variation and quan- ropäischen Binnenmollusken, 1. Kritische Anmerkun- titative shell traits. Mol. Ecol. 10: 2541–2554. gen und Berichtigungen zur Nomenklatur von Arttaxa 32) KOŠEL V. 1995. The first record of Anodonta woodiana der Clausiliidae. Heldia 2: 33–42. (Mollusca, Bivalvia) in Slovakia. Acta Zoologica Univer- 18) NORDSIECK H. 1993. Revision der Gattung Clausilia Dra- sitatis Comenianae 39: 3. parnaud, besonders der Arten in SW-Europa (Das Clau- 33) ŠTEFFEK J., ÈEJKA T., NAGEL K. O. 2002. The distribution silia rugosa-Problem) (Gastropoda: Stylommatophora: of Corbicula fluminea in the Slovakian parts of the river Clausiliidae). Arch. Moll. 119: 133–179. Danube. Soosiana 23/30: 72–73. 19) LOZEK V. 1964. Quartärmollusken der Tschechoslowa- 34) KOŠEL V. 2006. The first record of Sphaerium nucleus kei. Rozpr. Ústø. Úst. Geol., Praha. (Bivalvia) in Slovakia. Biologia 61: 524. 20) ÈEJKA T. 2000. First record of the 35) ŠTEFFEK J. 2005. First record of Pisidium hibernicum (Hebetodiscus) singleyanus inermis (Gastropoda, Punctidae) (Mollusca, Bivalvia) from Slovakia. Biologia 60: 136. in Slovakia. Biologia 55: 475–476. 36) HORSÁK M., NEUMANOVÁ K. 2004. Distribution of Pisi- 21) HUDEC V. 1964. Zum Vorkommen der Schnecke Aego- dium globulare Clessin, 1873 (Mollusca: Bivalvia) in the pinella epipedostoma (Fag.) und der anderen Arten der Czech Republic and Slovakia with notes to its ecology Gattung Aegopinella Lindh. in der Tschechoslowakei. Acta and morphological characters. J. Conch. 4: 373–381. Mus. Natl. Pragae, Ser. B 20: 119–132. 22) DVOØÁK L., HORSÁK M., VARGA A. 2004. Oxychilus (Medi- ACKNOWLEDGEMENTS terranea) hydatinus (Gastropoda: Zonitidae) new for Slo- vakia, with notes on its distribution in Hungary. Folia The manuscript preparation was partly supported Malacol. 12: 149–152 by the long-term research plans of Masaryk University, 23) DVOØÁK L., ÈEJKA T., HORSÁK M. 2003. Present knowl- Brno (MSM 0021622416). edge of distribution of (Hazay, 1881) in the Czech and Slovak Republics (Gastropoda: Milacidae). Malacological Newsletter 21: 37–43. 58 Tomáš Èejka, Libor Dvoøák, Michal Horsák, Jozef Šteffek

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